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Cryptomonads
The cryptomonads (or cryptophytes) are a group of algae, most of which have plastids. They are common in freshwater, and also occur in marine and brackish habitats. Each cell is around 10–50 μm in size and flattened in shape, with an anterior groove or pocket. At the edge of the pocket there are typically two slightly unequal flagella. Some may exhibit mixotrophy. Characteristics Cryptomonads are distinguished by the presence of characteristic extrusomes called ejectosomes, which consist of two connected spiral ribbons held under tension. If the cells are irritated either by mechanical, chemical or light stress, they discharge, propelling the cell in a zig-zag course away from the disturbance. Large ejectosomes, visible under the light microscope, are associated with the pocket; smaller ones occur underneath the periplast, the cryptophyte-specific cell surrounding. Except for the class ''Goniomonadea'',_which_lacks_plastids_entirely,_and_''Cryptomonas_paramecium''_(pre ...
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Rhodomonas
''Rhodomonas'' is a genus of cryptomonads. It is characterized by its red colour, the square-shaped plates of its inner periplast, its short furrow ending in a gullet, and a distinctly shaped chloroplast closely associated with its nucleomorph. Historically, ''Rhodomonas'' was characterized by its red chloroplast alone, but this no longer occurs as its taxonomy has become increasingly based on molecular and cellular data. Currently, there is some debate about the taxonomic validity of ''Rhodomonas'' as a genus and further research is needed to verify its taxonomic status. ''Rhodomonas'' is typically found in marine environments, although freshwater reports exist. It is commonly used as a live feed for various aquaculture species. History of Knowledge Discovery ''Rhodomonas'' was first described by G. Karsten in 1898 as a “strange, swimming organism”. The first indication of ''Rhodomonas’'' existence was Karsten's observation of its proliferation in a sample containing di ...
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Nucleomorph
Nucleomorphs are small, vestigial eukaryotic nuclei found between the inner and outer pairs of membranes in certain plastids. They are thought to be vestiges of primitive red and green algal nuclei that were engulfed by a larger eukaryote. Because the nucleomorph lies between two sets of membranes, nucleomorphs support the endosymbiotic theory and are evidence that the plastids containing them are complex plastids. Having two sets of membranes indicate that the plastid, a prokaryote, was engulfed by a eukaryote, an alga, which was then engulfed by another eukaryote, the host cell, making the plastid an example of secondary endosymbiosis. Organisms with known nucleomorphs So far, only two monophyletic groups of organisms are known to contain plastids with a vestigial nucleus or nucleomorph: the cryptomonads of the supergroup Chromista and the chlorarachniophytes of the supergroup Rhizaria, both of which have examples of sequenced nucleomorph genomes. Studies of the genomic organi ...
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Hacrobia
The cryptomonads-haptophytes assemblage is a proposed monophyletic grouping of unicellular eukaryotes that are not included in the SAR supergroup. Several alternative names have been used for the group, including Hacrobia (derived from "ha-" referring to Haptophyta, "-cr-" referring to cryptomonads, and "-bia" as a general suffix referring to life); CCTH (standing for Cryptophyta, Centrohelida, Telonemia and Haptophyta); and "Eukaryomonadae". , it is unclear whether this group is monophyletic or not; results of phylogenetic studies are "often dependent on the selection of taxa and gene data set". Two 2012 studies produced opposite results. Members In the past, heterokonts, haptophytes, and cryptomonads have sometimes been grouped together in a group known as chromists. Though the heterokonts are now split out, Cryptophyta and Haptophyta are considered in some studies to be closely related (and are sometimes simply referred to as the "Cryptophyta+Haptophyta" group). A 2009 pape ...
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Cryptophyceae
The cryptophyceae are a class of algae, most of which have plastids. About 220 species are known, and they are common in freshwater, and also occur in marine and brackish habitats. Each cell is around 10–50 μm in size and flattened in shape, with an anterior groove or pocket. At the edge of the pocket there are typically two slightly unequal flagella. Some exhibit mixotrophy. Characteristics Cryptophytes are distinguished by the presence of characteristic extrusomes called ejectosomes or ejectisomes, which consist of two connected spiral ribbons held under tension. If the cells are irritated either by mechanical, chemical or light stress, they discharge, propelling the cell in a zig-zag course away from the disturbance. Large ejectosomes, visible under the light microscope, are associated with the pocket; smaller ones occur underneath the periplast, the cryptophyte-specific cell surrounding. Except for '' Chilomonas'', which has leucoplasts, cryptophytes have one or ...
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Cryptophyta Cell Scheme
The cryptophyceae are a class of algae, most of which have plastids. About 220 species are known, and they are common in freshwater, and also occur in marine and brackish habitats. Each cell is around 10–50 μm in size and flattened in shape, with an anterior groove or pocket. At the edge of the pocket there are typically two slightly unequal flagella. Some exhibit mixotrophy. Characteristics Cryptophytes are distinguished by the presence of characteristic extrusomes called ejectosomes or ejectisomes, which consist of two connected spiral ribbons held under tension. If the cells are irritated either by mechanical, chemical or light stress, they discharge, propelling the cell in a zig-zag course away from the disturbance. Large ejectosomes, visible under the light microscope, are associated with the pocket; smaller ones occur underneath the periplast, the cryptophyte-specific cell surrounding. Except for '' Chilomonas'', which has leucoplasts, cryptophytes have one or ...
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Periplast
The periplast is one of three types of cell-covering of three classes of algae. The ''Cryptomonads'' have the periplast covering. The '' Dinophyceae'' have a type called the amphiesma, and the ''Euglena'' covering is the pellicle. Structure The periplast is a proteinaceous covering. It can be subdivided into an inner periplast and an outer periplast. Both of these components are variable in their composition. The inner periplast may be formed as a single sheet as in the Chilomonas paramecium, or as multiple plates of varying shape. The outer periplast surrounds the plasma membrane. The inner periplast below the plasma membrane may sometimes as in ''Komma caudata ''Komma caudata'' is a cryptomonad, and the only described species in the genus ''Komma'', although four or five more species may exist. Its cells are 4.5–5.5 μm wide by 7–10 μm long and bear two unequal flagella A flagellum ...'' have a hexagonal arrangement of superficial periplast plates, and t ...
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Chloroplast
A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, and stores it in the energy-storage molecules ATP and NADPH while freeing oxygen from water in the cells. The ATP and NADPH is then used to make organic molecules from carbon dioxide in a process known as the Calvin cycle. Chloroplasts carry out a number of other functions, including fatty acid synthesis, amino acid synthesis, and the immune response in plants. The number of chloroplasts per cell varies from one, in unicellular algae, up to 100 in plants like '' Arabidopsis'' and wheat. A chloroplast is characterized by its two membranes and a high concentration of chlorophyll. Other plastid types, such as the leucoplast and the chromoplast, contain little chlorophyll and do not carry out photosynthesis. Chloroplasts are highly dynamic—they cir ...
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Cryptomonada
Cryptomonada is a proposed super-class of Cryptists linking Cryptophyceae and Goniomonadea, which was first proposed by Cavalier-Smith in 2004. He made this proposal based on three shared morphological characteristics: Presence of a periplast, ejectisomes with secondary scroll, and mitochondrial cristae A crista (; plural cristae) is a fold in the inner membrane of a mitochondrion. The name is from the Latin for ''crest'' or ''plume'', and it gives the inner membrane its characteristic wrinkled shape, providing a large amount of surface area fo ... with flat tubules. Genetic studies as early as 1994 also supported the hypothesis that ''Goniomonas'' was sister to Cryptophyceae. A study in 2018 found strong evidence that the common ancestor of ''Cryptomonada'' was an autotrophic protist. References {{Taxonbar, from=Q65088966 Cryptista ...
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Goniomonadea
Goniomonadea is a proposed class of cryptomonads The cryptomonads (or cryptophytes) are a group of algae, most of which have plastids. They are common in freshwater, and also occur in marine and brackish habitats. Each cell is around 10–50 μm in size and flattened in shape, with an anteri ... which includes the orders Goniomonadida and Hemiarmida. Taxonomy * Order Goniomonadida Novarino & Lucas 1993 oniomonadales Novarino & Lucas 1993** Family Goniomonadidae Hill 1991 oniomonadaceae Hill 1991*** Genus '' Goniomonas'' von Stein 1878 * Order Hemiarmida Cavalier-Smith 2017 ** Family Hemiarmidae Cavalier-Smith 2017 *** Genus '' Hemiarma'' Shiratori & Ishida 2016 References Cryptomonads {{Hacrobia-stub ...
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Thomas Cavalier-Smith
Thomas (Tom) Cavalier-Smith, FRS, FRSC, NERC Professorial Fellow (21 October 1942 – 19 March 2021), was a professor of evolutionary biology in the Department of Zoology, at the University of Oxford. His research has led to discovery of a number of unicellular organisms (protists) and advocated for a variety of major taxonomic groups, such as the Chromista, Chromalveolata, Opisthokonta, Rhizaria, and Excavata. He was known for his systems of classification of all organisms. Life and career Cavalier-Smith was born on 21 October 1942 in London. His parents were Mary Maude (née Bratt) and Alan Hailes Spencer Cavalier Smith. He was educated at Norwich School, Gonville and Caius College, Cambridge (MA) and King's College London (PhD). He was under the supervision of Sir John Randall for his PhD thesis between 1964 and 1967; his thesis was entitled "''Organelle Development in'' Chlamydomonas reinhardii". From 1967 to 1969, Cavalier-Smith was a guest inves ...
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Cryptomonas
''Cryptomonas'' is the name-giving genus of the Cryptomonads established by German biologist Christian Gottfried Ehrenberg in 1831. The algae are common in freshwater habitats and brackish water worldwide and often form blooms in greater depths of lakes. The cells are usually brownish or greenish in color and are characteristic of having a slit-like furrow at the anterior. They are not known to produce any toxins. They are used to feed small zooplankton, which is the food source for small fish in fish farms. Many species of ''Cryptomonas'' can only be identified by DNA sequencing. ''Cryptomonas'' can be found in several marine ecosystems in Australia and South Korea. Etymology ''Cryptomonas'' has the meaning of hidden small flagellates from “crypto” and “monas”. Genome Structure Species within ''Cryptomonas'' contain four genomes: the nuclear, the nucleomorph, the plastid, and mitochondrial genomes. The plastid genome contains 118 kilobase pairs and is a resu ...
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Chlorophyll
Chlorophyll (also chlorophyl) is any of several related green pigments found in cyanobacteria and in the chloroplasts of algae and plants. Its name is derived from the Greek words , ("pale green") and , ("leaf"). Chlorophyll allow plants to absorb energy from light. Chlorophylls absorb light most strongly in the blue portion of the electromagnetic spectrum as well as the red portion. Conversely, it is a poor absorber of green and near-green portions of the spectrum. Hence chlorophyll-containing tissues appear green because green light, diffusively reflected by structures like cell walls, is less absorbed. Two types of chlorophyll exist in the photosystems of green plants: chlorophyll ''a'' and ''b''. History Chlorophyll was first isolated and named by Joseph Bienaimé Caventou and Pierre Joseph Pelletier in 1817. The presence of magnesium in chlorophyll was discovered in 1906, and was that element's first detection in living tissue. After initial work done by German ch ...
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