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Claosaurus
''Claosaurus'' ( ; Greek κλάω, ''klao'' meaning 'broken' and , ''sauros'' meaning 'lizard'; "broken lizard", referring to the odd position of the fossils when discovered) is a genus of hadrosauroid dinosaur that lived during the Late Cretaceous Period (Santonian-Campanian). Traditionally classified as an early member of the family Hadrosauridae, a 2008 analysis found ''Claosaurus agilis'' to be outside of the clade containing ''Hadrosaurus'' and other hadrosaurids, making it the closest non-hadrosaurid relative of true hadrosaurids within the clade Hadrosauria. Description Because of the insufficient fossil remains ("minority of skull and partial skeleton"), the size of an adult ''Claosaurus'' remains uncertain. However, Thomas Holtz gave a length estimate of and a mass estimate as that of a lion. Like other hadrosaurs, it was an herbivore. History Evidence of its existence was first found in the Niobrara Formation near the Smoky Hill River in Kansas, United States ...
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Edmontosaurus
''Edmontosaurus'' ( ) (meaning "lizard from Edmonton") is a genus of hadrosaurid (duck-billed) dinosaur. It contains two known species: '' Edmontosaurus regalis'' and '' Edmontosaurus annectens''. Fossils of ''E. regalis'' have been found in rocks of western North America that date from the late Campanian stage of the Cretaceous Period 73 million years ago, while those of ''E. annectens'' were found in the same geographic region but in rocks dated to the end of the Maastrichtian stage of the Cretaceous, 66 million years ago. ''Edmontosaurus'' was one of the last non- avian dinosaurs, and lived alongside dinosaurs like ''Triceratops'', ''Tyrannosaurus'', '' Ankylosaurus'', and ''Pachycephalosaurus'' shortly before the Cretaceous–Paleogene extinction event. ''Edmontosaurus'' included some of the largest hadrosaurid species, with ''E. annectens'' measuring up to in length and weighing around in average asymptotic body mass, although some individuals would have been larg ...
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Hadrosauridae
Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includes genera such as '' Edmontosaurus'' and '' Parasaurolophus'', was a common group of herbivores during the Late Cretaceous Period. Hadrosaurids are descendants of the Upper Jurassic/Lower Cretaceous iguanodontian dinosaurs and had a similar body layout. Hadrosaurs were among the most dominant herbivores during the Late Cretaceous in Asia and North America, and during the close of the Cretaceous several lineages dispersed into Europe, Africa, South America and Antarctica. Like other ornithischians, hadrosaurids had a predentary bone and a pubic bone which was positioned backwards in the pelvis. Unlike more primitive iguanodonts, the teeth of hadrosaurids are stacked into complex structures known as dental batteries, which acted as ef ...
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Thespesius
''Thespesius'' (meaning "wondrous one") is a nomen dubium, dubious genus of hadrosaurid dinosaur from the Maastrichtian, late Maastrichtian-age Upper Cretaceous Lance Formation of South Dakota. History In 1855 geologist Ferdinand Vandeveer Hayden sent a number of fossils to paleontologist Joseph Leidy in Philadelphia. Hayden had collected them from the surface of a rock formation then known as the Great Lignite Formation (now recognized as part of the Lance Formation) in the Nebraska Territory, near the Grand River (South Dakota), Grand River (present-day South Dakota). Among them were two tail, caudal vertebrae and a phalanx bones, phalanx. In 1856 Leidy named the type species ''Thespesius occidentalis'' for these three bones.Leidy, J. (1856). "Notice of extinct Vertebrata, discovered by F. V. Hayden during the expedition to the Sioux country under the command of Lieut. G.K. Warren." ''Proceedings of the Academy of Natural Science Philadelphia'', 8(December 30): 311-312. The gen ...
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Pierre Shale
The Pierre Shale is a geologic formation or series in the Upper Cretaceous which occurs east of the Rocky Mountains in the Great Plains, from Pembina Valley in Canada to New Mexico. The Pierre Shale was described by Meek and Hayden in 1862 in the Proceedings of the Academy of Sciences (Philadelphia). They described it as a dark-gray shale, fossiliferous, with veins and seams of gypsum, and concretions of iron oxide. The Pierre Shale is about 700 feet (210m) thick at the type locality. It overlies the Niobrara division and underlies the Fox Hills beds. It was named for an occurrence near Fort Pierre on the Missouri River in South Dakota. The Pierre Shale is of marine origin and was deposited in the Western Interior Seaway. It is correlative with other marine shales that occur farther west, such as the Bearpaw Shale, Mancos Shale and the Lewis Shale. It correlates with the Lea Park Formation in central Alberta. The Pierre is overlain by marginal marine deposits of the F ...
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Hadrosauroid
Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to ''Iguanodon''. Their remains have been recovered in Asia, Europe, Africa and the Americas. Many primitive hadrosauroids, such as the Asian ''Probactrosaurus'' and ''Altirhinus'', have traditionally been included in a paraphyletic (unnatural grouping) "Iguanodontidae". With cladistic analysis, the traditional Iguanodontidae has been largely disbanded, and probably includes only ''Iguanodon'' and perhaps its closest relatives. Classification The cladogram below follows an analysis by Andrew McDonald, 2012, and shows the position of Hadrosauroidea within Styracosterna. The cladogram below follows an analysis by Wu Wenhao and Pascal Godefroit (2012). Cladogram after Prieto-Marquez and Norell (2010). A phylogenetic analysis performed by Ramírez-Velasco ''et al.'' (2012) found a large polytomy be ...
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Othniel Charles Marsh
Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of Paleontology in Yale College and President of the National Academy of Sciences. He was one of the preeminent scientists in the field of paleontology. Among his legacies are the discovery or description of dozens of new species and theories on the origins of birds. Born into a modest family, Marsh was able to afford higher education thanks to the generosity of his wealthy uncle George Peabody. After graduating from Yale College in 1860 he travelled the world, studying anatomy, mineralogy and geology. He obtained a teaching position at Yale upon his return. From the 1870s to 1890s, he competed with rival paleontologist Edward Drinker Cope in a period of frenzied Western American expeditions known as the Bone Wars. Marsh's greatest legacy is the collection of Mesozoic reptiles, Cretaceous birds, and Mesozoic and Tertiary mammals that now constitute the backbone of the collections of Yale's ...
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Hadrosaurus
''Hadrosaurus'' (; ) is a genus of hadrosaurid ornithopod dinosaurs that lived in North America during the Late Cretaceous Period in what is now the Woodbury Formation about 80 million to 78 million years ago. The holotype specimen was found in fluvial marine sedimentation, meaning that the corpse of the animal was transported by a river and washed out to sea. They were large animals ranging from and . Most of the preserved elements are very robust, unusual traits in hadrosaurs. ''Hadrosaurus'' were ponderously-built animals equipped with keratinous beaks for cropping foliage and a specialized and complex dentition for food processing. ''Hadrosaurus foulkii'', the only species in this genus, is known from a single specimen consisting of much of the skeleton and parts of the skull. The specimen was collected in 1858 from the Woodbury Formation in New Jersey, US, representing the first dinosaur species known from more than isolated teeth to be identified in North America. Using r ...
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Gastrolith
A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In other species the rocks are ingested and pass through the digestive system and are frequently replaced. The grain size depends upon the size of the animal and the gastrolith's role in digestion. Other species use gastroliths as ballast.Rondeau, et aLarval Anurans Adjust Buoyancy in Response to Substrate IngestionCopeia: February 2005, Vol. 2005, No. 1, pp. 188-195. Particles ranging in size from sand to cobble have been documented. Etymology Gastrolith comes from the Greek γαστήρ (''gastēr''), meaning "stomach", and λίθος (''lithos''), meaning "stone". Occurrence Among living vertebrates, gastroliths are common among crocodiles, alligators, herbivorous birds, seals and sea lions. Domestic fowl require access to ''grit''. Stone ...
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Smoky Hill River
The Smoky Hill River is a river in the central Great Plains of North America, running through Colorado and Kansas.Smoky Hill River. (2009). In Encyclopædia Britannica. Retrieved July 22, 2009, from Encyclopædia Britannica Online: http://www.britannica.com/EBchecked/topic/550068/Smoky-Hill-River Names The Smoky Hill gets its name from the Smoky Hills region of north-central Kansas through which it flows. American Indians living along the Smoky Hill considered it and the Kansas River to be the same river, and their names for it included Chetolah and Okesee-sebo. Early maps of European explorers called the river (also in combination with the Kansas River) the River of the Padoucas as its source is located in what was then Padouca ( Comanche) territory. The USGS lists a number of other variant names for the Smoky Hill River, including Chitolah River, Fork of the Hill Buckaneuse, La Fourche de la Cote Boucaniere, La Touche de la Cote Bucanieus, Manoiyohe, Pe P'a, Sand River, S ...
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Kansas
Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the west. Kansas is named after the Kansas River, which in turn was named after the Kansa Native Americans who lived along its banks. The tribe's name (natively ') is often said to mean "people of the (south) wind" although this was probably not the term's original meaning. For thousands of years, what is now Kansas was home to numerous and diverse Native American tribes. Tribes in the eastern part of the state generally lived in villages along the river valleys. Tribes in the western part of the state were semi-nomadic and hunted large herds of bison. The first Euro-American settlement in Kansas occurred in 1827 at Fort Leavenworth. The pace of settlement accelerated in the 1850s, in the midst of political wars over the slavery deba ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessaril ...
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