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Caenagnathinae
Caenagnathidae is a family of bird-like maniraptoran theropod dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and close relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids had specialized beaks, long necks, and short tails, and would have been covered in feathers. The relationships of caenagnathids were long a puzzle. The family was originally named by Raymond Martin Sternberg in 1940 as a family of flightless birds. The discovery of skeletons of the related oviraptorids revealed that they were in fact non-avian theropods, and the discovery of more complete caenagnathid remains revealed that ''Chirostenotes pergracilis'', originally named on the basis of a pair of hands, and ''Citipes elegans'', originally thought to be an ornithomimid, named from a foot, were caenagnathids as well. Anatomy Overall, the anatomy of the caenagnathids is similar to that of the closely related Oviraptoridae, but there are a ...
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Citipes
''Citipes'' (meaning "fleet-footed") is an extinct genus of caenagnathid dinosaurs known from the Late Cretaceous (Campanian aged) Dinosaur Park of southern Alberta, Canada. It lived about 76.9–75.8 million years ago. The specializations of the beak in ''Citipes'' and other caenagnathids suggest that they were herbivores. The type species, ''C. elegans'' had been previously placed within the genera ''Chirostenotes'', ''Elmisaurus'', ''Leptorhynchos'' and ''Ornithomimus''. History The holotype material, found in 1926 at the Little Sandhill Creek, was originally thought to belong to an ornithomimid; William Arthur Parks William Arthur Parks (11 December 1868 – 3 October 1936) was a Canadian geologist and paleontologist, following in the tradition of Lawrence Lambe. Parks was born in Hamilton, Ontario. After graduating with a Bachelor of Arts in 1892, Parks ... in 1933 assigned it to a new species of ''Ornithomimus'': ''O. elegans''.Parks, W.A., (1933), "New species o ...
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Elmisaurus
''Elmisaurus'' (meaning "foot sole lizard") is an extinct genus of caenagnathid dinosaur from the Late Cretaceous Nemegt Formation of Mongolia. It was a theropod belonging to the Oviraptorosauria. Discovery In 1970, a paleontological Polish-Mongolian expedition discovered two fragmentary specimens of a small theropod in the Ömnögovĭ province of Mongolia. The type species, ''Elmisaurus rarus'', was named and described by Halszka Osmólska in 1981. The generic name is derived from Mongol ' or ', "foot sole", as the type specimen consisted of a metatarsus. The specific name means "rare" in Latin. The holotype, ZPAL MgD-I/172, consists of a left metatarsus fused with the tarsalia. There are two paratypes: ZPAL MgD-I/98, consisting of a right hand and foot, and ZPAL MgD-I/20, the upper part of the left metatarsus of a larger individual.Osmólska, H. (1981). Coossified tarsometatarsi in theropod dinosaurs and their bearing on the problem of bird origins. ''Palaeontologica Polo ...
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Leptorhynchos (dinosaur)
''Leptorhynchos'' (meaning "slender beak") is an extinct genus of caenagnathid dinosaurs known from the Late Cretaceous (Campanian-Maastrichtian aged) Aguja Formation of west Texas United States. It lived about 80.5–72 million years ago. It is distinguished from its relatives '' Chirostenotes'' and '' Anzu'' by its smaller size, and by a more strongly upturned mandible, similar to that of oviraptorids. The specializations of the beak in ''Leptorhynchos'' and other caenagnathids suggest that they were herbivores. The species ''L. elegans'' has since been transferred to the genus '' Citipes'', leaving only the type species ''L. gaddisi'' in the genus. ''Leptorhynchos'' has been placed in the Elmisaurinae Caenagnathidae is a family of bird-like maniraptoran theropod dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and close relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids .... See also * Timeline ...
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Hagryphus
''Hagryphus'' (meaning " Ha's griffin") is a monospecific genus of caenagnathid dinosaur from southern Utah that lived during the Late Cretaceous (upper Campanian stage, 75.95 Ma) in what is now the Kaiparowits Formation of the Grand Staircase–Escalante National Monument. The type and only species, ''Hagryphus giganteus'', is known only from an incomplete but articulated left manus and the distal portion of the left radius. It was named in 2005 by Lindsay E. Zanno and Scott D. Sampson. ''Hagryphus'' has an estimated length of 2.4–3 metres (8–10 feet) and weight of 50 kilograms (110 lbs). Discovery and naming To date, only a single species of ''Hagryphus'' has been named in 2005 by Lindsay Zanno and Scott Sampson, the type species ''Hagryphus giganteus''. The generic name is derived from Egyptian ''Ha'', the name of the god of the western desert and a Latinised Greek γρύψ (''gryps'') meaning 'griffin' (a mythological bird-like creature). The specific name means " ...
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Chirostenotes
''Chirostenotes'' ( ; named from Greek 'narrow-handed') is a genus of oviraptorosaurian dinosaur from the late Cretaceous (about 76.5 million years ago) of Alberta, Canada. The type species is ''Chirostenotes pergracilis''. History of discovery ''Chirostenotes'' has a confusing history of discovery and naming. The first fossils of ''Chirostenotes'', a pair of hands, were in 1914 found by George Fryer Sternberg near Little Sandhill Creek in the Campanian Dinosaur Park Formation of Canada, which has yielded the most dinosaurs of any Canadian formation. The specimens were studied by Lawrence Morris Lambe who, however, died before being able to formally name them. In 1924, Charles Whitney Gilmore adopted the name he found in Lambe's notes and described and named the type species ''Chirostenotes pergracilis''. The generic name is derived from Greek ''cheir'', "hand", and ''stenotes'', "narrowness". The specific name means "throughout", ''per~'', "gracile", ''gracilis'', in Lati ...
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Oviraptorid Clean
Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one and two metres long in most cases, though some possible oviraptorids were enormous. Oviraptorids are currently known only from the Late Cretaceous of Asia, with the most well-known species and complete specimens found only in the Gobi Desert of Mongolia and northwestern China. Description The most characteristic feature of this group is the skull structure. Oviraptorids had short snouts and very deep mandibles. Some taxa (such as ''Citipati'', ''Corythoraptor'', ''Rinchenia'') had a midline crest on top of the skull, resembling that of a cassowary. Other distinguishing characteristics include a bony spike intruding on the mandibular fenestra, nostrils placed very high and far back on the snout, an extremely thin bony bar beneath the e ...
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Oviraptoridae
Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one and two metres long in most cases, though some possible oviraptorids were enormous. Oviraptorids are currently known only from the Late Cretaceous of Asia, with the most well-known species and complete specimens found only in the Gobi Desert of Mongolia and northwestern China. Description The most characteristic feature of this group is the skull structure. Oviraptorids had short snouts and very deep mandibles. Some taxa (such as '' Citipati'', '' Corythoraptor'', '' Rinchenia'') had a midline crest on top of the skull, resembling that of a cassowary. Other distinguishing characteristics include a bony spike intruding on the mandibular fenestra, nostrils placed very high and far back on the snout, an extremely thin bony bar beneath t ...
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Oviraptorosauria
Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from ''Caudipteryx'', which was the size of a turkey, to the 8-meter-long, 1.4-ton ''Gigantoraptor''. The group (along with all maniraptoran dinosaurs) is close to the ancestry of birds. Some researchers such as Maryanska ''et al'' (2002) and Osmólska ''et al.'' (2004) have proposed that they may represent primitive flightless birds.Osmólska, Halszka, Currie, Philip J., Brasbold, Rinchen (2004) "The Dinosauria" Weishampel, Dodson, Osmólska. "Chapter 8 Oviraptorosauria" University of California Press. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is sparse.Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropod ...
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Epichirostenotes
''Epichirostenotes'' (meaning "above '' Chirostenotes''", because it lived after the latter genus) is a genus of oviraptorosaurian dinosaur from the late Cretaceous. ''Epichirostenotes'' is known from an incomplete skeleton found in 1923 at the Horseshoe Canyon Formation, in strata dated to about 72 million years ago. It was first named by Robert M. Sullivan, Steven E. Jasinski and Mark P.A. van Tomme in 2011 and the type species is ''Epichirostenotes curriei''. Its holotype, ROM 43250, had been assigned to ''Chirostenotes pergracilis'' by Hans-Dieter Sues Hans-Dieter Sues (born January 13, 1956) is a German-born American paleontologist who is Senior Scientist and Curator of Vertebrate Paleontology at the National Museum of Natural History of the Smithsonian Institution in Washington, DC. He receiv ... in 1997. See also * Timeline of oviraptorosaur research References Caenagnathids Late Cretaceous dinosaurs of North America Fossil taxa described in 2011 Horsesh ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species ...
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Paleognathae
Palaeognathae (; ) is a infraclass of birds, called paleognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant branches of flightless lineages (plus two extinct clades), termed ratites, and one flying lineage, the Neotropic tinamous.Wetmore, A. (1960) A Classification for Birds of the World. Smithsonian Miscellaneous Collection (Washington D.C.: Smithsonian Institution) 139: 1–37. There are 47 species of tinamous, five of kiwis (''Apteryx''), three of cassowaries (''Casuarius''), one of emus (''Dromaius'') (another became extinct in historic times), two of rheas (''Rhea'') and two of ostrich ('' Struthio'').Clements, J. C. ''et al''. (2010) Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radia ...
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Ostrich
Ostriches are large flightless birds of the genus ''Struthio'' in the order Struthioniformes, part of the infra-class Palaeognathae, a diverse group of flightless birds also known as ratites that includes the emus, rheas, and kiwis. There are two living species of ostrich: the common ostrich, native to large areas of sub-Saharan Africa and the Somali ostrich, native to the Horn of Africa. The common ostrich was also historically native to the Arabian Peninsula, and ostriches were present across Asia as far east as Mongolia during the Late Pleistocene and possibly into the Holocene. They lay the largest eggs of any living land animal. With the ability to run at 70 km/h (43.5 mph), they are the fastest birds on land. They are farmed worldwide, particularly for their feathers as they are used as decoration and feather dusters. Their skin is also used for leather products. They are the heaviest living birds. Taxonomic history The genus ''Struthio'' was first des ...
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