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Arthrolycosidae
Arthrolycosidae is an extinct family of arachnids, possibly spiders. Fossils placed in the family were found in the Carboniferous to Permian, . They were considered by Alexander Petrunkevitch to be "mesotheles", i.e. placed in the spider suborder Mesothelae. However, Paul A. Selden has stated they only have "the general appearance of spiders", with segmented abdomens (opisthosomae), but no definite spinnerets. At least some of the specimens placed in the family in the genus ''Arthrolycosa'' are considered to be spiders, whereas ''Eocteniza'' is "rather problematic as a spider". Genera Genera placed in the Arthrolycosidae are shown below, together with the location of the fossils. * †'' Arthrolycosa'' Harger, 1874 – United States ( Mazon Creek), Ukraine, Russia * †'' Eocteniza'' Pocock, 1911 – England (Coseley Coseley ( ) is a village in the north of the Dudley Metropolitan Borough, in the English West Midlands. Part of the Black Country, it is situated approx ...
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Mesothelae
The Mesothelae are a suborder of spiders (order Araneae) that includes a single extant family, Liphistiidae, and a number of extinct families. This suborder is thought to form the sister group to all other living spiders, and to retain ancestral characters, such as a segmented abdomen with spinnerets in the middle and two pairs of book lungs. Members of Liphistiidae are medium to large spiders with eight eyes grouped on a tubercle. They are found only in China, Japan, and southeast Asia. The oldest known Mesothelae spiders are known from the Carboniferous, over 300 million years ago. The Heptathelidae were once considered their own family; today they are considered a subfamily of the Liphistiidae. Description Members of Mesothelae have paraxial chelicerae, two pairs of coxal glands on the legs, eight eyes grouped on a nodule, two pairs of book lungs, and no endites on the base of the pedipalp. Most have at least seven or eight spinnerets near the middle of the abdomen. Latera ...
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Spider
Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species diversity among all orders of organisms. Spiders are found worldwide on every continent except for Antarctica, and have become established in nearly every land habitat. , 50,356 spider species in 132 families have been recorded by taxonomists. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900. Anatomically, spiders (as with all arachnids) differ from other arthropods in that the usual body segments are fused into two tagmata, the cephalothorax or prosoma, and the opisthosoma, or abdomen, and joined by a small, cylindrical pedicel, however, as there is currently neither paleontological nor embryological evidence that spiders ever had ...
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Eocteniza
''Eocteniza'' is an extinct genus of arachnids containing the sole species ''Eocteniza silvicola'', known from the Westphalian stage of the Carboniferous period in Coseley, England,, retrieved 21 March 2016 about . It was initially identified as a spider, but this is now doubted. Description The carapace (upper surface of the cephalothorax) is about 5 mm long by 4 mm wide. The abdomen is also about 5 mm long, slightly narrower at about 3.5 mm wide. The front end of the carapace is narrowed and raised, separated from the rest of the cephalothorax by shallow grooves, and bearing the eyes at the front. Pocock's description and drawing show in addition to the grooves separating the head from the rest of the thorax, three further pairs of radiating grooves, with the fovea towards the rear. This is different from mesothele spiders, whose fovea consists of a deep pit, with a transverse groove and deeper longitudinal groove. Taxonomy The genus and species were fi ...
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Arachnid
Arachnida () is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons. Almost all adult arachnids have eight legs, although the front pair of legs in some species has converted to a sensory function, while in other species, different appendages can grow large enough to take on the appearance of extra pairs of legs. The term is derived from the Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, who was turned into a spider. Almost all extant arachnids are terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 100,000 named species, of which 47,000 are species of spiders. Morphology Almost all adult arachnids have eight legs, unlike adult i ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian lin ...
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Permian
The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the Perm Governorate, region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the Sauropsida, sauropsids (reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditi ...
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Alexander Petrunkevitch
Alexander Ivanovitch Petrunkevitch ( Russian: Александр Иванович Петрункевич, December 22, 1875 in Plysky near Kyiv, now Ukraine – March 9, 1964 in New Haven) was an eminent Russian arachnologist of his time. From 1910 to 1939 he described over 130 spider species. One of his most famous essays was "The Spider and the Wasp." In it he uses effective word choices and some comic touch. Biography His aristocratic father, Ivan Illitch Petrunkevitch, was a liberal member of the First Duma and founded the Constitutional Democratic Party. After finishing his studies in Moscow and in Freiburg under August Weismann, Alexander settled in Yale in 1910, becoming a full professor in 1917. Apart from describing present-day species, he was a major figure in the study of fossil arachnids, including those in amber and from the Coal Measures. He also experimented with live specimens and worked on insects. Professor Petrunkevitch's formulation of the prin ...
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Paul Anthony Selden
Paul may refer to: *Paul (given name), a given name (includes a list of people with that name) *Paul (surname), a list of people People Christianity * Paul the Apostle (AD c.5–c.64/65), also known as Saul of Tarsus or Saint Paul, early Christian missionary and writer *Pope Paul (other), multiple Popes of the Roman Catholic Church *Saint Paul (other), multiple other people and locations named "Saint Paul" Roman and Byzantine empire *Lucius Aemilius Paullus Macedonicus (c. 229 BC – 160 BC), Roman general *Julius Paulus Prudentissimus (), Roman jurist *Paulus Catena (died 362), Roman notary *Paulus Alexandrinus (4th century), Hellenistic astrologer *Paul of Aegina or Paulus Aegineta (625–690), Greek surgeon Royals * Paul I of Russia (1754–1801), Tsar of Russia *Paul of Greece (1901–1964), King of Greece Other people * Paul the Deacon or Paulus Diaconus (c. 720 – c. 799), Italian Benedictine monk *Paul (father of Maurice), the father of Maurice, By ...
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Opisthosoma
The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma ( cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects to an abdomen (and is often referred to as such), the opisthosoma is differentiated by its inclusion of the respiratory organs ( book lungs or book gills) and the heart. Segments The number of segments and appendages on the opisthosoma vary. Scorpions have 13, but the first is only seen during its embryological development. Other arachnids have fewer; harvestmen, for instance, have only ten. In general, appendages are absent or reduced, although in horseshoe crabs they persist as large plate-like limbs, called opercula or branchiophores, bearing the book gills, and that function in locomotion and gas exchange. In most chelicerates the opisthosomal limbs are greatly reduced and persist only as specialized structures, such as the silk-produ ...
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Arthrolycosa
''Arthrolycosa'' (meaning '' wolf piderwith joints'') is an extinct genus of arachnids, possibly spiders, that lived about 300-250 million years ago. Fossils have been found Mazon Creek USA and in the Kirov Oblast region of Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eig .... ''A. antiqua'' is estimated to have a body length of about 2.17 cm and may have preyed upon insects and other smaller animals that lived alongside them. External links & References * Parker, Steve. Dinosaurus: the complete guide to dinosaurs. Firefly Books Inc, 2003. Pg. 75 Prehistoric arachnid genera Paleozoic arachnids Fossils of the United States Fossils of Russia Pennsylvanian first appearances Lopingian genus extinctions Animals described in 1874 Fossil taxa described in 1874 ...
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Mazon Creek Fossil Beds
The Mazon Creek fossil beds are a conservation ' found near Morris, in Grundy County, Illinois. The fossils are preserved in ironstone concretions, formed approximately in the mid- Pennsylvanian epoch of the Carboniferous period. These concretions frequently preserve both hard and soft tissues of animal and plant materials, as well as many soft-bodied organisms that do not normally fossilize. The quality, quantity and diversity of fossils in the area, known since the mid-nineteenth century, make the Mazon Creek ' important to paleontologists, in attempting to reconstruct the paleoecology of the sites. The locality was declared a National Historic Landmark in 1997..   Geology The Mazon Creek fossils are found in the Upper Carboniferous Francis Creek Shale; the type locality is the Mazon River (or Mazon Creek), a tributary of the Illinois River near Morris, Grundy County, Illinois. The 25 to 30 meters of shale were formed approximately , during the Pennsylvanian period ...
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