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The ruddy shelduck (''Tadorna ferruginea''), known in India as the Brahminy duck, is a bird
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Anatidae The Anatidae are the biological family (biology), family of water birds that includes ducks, goose, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted f ...
. It is a distinctive waterfowl, in length with a wingspan of . It has orange-brown body plumage with a paler head, while the tail and the flight feathers in the wings are black, contrasting with the white wing-coverts. It is a
migratory bird Bird migration is a seasonal movement of birds between breeding and wintering grounds that occurs twice a year. It is typically from north to south or from south to north. Migration is inherently risky, due to predation and mortality. Th ...
, wintering in the Indian subcontinent and breeding in southeastern Europe and central Asia, though there are small resident populations in North Africa. It has a loud honking call. The ruddy shelduck mostly inhabits inland water-bodies such as lakes, reservoirs and rivers. The male and female form a lasting pair bond and the nest may be well away from water, in a crevice or hole in a cliff, tree or similar site. A clutch of about eight eggs is laid and is incubated solely by the female for about four weeks. The young are cared for by both parents and
fledge Fledging is the stage in a flying animal's life between egg, hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnera ...
about eight weeks after hatching. In central and eastern Asia, populations are steady or rising, but in Europe they are generally in decline. Altogether, the birds have a wide range and large total population, and the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
has assessed their conservation status as being of
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
.


Taxonomy and etymology

The ruddy shelduck (''Tadorna ferruginea'') is a member of the shelduck
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Tadorna The shelducks, most species of which are found in the genus ''Tadorna'' (except for the Radjah shelduck, which is now found in its own monotypic genus ''Radjah''), are a group of large birds in the Tadorninae subfamily of the Anatidae, the biolog ...
'' in the wildfowl family Anatidae. The bird was first described in 1764 by the German zoologist and botanist
Peter Simon Pallas Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussia, Prussian zoologist, botanist, Ethnography, ethnographer, Exploration, explorer, Geography, geographer, Geology, geologist, Natura ...
who named it ''Anas ferruginea'', but later it was transferred to the genus ''Tadorna'' with the other shelducks. Some authorities place it in the genus '' Casarca'' along with the
South African shelduck The South African shelduck or Cape shelduck (''Tadorna cana'') is a species of shelduck, a group of large goose-like birds which are part of the bird family Anatidae, which also includes the swans, goose, geese and ducks. This is a common specie ...
(''T. cana''), the
Australian shelduck The Australian shelduck (''Tadorna tadornoides''), also known as the chestnut-breasted shelduck or mountain duck, is a shelduck, a group of large goose-like ducks part of the bird family Anatidae. The genus name ''Tadorna'' comes from Celtic ...
(''T. tadornoides'') and the
paradise shelduck The paradise shelduck (''Tadorna variegata''), also known as the paradise duck, or in Māori, is a species of shelduck, a group of goose-like ducks, which is endemic to New Zealand. Johann Friedrich Gmelin placed it in the genus '' Anas'' w ...
(''T. variegata'').
Phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analysis shows that it is most closely related to the South African shelduck. In captivity, the ruddy shelduck has been known to hybridise with several other members of ''Tadorna'', with several members of the dabbling duck genus ''
Anas ''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
'', and with the
Egyptian goose The Egyptian goose (''Alopochen aegyptiaca'') is an African member of the Anatidae family including ducks, geese, and swans. Because of their popularity chiefly as an ornamental bird, the species has also been introduced to Europe, the United ...
(''Alopochen aegyptiaca''). No
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised. The
generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
name ''Tadorna'' comes from the French , the
common shelduck The common shelduck (''Tadorna tadorna'') is a waterfowl species of the shelduck genus, ''shelduck, Tadorna''. It is widespread and common in the Euro-Siberian region of the Palearctic realm, Palearctic, mainly breeding in temperate and wintering ...
, and may originally derive from a Celtic word meaning "pied waterfowl". The
English English usually refers to: * English language * English people English may also refer to: Culture, language and peoples * ''English'', an adjective for something of, from, or related to England * ''English'', an Amish ter ...
name "sheld duck" dates from around 1700 and means the same. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''ferruginea'' is Latin for "rusty" and refers to the colour of the plumage.


Description

The ruddy shelduck grows to a length of and has a wingspan. The male has orange-brown body plumage and a paler, orange-brown head and neck, separated from the body by a narrow black collar. The rump, flight feathers, tail-coverts and tail feathers are black and there are iridescent green
speculum feathers The speculum is a patch, often distinctly coloured, on the secondary wing feathers, or ''remiges'', of some birds. Examples of the colour(s) of the speculum in a number of ducks are: * Common teal and green-winged teal: Iridescent green ed ...
on the inner surfaces of the wings. Both upper and lower wing-coverts are white, this feature being particularly noticeable in flight but hardly visible when the bird is at rest. The bill is black and the legs are dark grey. The female is similar but has a rather pale, whitish head and neck and lacks the black collar, and in both sexes, the colouring is variable and fades as the feathers age. The birds
moult In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is a process by which an animal casts off parts of its body to serve some beneficial purpose, either at ...
at the end of the breeding season and the male loses the black collar, but a further partial moult between December and April restores it. Juveniles are similar to the female but are a darker shade of brown. The call is a series of loud, nasal honking notes, it being possible to discern the difference between those produced by the male and the female. The calls are made both on the ground and in the air, and the sounds are variable according to the circumstances in which they are uttered.


Distribution and habitat

There are very small resident populations of this species in Northwest
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
and
Ethiopia Ethiopia, officially the Federal Democratic Republic of Ethiopia, is a landlocked country located in the Horn of Africa region of East Africa. It shares borders with Eritrea to the north, Djibouti to the northeast, Somalia to the east, Ken ...
, but the main breeding area of the bird is from southeast Europe across the
Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...
to
Lake Baikal Lake Baikal is a rift lake and the deepest lake in the world. It is situated in southern Siberia, Russia between the Federal subjects of Russia, federal subjects of Irkutsk Oblast, Irkutsk Oblasts of Russia, Oblast to the northwest and the Repu ...
, Mongolia, and western China. Eastern populations are mostly migratory, wintering in the
Indian subcontinent The Indian subcontinent is a physiographic region of Asia below the Himalayas which projects into the Indian Ocean between the Bay of Bengal to the east and the Arabian Sea to the west. It is now divided between Bangladesh, India, and Pakista ...
. This species has colonised the island of
Fuerteventura Fuerteventura () is one of the Canary Islands, in the Atlantic Ocean, geographically part of Macaronesia, and politically part of Spain. It is located away from the coast of North Africa. The island was declared a biosphere reserve by UNESCO i ...
in the
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
, first breeding there in 1994, and reaching a population of almost fifty pairs by 2008. The ruddy shelduck is a common winter visitor in India where it arrives by October and departs by April. Its typical breeding habitat is large wetlands and rivers with
mud flats Mudflats or mud flats, also known as tidal flats or, in Ireland, slob or slobs, are coastal wetlands that form in intertidal areas where sediments have been deposited by tides or rivers. A global analysis published in 2019 suggested that tidal f ...
and shingle banks, and it is found in large numbers on lakes and reservoirs. It breeds in high altitude lakes and swamps in
Jammu and Kashmir Jammu and Kashmir may refer to: * Jammu and Kashmir (union territory), a region administered by India as a union territory since 2019 * Jammu and Kashmir (state), a region administered by India as a state from 1952 to 2019 * Jammu and Kashmir (prin ...
. Outside the breeding season it prefers lowland streams, sluggish rivers, ponds, flooded grassland,
marsh In ecology, a marsh is a wetland that is dominated by herbaceous plants rather than by woody plants.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p More in genera ...
es and
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
lagoons. Although becoming quite rare in Southeastern
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
and southern
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
, the ruddy shelduck is still common across much of its Asian range. It may be this population which gives rise to vagrants as far west as
Iceland Iceland is a Nordic countries, Nordic island country between the Atlantic Ocean, North Atlantic and Arctic Oceans, on the Mid-Atlantic Ridge between North America and Europe. It is culturally and politically linked with Europe and is the regi ...
,
Great Britain Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...
and
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
. However, since the European population is declining, it is likely that most occurrences in Western Europe in recent decades are escapes or
feral A feral (; ) animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in som ...
birds. Although this bird is observed in the wild from time to time in eastern
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
, no evidence has been found that this is a genuine case of
vagrancy Vagrancy is the condition of wandering homelessness without regular employment or income. Vagrants usually live in poverty and support themselves by travelling while engaging in begging, waste picker, scavenging, or petty theft. In Western ...
. Feral ruddy shelduck have bred successfully in several European countries. In
Switzerland Switzerland, officially the Swiss Confederation, is a landlocked country located in west-central Europe. It is bordered by Italy to the south, France to the west, Germany to the north, and Austria and Liechtenstein to the east. Switzerland ...
the ruddy shelduck is considered an invasive species that threatens to displace native birds. Despite actions taken to reduce numbers, the population of ruddy shelduck in Switzerland increased from 211 to 1250 individuals in the period from 2006 to 2016. A stable population exists in Moscow, settling the city parks' ponds alongside the endemic
mallard The mallard () or wild duck (''Anas platyrhynchos'') is a dabbling duck that breeds throughout the temperate and subtropical Americas, Eurasia, and North Africa. It has been introduced to New Zealand, Australia, Peru, Brazil, Uruguay, Arge ...
s. It is theorized that the birds are feral descendants of escapees from the Moscow Zoo, the population most likely forming after 1948, when the policy of clipping the birds' wings was repealed. Unlike the wild population, these ducks are non-migratory, wintering instead in the non-freezing parts of the city's bodies of water. Population increased from 1990's to 2020's by over 20 times, reaching 3300 by 2025. This shelduck mostly frequents open locations on inland bodies of water such as lakes, reservoirs and rivers. It is seldom seen in forested areas but does occur in brackish water and saline lagoons. Though more common in the lowlands, it also inhabits higher altitudes and in central Asia is one of the few waterbirds, along with the
bar-headed goose The bar-headed goose (''Anser indicus'') is a goose that breeds in Central Asia in colonies of thousands near mountain lakes and winters in South Asia, as far south as peninsular India. It lays three to eight eggs at a time in a ground nest. It ...
(''Anser indicus''), to be found on lakes at .


Behaviour

The ruddy shelduck is a mainly
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
bird. It is
omnivorous An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize ...
and feeds on grasses, the young shoots of plants, grain and water plants as well as both aquatic and terrestrial
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s. On land it grazes on the foliage, in the water it dabbles in the shallows, and at greater depths, it up-ends, but it does not dive. The ruddy shelduck is usually found in pairs or small groups and rarely forms large flocks. However, moulting and wintering gatherings on chosen lakes or slow rivers can be very large. Gatherings of over four thousand birds have been recorded on the
Koshi Barrage The Koshi Barrage is a sluice across the Koshi river in the Nepalese Mithila region that carries vehicular, bicycle, and pedestrian traffic between Madhesh Province and Koshi Province of Nepal. It is near the International border with India. It ...
and in the
Koshi Tappu Wildlife Reserve The Koshi Tappu Wildlife Reserve is a protected area in the Terai of eastern Nepal covering of wetlands in the Sunsari District, Sunsari, Saptari District, Saptari and Udayapur Districts. It comprises extensive reed beds and freshwater marshes i ...
in Nepal, and over ten thousand at
Lake Düden Lake Düden, also known as Lake Kulu, ( or ''Kulu Gölü''), is a brackish water lake in Konya Province, Turkey. Lake Düden is located northwest of Lake Tuz and east of Kulu town in Konya Province at an elevation of . It is a shallow brackish ...
in Turkey. The birds arrive at their main breeding locations in central Asia in March and April. There is a strong pair bond between the male and female and it is thought they pair for life. In their breeding quarters, the birds are very aggressive towards their own kind and towards other species. The female in particular approaches intruders with head lowered and neck outstretched, uttering anger calls. If the intruder stands its ground, the female returns to the male and runs round him, inciting him to attack. He may or may not do so. Mating takes place on the water after a brief courtship ritual involving neck stretching, head dipping and tail raising. The nesting site is often far away from water in a hole in a tree or ruined building, a crevice in a cliff, among sand-dunes or in an animal burrow. The nest is constructed by the female using feathers and down and some grasses. A clutch of about eight eggs (range six to twelve) is laid between late April and early June. These have a dull gloss and are creamy-white, averaging . Incubation is done by the female while the male stands in attendance nearby. The eggs hatch after about twenty-eight days and both parents care for the young, which
fledge Fledging is the stage in a flying animal's life between egg, hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnera ...
in a further fifty-five days. After breeding the adults
moult In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is a process by which an animal casts off parts of its body to serve some beneficial purpose, either at ...
, losing the power of flight for about a month while they do so. Before moulting they move to large water bodies where they can more easily avoid
predation Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
while they are flightless. The family may stay together as a group for some time; the autumn migration starts around September and the young may mature in their second year. North African birds breed about five weeks earlier, and their breeding success is greater in wet summers.


Status

Buddhists Buddhism, also known as Buddhadharma and Dharmavinaya, is an Indian religion and philosophical tradition based on teachings attributed to the Buddha, a wandering teacher who lived in the 6th or 5th century BCE. It is the world's fourth ...
regard the ruddy shelduck as sacred and this gives the birds some protection in central and eastern Asia, where the population is thought to be steady or even rising. The Pembo Black-necked Crane Reserve in Tibet is an important wintering area for ruddy shelducks, and here they receive protection. In Europe on the other hand, populations are generally declining as wetlands are drained and the birds are hunted. However, in Europe and Italy the ruddy shelduck is protected. You can not hunt it. They are less vulnerable than some other waterfowl because of their adaptability to new habitats such as reservoirs. In 2023, the Ruddy Shelduck was selected as the
flagship species In conservation biology, a flagship species is a species chosen to raise support for biodiversity conservation in a given place or social context. Definitions have varied, but they have tended to focus on the strategic goals and the socio-economi ...
of Arjan International Wetland in Iran, due to its cultural visibility and conservation potential in engaging local communities. The ruddy shelduck has a very wide range and an estimated total population size of 170,000 to 225,000 individuals. The overall population trend is unclear as some local populations are increasing while others are decreasing. The bird does not appear to meet the higher criteria necessary to be considered threatened, and the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
assesses that its conservation status is of
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
. It is one of the species to which the
Agreement on the Conservation of African-Eurasian Migratory Waterbirds The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA), is an independent international treaty developed under the auspices of the United Nations Environment Programme's Convent ...
(AEWA) applies.


References


External links

* * * * * * {{Authority control
ruddy shelduck The ruddy shelduck (''Tadorna ferruginea''), known in India as the Brahminy duck, is a bird species in the family Anatidae. It is a distinctive waterfowl, in length with a wingspan of . It has orange-brown body plumage with a paler head, whil ...
Birds of North Africa Birds of Central Asia Birds of West Asia Birds of Mongolia Birds of Tibet
ruddy shelduck The ruddy shelduck (''Tadorna ferruginea''), known in India as the Brahminy duck, is a bird species in the family Anatidae. It is a distinctive waterfowl, in length with a wingspan of . It has orange-brown body plumage with a paler head, whil ...
Taxa named by Peter Simon Pallas