Gekkonidae - Phelsuma Klemmeri
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Gekkonidae (the common geckos) is the largest family of
gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates. They range from . Geckos are unique among lizards ...
s, containing over 950 described
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
in 62
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
. The Gekkonidae contain many of the most widespread gecko species, including house geckos (''
Hemidactylus ''Hemidactylus'' is a genus of the common gecko family, Gekkonidae. It has 194 described species, newfound ones being described every few years. These geckos are found in all the tropical regions of the world, extending into the subtropical parts ...
''), the tokay gecko (''
Gekko ''Gekko'' is a genus of Southeast Asian geckos, Common name, commonly known as true geckos or calling geckos, in the Family (biology), family Gekkonidae. Although species such as ''Gekko gecko'' (tokay gecko) are very widespread and common, some ...
''), day geckos (''
Phelsuma ''Phelsuma'' is a large genus of geckos in the Family (biology), family Gekkonidae. Species in the genus ''Phelsuma'' are commonly referred to as day geckos. Some day geckos are seriously endangered and some are common, but all ''Phelsuma'' spec ...
''), the
mourning gecko ''Lepidodactylus lugubris'', known as the mourning gecko or common smooth-scaled gecko, is a species of lizard, a gecko of the family Gekkonidae. Description ''Lepidodactylus lugubris'' measure 8.5–10 cm in length including tail (4–4. ...
(''
Lepidodactylus ''Lepidodactylus'' is a large genus of geckos, commonly known as scaly-toed geckos and closely related to house geckos or Gehyra, dtellas. Geographic range Species in the genus ''Lepidodactylus'' are found from Southeast Asia to Indo-Australi ...
''), and dtellas (''
Gehyra ''Gehyra'' is a genus of geckos, lizards in the family ''Gekkonidae''. Species within the genus are known as web-toed geckos or dtellas, and most bear close resemblance to geckos from the genus ''Hemidactylus''. Geographic range ''Gehyra'' spe ...
''). Gekkonid geckos occur globally and are particularly diverse in tropical areas. Many species of these geckos exhibit an adhering ability to surfaces through
Van der Waals forces In molecular physics and chemistry, the van der Waals force (sometimes van der Waals' force) is a distance-dependent interaction between atoms or molecules. Unlike ionic or covalent bonds, these attractions do not result from a chemical ele ...
utilizing intermolecular forces between molecules of their setae (foot hair) and molecules of the surface they are on. The genus ''Hemidactylus'' is one of the most species-rich and widely distributed of all reptile genera.Carranza, S., and E. Arnold. "Systematics, Biogeography, and Evolution of Hemidactylus Geckos (Reptilia: Gekkonidae) Elucidated Using Mitochondrial DNA Sequences." Molecular Phylogenetics and Evolution, vol. 38, no. 2, Elsevier Inc, 2006, pp. 531–45, .


Fossils

The family Gekkonidae is a member of the
infraorder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classific ...
Gekkota Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates. They range from . Geckos are unique among lizards f ...
, which seems to have first emerged during the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
period (201–145 million years ago). ''
Eichstaettisaurus ''Eichstaettisaurus'' (meaning "Eichstätt lizard") is a genus of lizards from the Late Jurassic and Early Cretaceous of Germany, Spain, and Italy. With a flattened head, forward-oriented and partially symmetrical feet, and tall claws, ''Eichstae ...
schroederi'' is recognized as one of the earliest examples of an ancestral gecko species. Members of the genus ''Eichstaettisaurus'' display morphological adaptations associated with climbing. ''Eichstaettisaurus'' has been ranked as a stem gekkotan, but its true taxonomic rank is unclear. The species ''
Hoburogekko ''Hoburogekko'' is an extinct genus of stem-group gecko that includes a single species, ''Hoburogekko suchanovi'', from the Early Cretaceous Dzunbain Formation of Mongolia.Alifanov, V.R. (1989) The oldest gecko (Lacertilia, Gekkonidae) from the L ...
suchanovi'' and ''
Gobekko ''Gobekko'' is an extinct genus of gecko or gecko-like lizard from the Late Cretaceous Djadokhta Formation of the Gobi Desert in Mongolia. ''Gobekko'' is either a basal member of Gekkota, the group that includes geckos and the legless pygopodid ...
cretacicus,'' dated to the
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
-
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
ages (121–100 million years ago) ages of the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
period, are unquestionably members of Gekkota. They are thought to be members of the Gekkonidae, as well, but their taxonomic rank is uncertain due to the incompleteness of the fossil remains. ''
Yantarogekko ''Yantarogekko'' is an extinct genus of gecko known from a single specimen found in Baltic amber from the Eocene Prussian Formation of Kaliningrad, Russia. The remains consist of the anterior half of a body with partially preserved limbs (includ ...
balticus'' is the earliest known gekkonid gecko. ''Y. balticus'' was discovered in
Baltic amber Baltic amber or succinite is amber from the Baltic region, home of its largest known deposits. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that this forested region provided the re ...
dated to the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
epoch (56–33.9 million years ago). This species was small, measuring only 20–22 mm from snout to vent. ''Y. balticus'' has enlarged, undivided
scansorial Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The habi ...
pads and a reduced but strongly clawed first digit, and lacks movable
eyelid An eyelid ( ) is a thin fold of skin that covers and protects an eye. The levator palpebrae superioris muscle retracts the eyelid, exposing the cornea to the outside, giving vision. This can be either voluntarily or involuntarily. "Palpebral ...
s. Together, these morphological traits are distinctive of the family Gekkonidae, and also indicate that ''Y. balticus'' may have had adhesive abilities similar to modern geckos.


Distribution

Species within the Gekkonidae inhabit every warm region. Furthermore, many genera are capable of widespread geographical habitation, and can be considered invasive in some areas; the genus ''Hemidactylus'' can be identified in all subtropical areas of the world. However, many genera in the family Gekkonidae are endemic. For example, genera ''
Afroedura ''Afroedura'' is a genus of African geckos, lizards in the family Gekkonidae. Member species are collectively known as rock geckos or flat geckos. In 2021, four new species of ''Afroedura'' were described from Angola. Species The genus ''Afroe ...
'' and ''
Afrogecko The marbled leaf-toed gecko (''Afrogecko porphyreus'') is a gecko found in southern and southwestern South Africa (including many offshore islands) and in Namibia. It is a flat, medium-sized gecko. Description ''Afrogecko porphyreus'' has a m ...
'' are found only in
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
. The range of genus ''Lepidodactylus'' stretches from South Asia to Oceania; this genus also accounts for several island species. The genera listed here do not account for all Gekkonidae subspecies and are used only as examples of distribution. For a full list, see section 'Genera'.


Genera

Gekkonidae contains these genera: Possibly belonging to the family: ''
Yantarogekko ''Yantarogekko'' is an extinct genus of gecko known from a single specimen found in Baltic amber from the Eocene Prussian Formation of Kaliningrad, Russia. The remains consist of the anterior half of a body with partially preserved limbs (includ ...
'' (fossil)


Phylogeny

Pyron, et al. (2013)Pyron, R Alexander, Frank T Burbrink and John J Wiens. 2013. A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evolutionary Biology 2013 13:93. presents the following classification of ''Gekkonidae'' genera, based on
molecular phylogenetics Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
.


References

{{Authority control Lizard families Taxa named by John Edward Gray Geckos Extant Eocene first appearances bcl:Tukô