''Yongjinglong'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

titanosauriform Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located (hence ...

sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...

known from the

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

of Lanzhou-Minhe Basin of

Gansu Province Gansu is a provinces of China, province in Northwestern China. Its capital and largest city is Lanzhou, in the southeastern part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibetan Plateau, Ti ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

. It contains a single species, ''Yongjinglong datangi''.

Discovery

''Yongjinglong'' was first described and named by Li-Guo Li, Da-Qing Li, Hai-Lu You and

Peter Dodson Peter Dodson (born August 20, 1946) is an American paleontologist who has published many papers and written and collaborated on books about dinosaurs. An authority on Ceratopsians, he has also authored several papers and textbooks on hadrosaurs a ...

2014 The year 2014 was marked by the surge of the Western African Ebola epidemic, West African Ebola epidemic, which began in 2013, becoming the List of Ebola outbreaks, most widespread outbreak of the Ebola, Ebola virus in human history, resul ...

and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Yongjinglong datangi''. The generic name is derived from the name of the historical

Yongjing County Yongjing (, Xiao'erjing: ) is a county in Linxia Hui Autonomous Prefecture in China's Gansu Province. The county seat, the town (''zhen'') of Liujiaxia (), is located about 80 km south-west from the provincial capital city, Lanzhou. As is ...

, near where the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

of ''Yongjinglong'' and numerous dinosaur track fossils were collected, and from ''long'', meaning "

dragon A dragon is a Magic (supernatural), magical legendary creature that appears in the folklore of multiple cultures worldwide. Beliefs about dragons vary considerably through regions, but European dragon, dragons in Western cultures since the Hi ...

" in

Chinese Chinese may refer to: * Something related to China * Chinese people, people identified with China, through nationality, citizenship, and/or ethnicity **Han Chinese, East Asian ethnic group native to China. **'' Zhonghua minzu'', the supra-ethnic ...

. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''datangi'', honors the

Tang dynasty The Tang dynasty (, ; zh, c=唐朝), or the Tang Empire, was an Dynasties of China, imperial dynasty of China that ruled from 618 to 907, with an Wu Zhou, interregnum between 690 and 705. It was preceded by the Sui dynasty and followed ...

and also Mr. Zhi-Lu Tang from the

IVPP The Institute of Vertebrate Paleontology and Paleoanthropology (IVPP; ) of China is a research institution and collections repository for fossils, including many dinosaur and pterosaur specimens (many from the Yixian Formation). As its name sugges ...

, for his contributions to the study of

s. ''Yongjinglong'' is known solely from the

GSGM ZH(08)-04, a partial

postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...

and three

teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

, currently housed at the

Gansu Geological Museum Gansu is a province in Northwestern China. Its capital and largest city is Lanzhou, in the southeastern part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibetan and Loess plateaus and borders ...

. The postcranial remains include one fragmentary dorsal

rib In vertebrate anatomy, ribs () are the long curved bones which form the rib cage, part of the axial skeleton. In most tetrapods, ribs surround the thoracic cavity, enabling the lungs to expand and thus facilitate breathing by expanding the ...

, the left

scapulocoracoid The scapulocoracoid is the unit of the pectoral girdle that contains the coracoid and scapula. The coracoid itself is a beak-shaped bone that is commonly found in most vertebrates with a few exceptions. The scapula is commonly known as the ''shoulde ...

, the right

ulna The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...

and

radius In classical geometry, a radius (: radii or radiuses) of a circle or sphere is any of the line segments from its Centre (geometry), center to its perimeter, and in more modern usage, it is also their length. The radius of a regular polygon is th ...

, as well as eight presacral

vertebrae Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...

including one caudal

cervical vertebra In tetrapods, cervical vertebrae (: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In sauropsid s ...

, four cranial

dorsal vertebrae In vertebrates, thoracic vertebrae compose the middle segment of the vertebral column, between the cervical vertebrae and the lumbar vertebrae. In humans, there are twelve thoracic vertebra (anatomy), vertebrae of intermediate size between the ce ...

, and three articulated middle dorsal vertebrae. It represents a subadult individual. GSGM ZH(08)-04 was by Li Daqing and You Hailu discovered in 2008 beside the G75 Highway, less than a

kilometer The kilometre ( SI symbol: km; or ), spelt kilometer in American and Philippine English, is a unit of length in the International System of Units (SI), equal to one thousand metres (kilo- being the SI prefix for ). It is the preferred mea ...

from the quarries of ''

Daxiatitan ''Daxiatitan'' is a genus of sauropod dinosaur known from the Lower Cretaceous of Gansu, China. Its type and only species is ''Daxiatitan binglingi''. It is known from a single partial skeleton consisting of most of the neck and back vertebrae, ...

'' and '' Huanghetitan liujiaxiaensis'', near

Zhongpu Zhongpu Township or Jhongpu Township () is a rural township in Chiayi County, Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', li ...

. It was collected from the upper

Hekou Group The Hekou Group is a geological group in Gansu Province, China. It is Early Cretaceous in age. Many dinosaur fossils have been recovered from the Hekou Group, including iguanodonts, large sauropods, and Thyreophora, armored dinosaurs. Fossil eggs ...

, in the southeastern part of the Lanzhou-Minhe Basin,

, dating to the

Description

''Yongjinglong'' was a medium-sized sauropod. The describers established some diagnostic traits. The premaxillary teeth are long and spoon-shaped. The neck vertebrae and anterior dorsal vertebrae possess large and deep pleurocoels, pneumatic depressions, that cover the entire sides of the centra. The ridges on the sides of the middle dorsal vertebrae form a pattern shaped like a "XI", with the "X" in front. The spine of at least one middle dorsal vertebra is low and not bifurcated and forms with the postzygapophyses a triangular bone plate, directed upwards. The shoulder blade is extremely long, with, in side view, exceptionally straight front and rear edges.

Phylogeny

''Yongjinglong'' was placed in the

Titanosauria Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of Sauropoda, sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with tax ...

by Li ''et al.'' (2014). Their

cladistic Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...

analysis showed it was a member of the

Somphospondyli Somphospondyli is an extinct clade of titanosauriformes, titanosauriform sauropods that lived from the Late Jurassic until the end of the Late Cretaceous, comprising all titanosauriforms more closely related to Titanosauria proper than Brachiosau ...

and a

sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

of ''

Opisthocoelicaudia ''Opisthocoelicaudia'' is a genus of sauropod dinosaur of the Late Cretaceous Period discovered in the Gobi Desert of Mongolia. The type species is ''Opisthocoelicaudia skarzynskii''. A well-preserved skeleton lacking only the head and neck wa ...

''. However, a 2019 revision of Chinese "titanosaurs" resolved ''Yongjinglong'' deeply nested within

Euhelopodidae Euhelopodidae is a family of sauropod dinosaurs of disputed membership and affinities, which contains '' Euhelopus'' and its close relatives. Most proposed euhelopodids are from East Asia. Euhelopodidae was first recognized by Carl Wiman in 1929 ...

, sister taxon to ''

Huanghetitan ''Huanghetitan'' (meaning "Yellow River titan"), is a genus of sauropod dinosaur from the early Cretaceous Period. It was a basal titanosauriformes, titanosauriform which lived in what is now Gansu, China. History The type species, ''Huangh ...

'' and '' Ruyangosaurus'', or a clade including ''

Euhelopus ''Euhelopus'' is a genus of sauropod dinosaur that lived between 145 and 133 million years ago during the Berriasian and Valanginian ages of the Early Cretaceous in what is now Shandong Province in China. It was a large quadrupedal herbivore. Lik ...

'', '' Erketu'' and ''

Gobititan ''Gobititan'' is a genus of herbivorous sauropod dinosaur from the Aptian faunal stage of the Early Cretaceous. The name of this genus is derived from the Gobi Desert region and the Titans of Greek mythology, which is a reference to its large bo ...

''.

References

{{Taxonbar, from=Q15690935 Titanosauria Dinosaur genera Albian dinosaurs Dinosaurs of China Fossil taxa described in 2014 Taxa named by Hailu You Taxa named by Peter Dodson