''Yongjinglong'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of titanosauriform

sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

known from the

Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pro ...

of Lanzhou-Minhe Basin of

Gansu Province Gansu (, ; alternately romanized as Kansu) is a province in Northwest China. Its capital and largest city is Lanzhou, in the southeast part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibeta ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

. It contains a single species, ''Yongjinglong datangi''.

Discovery

''Yongjinglong'' was first described and named by Li-Guo Li, Da-Qing Li, Hai-Lu You and Peter Dodson in

2014 File:2014 Events Collage.png, From top left, clockwise: Stocking up supplies and personal protective equipment (PPE) for the Western African Ebola virus epidemic; Citizens examining the ruins after the Chibok schoolgirls kidnapping; Bundles of wat ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Yongjinglong datangi''. The generic name is derived from the name of the historical Yongjing County, near where the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

of ''Yongjinglong'' and numerous dinosaur track fossils were collected, and from ''long'', meaning "

dragon A dragon is a reptilian legendary creature that appears in the folklore of many cultures worldwide. Beliefs about dragons vary considerably through regions, but dragons in western cultures since the High Middle Ages have often been depicted as ...

" in

Chinese Chinese can refer to: * Something related to China * Chinese people, people of Chinese nationality, citizenship, and/or ethnicity **''Zhonghua minzu'', the supra-ethnic concept of the Chinese nation ** List of ethnic groups in China, people of va ...

. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''datangi'', honors the

Tang dynasty The Tang dynasty (, ; zh, t= ), or Tang Empire, was an Dynasties in Chinese history, imperial dynasty of China that ruled from 618 to 907 AD, with an Zhou dynasty (690–705), interregnum between 690 and 705. It was preceded by the Sui dyn ...

and also Mr. Zhi-Lu Tang from the

IVPP The Institute of Vertebrate Paleontology and Paleoanthropology (IVPP; ) of China is a research institution and collections repository for fossils, including many dinosaur and pterosaur specimens (many from the Yixian Formation). As its name suggest ...

, for his contributions to the study of

s. ''Yongjinglong'' is known solely from the

GSGM ZH(08)-04, a partial

postcrania Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated sk ...

skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...

and three

teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

, currently housed at the Gansu Geological Museum,

. The postcranial remains include one fragmentary dorsal

rib In vertebrate anatomy, ribs ( la, costae) are the long curved bones which form the rib cage, part of the axial skeleton. In most tetrapods, ribs surround the chest, enabling the lungs to expand and thus facilitate breathing by expanding the ches ...

, the left scapulocoracoid, the right

ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...

and

radius In classical geometry, a radius ( : radii) of a circle or sphere is any of the line segments from its center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', meaning ray but also the ...

, as well as eight presacral

vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic i ...

including one caudal cervical vertebra, four cranial dorsal vertebrae, and three articulated middle dorsal vertebrae. It represents a subadult individual. GSGM ZH(08)-04 was by Li Daqing and You Hailu discovered in 2008 beside the G75 Highway, less than a kilometer from the quarries of '' Daxiatitan'' and '' Huanghetitan liujiaxiaensis'', near Zhongpu. It was collected from the upper

Hekou Group The Hekou Group is a geological group in Gansu Province, China. It is Early Cretaceous in age. Dinosaur body fossils have also been recovered from the Hekou Group, including the iguanodont '' Lanzhousaurus'' and the titanosaurs ''Daxiatitan'', ...

, in the southeastern part of the Lanzhou-Minhe Basin,

, dating to the

Description

''Yongjinglong'' was a medium-sized sauropod. The describers established some diagnostic traits. The premaxillary teeth are long and spoon-shaped. The neck vertebrae and anterior dorsal vertebrae possess large and deep pleurocoels, pneumatic depressions, that cover the entire sides of the centra. The ridges on the sides of the middle dorsal vertebrae form a pattern shaped like a "XI", with the "X" in front. The spine of at least one middle dorsal vertebra is low and not bifurcated and forms with the postzygapophyses a triangular bone plate, directed upwards. The shoulder blade is extremely long, with, in side view, exceptionally straight front and rear edges.

Phylogeny

''Yongjinglong'' was placed in the Titanosauria by Li ''et al.'' (2014). Their cladistic analysis showed it was a member of the Somphospondyli and a sister species of '' Opisthocoelicaudia''. However, a 2019 revision of Chinese "titanosaurs" resolved ''Yongjinglong'' deeply nested within Euhelopodidae, sister taxon to '' Huanghetitan'' and '' Ruyangosaurus'', or a clade including '' Euhelopus'', ''

Erketu ''Erketu'' (meaning "Erketü Tengri") is a genus of somphospondylan sauropod dinosaur that lived in Asia during the Late Cretaceous roughly between 96 million and 89 million years ago. Its fossils were found in Mongolia between 2002 and 2003 du ...

'' and ''

Gobititan ''Gobititan'' is a genus of herbivorous sauropod dinosaur from the Aptian faunal stage of the Early Cretaceous. The name of this genus is derived from the Gobi desert region and the Titans of Greek mythology, which is a reference to its large ...

''.

References

{{Taxonbar, from=Q15690935 Early Cretaceous dinosaurs of Asia Fossil taxa described in 2014 Titanosaurs Taxa named by Peter Dodson Paleontology in Gansu