''Xiphactinus'' (from
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
and
Greek
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*Greek language, a branch of the Indo-European language family.
**Proto-Greek language, the assumed last common ancestor ...
for "
sword
A sword is an edged, bladed weapon intended for manual cutting or thrusting. Its blade, longer than a knife or dagger, is attached to a hilt and can be straight or curved. A thrusting sword tends to have a straighter blade with a pointed t ...
-ray") is an
extinct genus of large (
[Shimada, Kenshu, and Michael J. Everhart. "Shark-bitten Xiphactinus audax (Teleostei: Ichthyodectiformes) from the Niobrara Chalk (Upper Cretaceous) of Kansas." The Mosasaur 7 (2004): 35-39.]) predatory marine
bony fish
Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartila ...
that lived during the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
(
Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± ...
to
Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interv ...
).
Species in the genus bore a superficial resemblance to a gargantuan, fanged
tarpon
Tarpons are fish of the genus ''Megalops''. They are the only members of the family Megalopidae. Of the two species, one (''M. atlanticus'') is native to the Atlantic, and the other (''M. cyprinoides'') to the Indo-Pacific Oceans.
Species an ...
.
The species ''Portheus molossus'' described by Cope is a
junior synonym of ''X. audax''. Skeletal remains of ''Xiphactinus'' have come from the
Carlile Shale
The Carlile Shale is a Turonian age Upper Cretaceous, Upper/Late Cretaceous series shale geologic Formation (geology), formation in the central-western United States, including in the Great Plains region of Colorado, Kansas, Nebraska, New Mexico ...
and
Greenhorn Limestone
The Greenhorn Limestone or Greenhorn Formation is a geologic formation in the Great Plains Region of the United States, dating to the Cenomanian and Turonian ages of the Late Cretaceous period. The formation gives its name to the Greenhorn cyc ...
of
Kansas
Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to ...
(where the first ''Xiphactinus'' fossil was discovered during the 1850s in the
Niobrara Chalk
The Niobrara Formation , also called the Niobrara Chalk, is a geologic formation in North America that was deposited between 87 and 82 million years ago during the Coniacian, Santonian, and Campanian stages of the Late Cretaceous. It is compose ...
),
[''Xiphactinus''](_blank)
at Fossilworks
Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world.
History
Fossilworks was cre ...
.org and Cretaceous formations all over the
East Coast
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Entertainment
* East Coast hip hop, a subgenre of hip hop
* "East Coast" (ASAP Ferg song), 2017
* "East Coast" (Saves the Day song), 2004
* East Coast FM, a radio station in Co. Wicklow, Ireland
* East Coast Swing, a f ...
(most notably
Georgia
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* Related to t ...
,
Alabama
(We dare defend our rights)
, anthem = " Alabama"
, image_map = Alabama in United States.svg
, seat = Montgomery
, LargestCity = Huntsville
, LargestCounty = Baldwin County
, LargestMetro = Greater Birmingham
, area_total_km2 = 135,7 ...
,
North Carolina
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, and
New Jersey
New Jersey is a U.S. state, state in the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. It is bordered on the north and east by the state of New York (state), New York; on the ea ...
) in the
United States
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,
as well as
Europe
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,
Australia, the
Kanguk and
Ashville Formations of
Canada
Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
,
La Luna Formation
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of
Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in ...
and the
Salamanca Formation
The Salamanca Formation is a geologic formation in the Golfo San Jorge Basin of central Patagonia that yields well-preserved, well-dated fossils from the early Paleocene. Studies of these fossils are providing new data on plant and animal diversit ...
in
Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
.
Paleobiology

Species of ''Xiphactinus'' were voracious predatory fish. At least a dozen specimens of ''X. audax'' have been collected with the remains of large, undigested or partially digested prey in their stomachs. In particular, one fossil "Fish-Within-A-Fish" specimen was collected by
George F. Sternberg with another, nearly perfectly preserved long ichthyodectid ''
Gillicus arcuatus
''Gillicus'' was a relatively small, 2-metre long ichthyodectiform fish that lived in the Western Interior Seaway, in what is now central North America, during the Late Cretaceous.
Description
Like its larger relative, ''Ichthyodectes ctenodo ...
'' inside of it. The larger fish apparently died soon after eating its prey, most likely owing to the smaller prey's struggling and rupturing an organ as it was being swallowed. This fossil is on display at the Sternberg Museum of Natural History in
Hays, Kansas
Hays is a city in and the county seat of Ellis County, Kansas, United States. The largest city in northwestern Kansas, it is the economic and cultural center of the region. As of the 2020 census, the population of the city was 21,116. It is ...
.
Like many other species in the Late Cretaceous oceans, a dead or injured individual was likely to be scavenged by sharks (''
Cretoxyrhina
''Cretoxyrhina'' (; meaning 'Cretaceous sharp-nose') is an extinct genus of large mackerel shark that lived about 107 to 73 million years ago during the late Albian to late Campanian of the Late Cretaceous. The type species, ''C. mantelli'', is m ...
'' and ''
Squalicorax
''Squalicorax'', commonly known as the crow shark, is a genus of extinct lamniform shark known to have lived during the Cretaceous period. The genus had a global distribution in the Late Cretaceous epoch. Multiple species within this genus are c ...
''). The remains of a ''Xiphactinus'' were found within a large specimen of ''
Cretoxyrhina
''Cretoxyrhina'' (; meaning 'Cretaceous sharp-nose') is an extinct genus of large mackerel shark that lived about 107 to 73 million years ago during the late Albian to late Campanian of the Late Cretaceous. The type species, ''C. mantelli'', is m ...
'' collected by Charles H. Sternberg. The specimen is on display at the University of Kansas Museum of Natural History.
Virtually nothing is known about the larval or juvenile stages. The smallest fossil specimen of ''X. audax'' consists of a tooth-bearing
premaxilla
The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...
and lower jaws of an individual estimated to be about long.
The species and all other ichthyodectids became extinct near the end of the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
– see
Cretaceous–Paleogene extinction event
The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden extinction event, mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million y ...
.
An incomplete
skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, t ...
of what may be a new species of ''Xiphactinus'' was found in 2002 in the
Czech Republic
The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. Th ...
, in a small town called
Šachov next to
Borohrádek
Borohrádek (; german: Heideburg) is a town in Rychnov nad Kněžnou District in the Hradec Králové Region of the Czech Republic. It has about 2,100 inhabitants.
Administrative parts
The village of Šachov is an administrative part of Borohr ...
city, by then 16-year-old student Michal Matějka.
In July 2010 the bones of a ''Xiphactinus'' were discovered near Morden, Manitoba, Canada. The specimen was about long and was found with the flipper of a
mosasaur
Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Ancient Greek, Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Ma ...
between its jaws.
"Unicerosaurus"
In 1982, a former Baptist minister,
Carl Baugh
Carl Edward Baugh (born October 21, 1936) is an American young Earth creationist. Baugh has claimed to have discovered human footprints alongside dinosaur footprints near the Paluxy River in Texas. Baugh promoted creationism as the former host o ...
, began excavations on the limestone beds of the
Paluxy River, near
Glen Rose, Texas
Glen Rose is a city in and the county seat of Somervell County, Texas, United States. As of the 2012 census estimate, the city population was 2,502.
History
19th century
The area was first settled in 1849 by Charles Barnard, who opened a tradi ...
, famous for its dinosaur tracks. Some of the tracks resembled human footprints and had been proclaimed since 1900 as evidence that dinosaurs and modern humans had once lived alongside one another. Scientists' investigations found the supposed human footprints to be "forms of elongate dinosaur tracks, while others were selectively highlighted erosional markings, and still others (on loose blocks) probable carvings." While excavating, he found a solitary "Y-shaped" fossil that he informally called "Unicerosaurus". In a 1987 popular article, John Armstrong described the fossil as a "Y-shaped petrified bone that appears to be the neural spine from a huge fish like the ''Portheus'' of
Niobrara Chalk
The Niobrara Formation , also called the Niobrara Chalk, is a geologic formation in North America that was deposited between 87 and 82 million years ago during the Coniacian, Santonian, and Campanian stages of the Late Cretaceous. It is compose ...
" that Baugh's museum "declared to be the forehead horn of a newly discovered dinosaur genus". The museum's exhibit told visitors that the "horn" belonged to "the
unicorn
The unicorn is a legendary creature that has been described since Classical antiquity, antiquity as a beast with a single large, pointed, spiraling horn (anatomy), horn projecting from its forehead.
In European literature and art, the unicor ...
of Job 38, one of three dinosaurs mentioned in Scripture; the others being behemoth and leviathan of Job 40 and 41", and that the horn was able to fold back like the blade of a jack knife. Although some Young Earth Creationists shared Baugh's interpretations of the biblical
Behemoth
Behemoth (; he, בְּהֵמוֹת, ''bəhēmōṯ'') is a beast from the biblical Book of Job, and is a form of the primeval chaos-monster created by God at the beginning of creation; he is paired with the other chaos-monster, Leviathan, and ...
and
Leviathan
Leviathan (; he, לִוְיָתָן, ) is a sea serpent noted in theology and mythology. It is referenced in several books of the Hebrew Bible, including Psalms, the Book of Job, the Book of Isaiah, the Book of Amos, and, according to so ...
, Baugh's claims were not taken seriously either by Christian organizations or the scientific community.
In popular culture

In October 2010, Kansas House Rep. Tom Sloan (R-Lawrence) announced that he would introduce legislation to make ''Xiphactinus audax'', a.k.a. the "X-fish", the state fossil of Kansas.
Notes
References
External links
BBCCanadian Fossil Discovery Centre
{{Taxonbar, from=Q131444
Ichthyodectidae
Marine fish genera
Prehistoric ray-finned fish genera
Late Cretaceous fish of North America
Cretaceous Canada
Fossils of Canada
Cretaceous United States
Fossils of the United States
Cretaceous fish of Europe
Prehistoric fish of Australia
Prehistoric fish of South America
Cretaceous Venezuela
Fossils of Venezuela
Albian genus first appearances
Cenomanian genera
Turonian genera
Coniacian genera
Santonian genera
Campanian genera
Maastrichtian genus extinctions
Demopolis Chalk
Mooreville Chalk
Fossil taxa described in 1870
Taxa named by Joseph Leidy
Cretaceous Argentina
Golfo San Jorge Basin
Salamanca Formation