Xinjiangtitan (adjusted)
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''Xinjiangtitan'' ( zh, s=新疆巨龙, p=Xīnjiāngjùlóng) is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of mamenchisaurid
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
that lived during the
Middle Jurassic The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
of what is now
Xinjiang Xinjiang,; , SASM/GNC romanization, SASM/GNC: Chinese postal romanization, previously romanized as Sinkiang, officially the Xinjiang Uygur Autonomous Region (XUAR), is an Autonomous regions of China, autonomous region of the China, People' ...
, northwestern
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. Its type and only species is ''Xinjiangtitan shanshanesis'' ( zh, s=鄯善新疆巨龙, p=Shànshàn Xīnjiāngjùlóng), known from a single incomplete skeleton recovered from the Qiketai Formation. The holotype preserves one of the most complete vertebral columns of any sauropod found in Asia, and has the longest complete neck known for any animal.


Discovery

The
type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
of ''Xinjiangtitan shanshanesis'' was discovered by a joint expedition of Jilin University, Shenyang Normal University, and Xinjiang Geological Survey Institutute in 2012, from a quarry south of Qiketai,
Xinjiang Xinjiang,; , SASM/GNC romanization, SASM/GNC: Chinese postal romanization, previously romanized as Sinkiang, officially the Xinjiang Uygur Autonomous Region (XUAR), is an Autonomous regions of China, autonomous region of the China, People' ...
. In
2013 2013 was the first year since 1987 to contain four unique digits (a span of 26 years). 2013 was designated as: *International Year of Water Cooperation *International Year of Quinoa Events January * January 5 – 2013 Craig, Alask ...
, before the specimen had been fully excavated, Wu Wen-hao, Zhou Chang-fu, Oliver Wings, Toru Sekiha and
Dong Zhiming Dong Zhiming (Chinese language, Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; January 1937 – 20 October 2024) was a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) ...
described it as a new genus and species, ''Xinjiangtitan shanshanesis''. The generic name is derived from Xinjiang, and from ''
titan Titan most often refers to: * Titan (moon), the largest moon of Saturn * Titans, a race of deities in Greek mythology Titan or Titans may also refer to: Arts and entertainment Fictional entities Fictional locations * Titan in fiction, fictiona ...
'', giant in
Greek mythology Greek mythology is the body of myths originally told by the Ancient Greece, ancient Greeks, and a genre of ancient Greek folklore, today absorbed alongside Roman mythology into the broader designation of classical mythology. These stories conc ...
. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''shanshanesis'', is derived from an alternative name for the county where it was found, Shanshan, named after the ancient Shanshan Kingdom. The specific name is occasionally misspelled as "shanshanensis" or "shashaensis", which are invalid spellings even though the correct form would indeed have been "shanshanensis" rather than ''shanshanesis'', as the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
suffix "-ensis" meaning "from", was used to create the name. Continued excavation of the type specimen revealed that the neck and tail were nearly complete.


Fossils

''Xinjiangtitan shanshanesis'' is known from a single specimen, the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
SSV12001, which consists of a nearly complete vertebral column preserved in articulation, as well as a partial skull, partial pelvis, and most of the left hind limb. The neck, of which all 18 vertebrae are present, is nearly 15 meters long and is the longest complete neck ever discovered of any animal. The vertebral column is overall among the most complete of any sauropod specimen from Asia. The cervical and dorsal vertebrae have been described in detail. The specimen was originally reported as being from the
Qigu Formation The Qigu Formation is a Late Jurassic ( Oxfordian) geologic formation in the Southern Junggar Basin in China. Indeterminate Dinosaur remains are among the fossils that have been recovered from the formation, including the stegosaur (" Angustungu ...
(
Late Jurassic The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
), but it was subsequently considered to be from the Qiketai Formation (
Middle Jurassic The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
).


Description

''Xinjiangtitan'' was one of the longest known sauropods, measuring long and weighing up to . ''Xinjiangtitan'' was diagnosed based on the following traits: the presence of a ventral keel on the penultimate cervical centrum that forms a small
semicircular In mathematics (and more specifically geometry), a semicircle is a one-dimensional locus of points that forms half of a circle. It is a circular arc that measures 180° (equivalently, radians, or a half-turn). It only has one line of symmetr ...
process under the distal articular facet; both cervical vertebrae are relatively elongated; the sacricostal yoke except the first sacral rib; and an extremely robust femur. The series of dorsal vertebrae has a length of . The thighbone is long, and the tibia is long. Its long neck is measured at least in length.


Phylogeny

Among sauropods, a
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analysis places ''Xinjiangtitan'' as the
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
of ''
Mamenchisaurus ''Mamenchisaurus'' ( , or spelling pronunciation ) is an extinct genus of sauropod dinosaurs known for their remarkably long necks which made up nearly half the total body length. Numerous species have been assigned to the genus; however, the val ...
'', the only other mamenchisaurid included. ''Xinjiangtitan'' shares several derived characters with
diplodocid Diplodocids, or members of the family Diplodocidae ("double beams"), are a group of sauropod dinosaurs. The family includes some of the longest creatures ever to walk the Earth, including '' Diplodocus'' and '' Supersaurus'', some of which may ha ...
s, including prominent ambiens process of pubis, relatively short hind limb and
fourth trochanter The fourth trochanter is a shared characteristic common to archosaurs. It is a protrusion on the posterior-medial side of the middle of the femur shaft that serves as a muscle attachment, mainly for the '' musculus caudofemoralis longus'', the m ...
on the femur that is caudomedially developed.


References

{{Taxonbar, from=Q15144358 Mamenchisauridae Dinosaur genera Callovian dinosaurs Dinosaurs of China Fossil taxa described in 2013 Taxa named by Dong Zhiming