''Xenodon merremii'', also known commonly as Wagler's snake, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...

in the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Dipsadinae Dipsadinae is a large subfamily of colubroid snakes, sometimes referred to as a family (Dipsadidae). Species of the subfamily Dipsadinae are found in most of the Americas, including the West Indies, and are most diverse in South America. There a ...

of the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Colubridae Colubridae (, commonly known as colubrids , from , 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest fossil species of the family date back to the Late Eocene epoch, with earlier origins suspected. C ...

. The species is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

and is widespread in the eastern half of the continent.

Geographic range

''X. merremii'' occurs in

Venezuela Venezuela, officially the Bolivarian Republic of Venezuela, is a country on the northern coast of South America, consisting of a continental landmass and many Federal Dependencies of Venezuela, islands and islets in the Caribbean Sea. It com ...

Guyana Guyana, officially the Co-operative Republic of Guyana, is a country on the northern coast of South America, part of the historic British West Indies. entry "Guyana" Georgetown, Guyana, Georgetown is the capital of Guyana and is also the co ...

Suriname Suriname, officially the Republic of Suriname, is a country in northern South America, also considered as part of the Caribbean and the West Indies. It is a developing country with a Human Development Index, high level of human development; i ...

French Guiana French Guiana, or Guyane in French, is an Overseas departments and regions of France, overseas department and region of France located on the northern coast of South America in the Guianas and the West Indies. Bordered by Suriname to the west ...

Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...

Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...

Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...

Uruguay Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...

, and northern

Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...

.. www.reptile-database.org.

Etymology

The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''merremii'', is in honor of German

herpetologist Herpetology (from Ancient Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is a branch of zoology concerned with the study of amphibians (including frogs, salamanders, and caecilians (Gymnophiona)) and reptiles (in ...

Blasius Merrem Blasius Merrem (4 February 1761 – 23 February 1824) was a German natural history, naturalist, zoologist, Ornithology, ornithologist, mathematician, and Herpetology, herpetologist. In 1804, he became the professor of political economy and botan ...

Description

Adults of ''X. merremii'' are usually or less in total length (including tail). Freiberg M (1982). Its color pattern is very variable. Some "red phase" specimens are a uniform reddish tan. Other specimens are pale brown

dorsally Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provi ...

, with broad dark-brown crossbands, which are edged with black, and are narrower or interrupted in the middle. Boulenger GA (1894). The latter color pattern resembles that of the

venomous snake ''Venomous snakes'' are species of the suborder Serpentes that are capable of producing venom, which they use for killing prey, for defense, and to assist with digestion of their prey. The venom is typically delivered by injection using hollow ...

Bothrops alternatus ''Bothrops alternatus'', known by the common names crossed pit viper''U.S. Navy. 1991. ''Poisonous Snakes of the World''. US Govt. New York: Dover Publications Inc. 203 pp. .'', yarará grande, and urutu,Campbell JA, Lamar WW. 2004. ''The Venomo ...

''.

Diet

''X. merremii'' preys on insects, frogs, toads, lizards, and sometimes snakes. Like other rear-fanged toad-eaters of the

genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...

Heterodon ''Heterodon'' is a genus of harmless colubrid snakes endemic to North America.Platt, Dwight R. 1983. Heterodon'. Catalogue of American Amphibians and Reptiles. Society for the Study of Amphibians and Reptiles, 351: 1-2. They are stout with upt ...

'' and ''

Lystrophis ''Lystrophis'' is the genus of tricolored South American hognose snakes. They mimic Milk Snakes, milk snakes or coral snakes with their red, black, and white ringed patterns. Species The genus ''Lystrophis'' contains five species that are rec ...

'', ''X. merremii'' uses its enlarged posterior maxillary teeth to puncture and deflate toads which have defensively puffed themselves up, thereby making them easier to swallow.

Defensive behavior

When threatened, ''X. merremii'' raises the anterior part of its body, inflating and spreading its neck, similar to a cobra.

Reproduction

''X. merremii'' is

oviparous Oviparous animals are animals that reproduce by depositing fertilized zygotes outside the body (i.e., by laying or spawning) in metabolically independent incubation organs known as eggs, which nurture the embryo into moving offsprings kno ...