
Wildlife crossings are structures that allow
animal
Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s to cross
human
Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
-made barriers safely. Wildlife crossings may include
underpass tunnels or wildlife tunnels,
viaducts, and
overpasses or green bridges (mainly for large or
herd-type animals);
amphibian tunnels;
fish ladders;
canopy bridges (especially for monkeys and squirrels);
tunnels and
culverts (for small mammals such as
otters,
hedgehogs, and
badgers); and
green roofs (for butterflies and birds).
Wildlife crossings are a practice in
habitat conservation, allowing connections or reconnections between
habitat
In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s, combating
habitat fragmentation. They also assist in avoiding
collisions between vehicles and animals, which in addition to killing or injuring wildlife may cause injury or death to humans and
property damage.
Similar structures can be used for domesticated animals, such as
cattle creeps.
Roads and habitat fragmentation
Habitat fragmentation occurs when human-made barriers such as
road
A road is a thoroughfare used primarily for movement of traffic. Roads differ from streets, whose primary use is local access. They also differ from stroads, which combine the features of streets and roads. Most modern roads are paved.
Th ...
s,
railroad
Rail transport (also known as train transport) is a means of transport using wheeled vehicles running in railway track, tracks, which usually consist of two parallel steel railway track, rails. Rail transport is one of the two primary means of ...
s,
canal
Canals or artificial waterways are waterways or engineered channels built for drainage management (e.g. flood control and irrigation) or for conveyancing water transport vehicles (e.g. water taxi). They carry free, calm surface ...
s,
electric power lines, and
pipelines penetrate and divide wildlife habitat.
Of these, roads have the most widespread and detrimental effects.
Scientists estimate that the system of roads in the
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
affects the ecology of at least one-fifth of the land area of the country. For many years ecologists and conservationists have documented the adverse relationship between roads and wildlife, and identify four ways that roads and traffic detrimentally affect wildlife populations: (1) they decrease habitat amount and quality, (2) they increase mortality due to
wildlife-vehicle collisions (road kill), (3) they prevent access to resources on the other side of the road, and (4) they subdivide wildlife populations into smaller and more vulnerable sub-populations (fragmentation). Habitat fragmentation can lead to extinction or extirpation if a population's gene pool is restricted enough.
The first three effects (loss of habitat, road kill, and isolation from resources) exert pressure on various animal populations by reducing available resources and directly killing individuals in a population. For instance, found that road kills do not pose a significant threat to healthy populations but can be devastating to small, shrinking, or threatened populations. Road mortality has significantly affected a number of prominent species in the United States, including
white-tailed deer (''Odocoileus virginianus''),
Florida panthers (''Puma concolor coryi''), and
black bears (''Ursus americanus''). In addition, habitat loss can be direct, if habitat is destroyed to make room for a road, or indirect, if habitat quality close to roads is compromised due to emissions from the roads (e.g. noise, light, runoff, pollution, etc.). Finally, species that are unable to migrate across roads to reach resources such as food, shelter and mates will experience reduced reproductive and survival rates, which can compromise population viability.
In addition to the first three factors, numerous studies have shown that the construction and use of roads is a direct source of habitat fragmentation.
As mentioned above, populations surrounded by roads are less likely to receive immigrants from other habitats and as a result, they suffer from a lack of
genetic diversity
Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species. It ranges widely, from the number of species to differences within species, and can be correlated to the span of survival for a species. It is d ...
. These small populations are particularly vulnerable to
extinction
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
due to demographic, genetic, and
environmental stochasticity because they do not contain enough
alleles to adapt to new selective pressures such as changes in temperature, habitat, and food availability.
The relationship between roads and habitat fragmentation is well documented. One study found that roads contribute more to fragmentation in forest habitats than clear cuts. Another study concluded that road fragmentation of formerly contiguous forest in eastern North America is the primary cause for the decline of forest bird species and has also significantly harmed small mammals, insects, and reptiles in the United States.
After years of research, biologists agree that roads and traffic lead to habitat fragmentation, isolation and road kill, all of which combine to significantly compromise the viability of wildlife populations throughout the world.
Wildlife-vehicle collisions
Wildlife-vehicle collisions have a significant cost for human populations because collisions damage property and injure and kill passengers and drivers. Research in the 1990s estimated the number of collisions with
ungulates in traffic in Europe at 507,000 per year, resulting in 300 people killed, 30,000 injured,
and property damage exceeding $1 billion. In parallel, 1.5 million traffic accidents involving deer in the United States cause an estimated $1.1 billion in vehicle damage each year.
On a larger scale, research indicates that wildlife-vehicle collisions in the United States result in 29,000 injuries and more than 200 fatalities per year.
The conservation issues associated with roads (wildlife mortality and habitat fragmentation) coupled with the substantial human and economic costs resulting from wildlife-vehicle collisions have caused scientists, engineers, and transportation authorities to consider a number of mitigation tools for reducing the conflict between roads and wildlife. Of the currently available options, structures known as wildlife crossings have been the most successful at reducing both habitat fragmentation and wildlife-vehicle collisions caused by roads.

Wildlife crossings are structural passages beneath or above roadways that are designed to facilitate safe wildlife movement across roadways.
In recent years, conservation biologists and wildlife managers have advocated wildlife crossings coupled with roadside fencing as a way to increase road permeability and habitat connectivity while decreasing wildlife-vehicle collisions. Wildlife crossing is the umbrella term encompassing underpasses, overpasses, ecoducts, green bridges, amphibian/small mammal tunnels, and wildlife viaducts . All of these structures are designed to provide semi-natural corridors above and below roads so that animals can safely cross without endangering themselves and motorists.
History
Written reports of rough fish ladders date to 17th-century France, where bundles of branches were used to create steps in steep channels to bypass obstructions. A version was patented in 1837 by Richard McFarlan of
Bathurst, New Brunswick
Bathurst () is a city in northern New Brunswick with a population of 12,157 and the 4th largest metropolitan area in New Brunswick as defined by Census Canada with a population of 31,387 as of 2021. The City of Bathurst overlooks Nepisiguit Ba ...
, Canada, who designed a fishway to bypass a dam at his water-powered lumber mill. In 1880, the first fish ladder was built in
Rhode Island
Rhode Island ( ) is a state in the New England region of the Northeastern United States. It borders Connecticut to its west; Massachusetts to its north and east; and the Atlantic Ocean to its south via Rhode Island Sound and Block Is ...
, United States, on the
Pawtuxet Falls Dam. As the
Industrial Age advanced, dams and other river obstructions became larger and more common, leading to the need for effective fish by-passes.
The first overland wildlife crossings were constructed in France during the 1950s.
European countries including the Netherlands, Switzerland, Germany, and France have been using various crossing structures to reduce the conflict between wildlife and roads for several decades and use a variety of overpasses and underpasses to protect and re-establish
wildlife such as: amphibians, badgers, ungulates, invertebrates, and other small mammals.
The
Humane Society of the United States reported in 2007 that the more than 600 tunnels installed under major and minor roads in the
Netherlands
, Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...
have helped to substantially increase population levels of the endangered
European badger.
[Wildlife crossings - Wild animals and roads](_blank)
The Humane Society of the United States. Archived fro
the original
on September 27, 2007. The longest "ecoduct" overpass, Natuurbrug Zanderij Crailoo, in the Netherlands, runs and spans a highway, railway and
golf course.
Wildlife crossings are becoming increasingly common in
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
and the United States. Recognizable wildlife crossings are found in
Banff National Park in
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
, where vegetated overpasses provide safe passage over the
Trans-Canada Highway
The Trans-Canada Highway (Canadian French, French: ; abbreviated as the TCH or T-Can) is a transcontinental federal–provincial highway system that travels through all ten provinces of Canada, from the Pacific Ocean on the west coast to the A ...
for bears, moose, deer, wolves, elk, and many other species.
The 24 wildlife crossings in Banff were constructed as part of a road improvement project in 1978.
In the United States, thousands of wildlife crossings have been built in the past 30 years, including culverts, bridges, and overpasses. These have been used to protect
mountain goats in Montana,
spotted salamander
The spotted salamander (''Ambystoma maculatum''), also known commonly as the yellow-spotted salamander, is a species of mole salamander in the family Ambystomatidae. The species is native to the eastern United States and Canada. It is the sta ...
s in Massachusetts,
bighorn sheep in Colorado,
desert tortoises in
California
California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
, and endangered
Florida panthers in
Florida
Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
.
The
Henry Street salamander tunnels are tunnels under Henry Street in
North Amherst, Massachusetts: they help salamanders cross Henry Street to get to
vernal pools that the salamanders use for breeding.
The first wildlife crossing in the Canadian province of
Ontario
Ontario is the southernmost Provinces and territories of Canada, province of Canada. Located in Central Canada, Ontario is the Population of Canada by province and territory, country's most populous province. As of the 2021 Canadian census, it ...
was built in 2010, along
Ontario Highway 69 between
Sudbury and
Killarney, as part of the route's ongoing freeway conversion.
Costs and benefits
The benefits derived from constructing wildlife crossings to extend wildlife migration corridors over and under major roads appear to outweigh the costs of construction and maintenance. One study estimates that adding wildlife crossings to a road project is a 7–8% increase in the total cost of the project . Theoretically, the monetary costs associated with constructing and maintaining wildlife crossings in ecologically important areas are trumped by the benefits associated with protecting wildlife populations, reducing property damage to vehicles, and saving the lives of drivers and passengers by reducing the number of collisions caused by wildlife.
A study completed for the Virginia Department of Transportation estimated that underpasses for wildlife become cost effective, in terms of property damage, when they prevent between 2.6 and 9.2 deer-vehicle collisions per year, depending on the cost of the underpass. Approximately 300 deer crossed through the underpasses in the year the study took place .
Effectiveness
A number of studies have been conducted to determine the effectiveness of
wildlife corridors at providing habitat connectivity (by providing viable migration corridors) and reducing wildlife-vehicle collisions. The effectiveness of these structures appears to be highly site-specific (due to differences in location, structure, species, habitat, etc.), and also dependent on design, but crossings have been beneficial to a number of species in a variety of locations.
Examples
Banff National Park
Banff National Park offers one of the best opportunities to study the effectiveness of wildlife crossings because the park contains a wide variety of species and is bisected by the
Trans-Canada Highway
The Trans-Canada Highway (Canadian French, French: ; abbreviated as the TCH or T-Can) is a transcontinental federal–provincial highway system that travels through all ten provinces of Canada, from the Pacific Ocean on the west coast to the A ...
(TCH), a large commercial road. To reduce the effects of the four-lane TCH, 24 wildlife crossings (22 underpasses and two overpasses) were built to ensure habitat connectivity and protect motorists . In 1996,
Parks Canada developed a contract with university researchers to assess the effectiveness of the crossings. Subsequently, a number of publications have analyzed the crossings' effect on various species and overall wildlife mortality (see , , and ).

Using a variety of techniques to monitor the crossings since the early 1980s, scientists report that 10 species of large mammals (including deer, elk, black bear,
grizzly bear, mountain lion, wolf, moose, and
coyote) have used the 24 crossings in Banff a total of 84,000 times as of January 2007 . The research also identified a "
learning curve" such that animals need time to acclimate to the structures before they feel comfortable using them. For example, grizzly bear crossings increased from seven in 1996 to more than 100 in 2006, although the actual number of individual bears using the structures remained constant over this time at between two and four bears (Parks Canada, unpublished results). A similar set of observations was made for wolves, with crossings increasing from two to approximately 140 over the same 10-year period. However, in this case the actual number of wolves in the packs using the crossings increased dramatically, from a low of two up to a high of over 20 individuals.
reported that the use of wildlife crossings and fencing reduced traffic-induced mortality of large ungulates on the TCH by more than 80 percent. Recent analysis for carnivores showed results were not as positive however, with bear mortality increasing by an average of 116 percent in direct parallel to an equal doubling of traffic volumes on the highway, clearly showing no effect of fencing to reduce bear mortality (Hallstrom, Clevenger, Maher and Whittington, in prep). Research on the crossings in Banff has thus shown mixed value of wildlife crossings depending on the species in question.
Parks Canada is currently planning to build 17 additional crossing structures across the TCH to increase driver safety near the hamlet of
Lake Louise. Lack of effectiveness of standard fencing in reducing bear mortality demonstrates that additional measures such as wire 'T-caps' on the fence may be needed for fencing to mitigate effectively for bears (Hallstrom, Clevenger, Maher and Whittington, in prep).
Collier and Lee Counties in Florida
Twenty-four wildlife crossings (highway underpasses) and 12 bridges modified for wildlife have been constructed along a 40-mile stretch of
Interstate 75 in
Collier and
Lee Counties in
Florida
Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
. These crossings are specifically designed to target and protect the endangered Florida panther, a subspecies of cougar found in the
Southeastern United States. Scientists estimate that there are only 80–100 Florida panthers alive in the wild, which makes them one of the most endangered large mammals in North America . The Florida panther is particularly vulnerable to wildlife-vehicle collisions, which claimed 11 panthers in 2006 and 14 in 2007 .
The
Florida Fish and Wildlife Conservation Commission (FWC) has used a number of mitigation tools in an effort to protect Florida panthers and the combination of wildlife crossings and fences have proven the most effective . As of 2007, no panthers have been killed in areas equipped with continuous fencing and wildlife crossings and the FWC is planning to construct many more crossing structures in the future. The underpasses on I-75 also appeared to benefit bobcats, deer, and raccoons by significantly reducing wildlife-vehicle collisions along the interstate .
Southern California
Wildlife crossings have also been important for protecting
biodiversity in several areas of
southern California
Southern California (commonly shortened to SoCal) is a geographic and Cultural area, cultural List of regions of California, region that generally comprises the southern portion of the U.S. state of California. Its densely populated coastal reg ...
. In
San Bernardino County, biologists have erected fences along
State Route 58 to complement underpasses (culverts) that are being used by the threatened
desert tortoise. Tortoise deaths on the highway declined by 93% during the first four years after the introduction of the fences, proving that even makeshift wildlife crossings (storm-drainage culverts in this case) have the ability to increase highway permeability and protect sensitive species . Studies by and report that underpasses in Orange, Riverside, and Los Angeles Counties have drawn significant use from a variety of species including
bobcats,
coyotes,
gray fox,
mule deer
The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer.
Unlike the related whit ...
, and
long-tailed weasels. These results could be extremely important for wildlife conservation efforts in the region's
Puente Hills and
Chino Hills links, which have been increasingly fragmented by road construction . Los Angeles County's first wildlife-purpose built underpass is at Harbor Boulevard. It was built in partnership between Los Angeles County, California State Parks and the Puente Hills Habitat Preservation Authority.
The
Wallis Annenberg Wildlife Crossing in
Agoura Hills, California, will be the world's largest wildlife crossing once completed in 2026.
Ecoducts, Netherlands
The Netherlands has over 66 wildlife crossings (overpasses and ecoducts) that have been used to protect the endangered
European badger, as well as populations of
wild boar,
red deer, and
roe deer. As of 2012, the
Veluwe, of woods, heathland and drifting sands, the largest lowland nature area in North Western Europe, contains nine ecoducts, wide on average, that are used to shuttle wildlife across highways that transect the Veluwe. The first two ecoducts on the Veluwe were built in 1988 across the
A50 when the highway was constructed. Five of the other ecoducts on the Veluwe were built across existing highways, one was built across a two lane provincial road. The two ecoducts across the A50 were used by nearly 5,000 deer and wild boar during a one-year period .
The Netherlands also boasts the world's longest ecoduct-wildlife overpass called the Natuurbrug Zanderij Crailoo (sand quarry nature bridge at Crailo) . The massive structure, completed in 2006, is wide and over long and spans a railway line,
business park, roadway, and sports complex . Monitoring is currently underway to examine the effectiveness of this innovative project combining wildlife protection with urban development. The oldest wildlife passage is Zeist West - A 28, opened in 1988.
Slaty Creek Wildlife Underpass, Calder Freeway, Black Forest, Australia
Another case study of the effectiveness of wildlife crossings comes from an underpass built to minimize the ecological effect of the
Calder Freeway as it travels through the Black Forest in Victoria, Australia. In 1997, the
Victorian Government Roads Corporation built Slaty Creek wildlife underpass at a cost of $3 million . Scientists used 14 different techniques to monitor the underpass for 12 months in order to determine the abundance and diversity of species using the underpass . During the 12-month period, 79 species of fauna were detected in the underpass (compared with 116 species detected in the surrounding forest) including amphibians,
bats, birds,
koala
The koala (''Phascolarctos cinereus''), sometimes inaccurately called the koala bear, is an arboreal herbivorous marsupial native to Australia. It is the only Extant taxon, extant representative of the Family (biology), family ''Phascolar ...
s,
wombats,
gliders, reptiles, and
kangaroos . The results indicate that the underpass could be useful to a wide array of species but the authors suggest that Slaty Creek could be improved by enhanced design and maintenance of fencing to minimise road kill along the Calder Freeway and by attempting to exclude introduced predators such as cats and foxes from the area.
I-70 Vail Pass, Colorado
In 2005, area environmental groups floated the idea of a wildlife overpass west of
Vail Pass. In 2010, ARC Solutions – an interdisciplinary partnership – initiated the International Wildlife Crossing Infrastructure Design Competition for a wildlife crossing over
Interstate 70 in the high country west of
Denver
Denver ( ) is a List of municipalities in Colorado#Consolidated city and county, consolidated city and county, the List of capitals in the United States, capital and List of municipalities in Colorado, most populous city of the U.S. state of ...
,
Colorado
Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...
; designers had to account for challenges unique to the area, including snow and severe weather, high elevation and steep grades, a six-lane roadway, a bike path, and high traffic volumes, as well as multiple species of wildlife, including
lynx.
After receiving 36 submissions from nine countries, a jury of international experts in
landscape architecture, engineering, architecture, ecology and transportation selected five finalists in November 2010 to further develop their conceptual designs for a wildlife crossing structure. In January 2011, the team led by HNTB with Michael Van Valkenburgh & Associates (New York) were selected as the winners. The design features a single 100 m (328 ft) concrete span across the highway that is planted with a variety of vegetation types, including a pine-tree forest and meadow grasses, to attract different species to cross. A modular precast concrete design means that much of the bridge can be constructed offsite and moved into place.
In late 2020, Summit County Safe Passages released the "I-70 East Vail Pass Wildlife Crossings Feasibility Study" for a wildlife overpass.
I-90 near Snoqualmie Pass
In 2005, the
Washington State Department of Transportation
The Washington State Department of Transportation (WSDOT or WashDOT, both ) is a governmental agency that constructs, maintains, and regulates the use of transportation infrastructure in the U.S. state of Washington (state), Washington. Establi ...
received approval to begin a safety improvement project through the
Snoqualmie Pass area along the
Interstate 90 corridor from Hyak to
Easton, through the
Central Cascades and
Mountains to Sound Greenway National Heritage Area, including a series of wildlife crossings. Wildlife habitat on either side of I-90 will be reconnected with the installation of new bridges and culverts, protecting both wildlife and the traveling public. The construction of the wildlife overcrossing began in 2015 and was completed in late 2019. Work to restore habitat on the wildlife bridge over I-90 has continued throughout 2020, with 90,000 trees and shrubs planted on the overcrossing.
Interstate 80 in Parleys Canyon
In 2018, the
Utah Department of Transportation announced a wildlife crossing over
Interstate 80 in
Parleys Canyon.
The project was completed in early 2019 and measures long by wide. It is currently the only wildlife overpass in the state, though Utah has more than 50 wildlife underpasses.
Robert L.B. Tobin Land Bridge
On December 11, 2020, the Robert L.B. Tobin Land Bridge opened over
Wurzbach Parkway in
San Antonio, Texas' Phil Hardberger Park. The project cost $23 million and is designed for both wildlife and pedestrians. Construction began on November 26, 2018, originally expected to end in April 2020, and opened in December 2020. At long and wide, it was the largest wildlife bridge in the United States when it was constructed.
Canopy Bridge in Anamalai Tiger Reserve
Many endangered
lion-tailed macaques used to be killed while crossing the highway at Puduthotam in
Valparai, South India. Thanks to the efforts of NGOs and the forest department, several canopy bridges were installed, connecting trees on either side of the road. This helped to lower the numbers of lion-tailed macaques killed in the region. The
Environment Conservation Group had initiated a national mission to increase awareness on the importance of adopting roadkill mitigation methods through their mission PATH traveling more than across 22 states.
See also
* Colored walls or corridors
**
Aquatic organism passage
**
Emerald network
**
Wildlife corridor, green corridor
* Animal passages
**
Amphibian and reptile tunnel
**
Bat bridge
**
Squirrel bridge
**
Toad tunnel
*
Ecological network
**
Habitat destruction
**
Rewilding
**
Landscape connectivity
References
Bibliography
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*Hallstrom, W., A. P. Clevenger, A. Maher and J Whittington. 2008. Effectiveness of highway mitigation fencing for ungulates and carnivores. Journal of Applied Ecology - In Review.
*.
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External links
Eco-Logical: An Ecosystem Approach to Developing Infrastructure Projects- Federal Highway Administration (FHWA)
Wildlife Crossing Structures-
Yellowstone to Yukon Conservation InitiativeWildlife Crossings in Banff National ParkRoad Ecology Center, UC DavisCalifornia Roadkill Observation SystemMaine Audubon Wildlife Road WatchAn Assessment of Wildlife Habitat Linkages on Interstate 70, UtahWildlife Consulting Resources Wildlife Crossing and Linkage Information for New Highway Projects
Wildlife Crossings Project- The Wildlife Crossings Project provides information about georreferenced wildlife crossings all around the world, and allow specialists to publish them.
{{Authority control
Bridges
Conservation projects
Ecological connectivity
Ecological restoration
Roadkill
Road traffic management
Subway (underpass)
Tunnels