Wildcat Creek Marsh
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The wildcat is a
species complex In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
comprising two small wild cat
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
: the
European wildcat The European wildcat (''Felis silvestris'') is a small wildcat species native to continental Europe, Great Britain, Turkey and the Caucasus. Its fur is brownish to grey with stripes on the forehead and on the sides and has a bushy tail with a bl ...
(''Felis silvestris'') and the
African wildcat The African wildcat (''Felis lybica'') is a small wildcat species with sandy grey fur, pale vertical stripes on the sides and around the face. It is native to Africa, West and Central Asia, and is distributed to Rajasthan in India and Xinjiang ...
(''F. lybica''). The European wildcat inhabits
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
s in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
,
Anatolia Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
and the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
, while the African wildcat inhabits semi-
arid Aridity is the condition of geographical regions which make up approximately 43% of total global available land area, characterized by low annual precipitation, increased temperatures, and limited water availability.Perez-Aguilar, L. Y., Plata ...
landscapes and
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropica ...
s in
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, the
Arabian Peninsula The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...
,
Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...
, into western
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
and western
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. The wildcat species differ in fur pattern, tail, and size: the European wildcat has long fur and a bushy tail with a rounded tip; the smaller African wildcat is more faintly striped, has short sandy-gray fur and a tapering tail; the
Asiatic wildcat The Asiatic wildcat (''Felis lybica ornata''), also known as the Indian desert cat, is an African wildcat subspecies that occurs from the eastern Caspian Sea north to Kazakhstan, into western India, western China and southern Mongolia. There is ...
(''F. lybica ornata'') is spotted. The wildcat and the other members of the cat family had a
common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. According to modern evolutionary biology, all living beings could be descendants of a unique ancestor commonl ...
about 10–15 million years ago. The European wildcat
evolved Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
during the
Cromerian Stage The Cromerian Stage or Cromerian Complex, also called the Cromerian (), is a stage in the Pleistocene glacial history of north-western Europe, mostly occurring more than half a million years ago. It is named after the East Anglian town of Cromer in ...
about 866,000 to 478,000 years ago; its direct ancestor was ''
Felis lunensis ''Felis lunensis,'' or the Martelli's cat is an extinct felid of the subfamily Felinae. Evolution and taxonomy Around 12 million years ago, the genus ''Felis'' appeared and eventually gave rise to many of the modern small cats. ''Felis lunen ...
''. The ''silvestris'' and ''lybica'' lineages probably diverged about 173,000 years ago. The wildcat is categorized as
Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
since 2002, since it is widely distributed in a stable global population exceeding 20,000 mature individuals. Some local populations are threatened by introgressive hybridisation with the
domestic cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small Domestication, domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have sh ...
(''F. catus''), contagious disease, vehicle collisions and persecution. The association of African wildcats and humans appears to have developed along with the establishment of settlements during the
Neolithic Revolution The Neolithic Revolution, also known as the First Agricultural Revolution, was the wide-scale transition of many human cultures during the Neolithic period in Afro-Eurasia from a lifestyle of hunter-gatherer, hunting and gathering to one of a ...
, when
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s in grain stores of early
farmer A farmer is a person engaged in agriculture, raising living organisms for food or raw materials. The term usually applies to people who do some combination of raising field crops, orchards, vineyards, poultry, or other livestock. A farmer ...
s attracted wildcats. This association ultimately led to it being
tamed A tame animal is an animal that is relatively tolerant of human presence. Tameness may arise naturally (as in the case, for example, of island tameness) or due to the deliberate, human-directed process of animal training, training an animal agai ...
and
domesticated Domestication is a multi-generational mutualistic relationship in which an animal species, such as humans or leafcutter ants, takes over control and care of another species, such as sheep or fungi, to obtain from them a steady supply of reso ...
: the domestic cat is the direct descendant of the African wildcat. It was one of the revered
cats in ancient Egypt In ancient Egypt, cats were represented in social and religious scenes dating as early as 1980 BC. Several ancient Egyptian deities were depicted and sculptured with cat-like heads such as Mafdet, Bastet and Sekhmet, representing justice, fertili ...
. The European wildcat has been the subject of
mythology Myth is a genre of folklore consisting primarily of narratives that play a fundamental role in a society. For scholars, this is very different from the vernacular usage of the term "myth" that refers to a belief that is not true. Instead, the ...
and
literature Literature is any collection of Writing, written work, but it is also used more narrowly for writings specifically considered to be an art form, especially novels, Play (theatre), plays, and poetry, poems. It includes both print and Electroni ...
.


Taxonomy

''Felis (catus) silvestris'' was the
scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
used in 1777 by Johann von Schreber when he described the European wildcat based on descriptions and names proposed by earlier naturalists such as
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...
,
Ulisse Aldrovandi Ulisse Aldrovandi (11 September 1522 – 4 May 1605) was an Italian naturalist, the moving force behind Bologna's botanical garden, one of the first in Europe. Carl Linnaeus and the comte de Buffon reckoned him the father of natural history stud ...
and
Conrad Gessner Conrad Gessner (; ; 26 March 1516 – 13 December 1565) was a Swiss physician, naturalist, bibliographer, and philologist. Born into a poor family in Zürich, Switzerland, his father and teachers quickly realised his talents and supported him t ...
. ''Felis lybica'' was the name proposed in 1780 by
Georg Forster Johann George Adam Forster, also known as Georg Forster (; 27 November 1754 – 10 January 1794), was a German geography, geographer, natural history, naturalist, ethnology, ethnologist, travel literature, travel writer, journalist and revol ...
, who described an African wildcat from
Gafsa Gafsa (; ; ') is the capital of Gafsa Governorate in Tunisia. With a population of 120,739, Gafsa is the ninth-largest Tunisian city and is 335 km from the country's capital, Tunis. Overview Gafsa is the capital of Gafsa Governorate, in ...
on the
Barbary Coast The Barbary Coast (also Barbary, Berbery, or Berber Coast) were the coastal regions of central and western North Africa, more specifically, the Maghreb and the Ottoman borderlands consisting of the regencies in Algiers, Tunis, and Tripoli, a ...
. In subsequent decades, several naturalists and explorers described 40 wildcat
specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository f ...
s collected in European, African and Asian range countries. In the 1940s, the taxonomist
Reginald Innes Pocock Reginald Innes Pocock, (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock and Edith Prichard. He began showing interest in natural history at St. Edward's ...
reviewed the collection of wildcat skins and skulls in the
Natural History Museum, London The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum (Lo ...
, and designated seven ''F. silvestris''
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
from Europe to
Asia Minor Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
, and 25 ''F. lybica'' subspecies from
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, and
West West is one of the four cardinal directions or points of the compass. It is the opposite direction from east and is the direction in which the Sun sets on the Earth. Etymology The word "west" is a Germanic word passed into some Romance langu ...
to
Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...
. Pocock differentiated the: * Forest wildcat subspecies (''silvestris'' group) * Steppe wildcat subspecies (''ornata''-''caudata'' group): is distinguished from the forest wildcat by being smaller, with comparatively lighter fur colour, and longer and more sharply-pointed tails. The domestic cat is thought to have derived from this group. * Bush wildcat subspecies (''ornata''-''lybica'' group): is distinguished from the steppe wildcat by paler fur, well-developed spot patterns and bands. In 2005, 22 subspecies were recognized by the authors of ''
Mammal Species of the World ''Mammal Species of the World: A Taxonomic and Geographic Reference'' is a standard reference work in mammalogy giving descriptions and Bibliographic database, bibliographic data for the known species of mammals. It is now in its third edition, ...
'', who allocated subspecies largely in line with Pocock's assessment. In 2006, the
Chinese mountain cat The Chinese mountain cat (''Felis bieti''), also known as Chinese desert cat and Chinese steppe cat, is a small wild '' Felis'' species with sand-coloured fur, faint dark stripes on the face and legs and black tipped ears. It is endemic to the ...
was placed within the wildcat lineage, being found to be more closely related to the Asiatic wildcat and the domestic cat than to the European wildcat through nuclear DNA, while being placed as an outgroup to the wildcat clade through mitochondrial DNA. In 2017, the Cat Classification Task Force revised the
taxonomy image:Hierarchical clustering diagram.png, 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme o ...
of the Felidae, and recognized the following as valid taxa:


Evolution

The wildcat is a member of the Felidae, a family that had a
common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. According to modern evolutionary biology, all living beings could be descendants of a unique ancestor commonl ...
about 10–15 million years ago. ''Felis'' species diverged from the Felidae around 6–7 million years ago. The European wildcat diverged from ''Felis'' about 1.09 to 1.4 million years ago. The European wildcat's direct ancestor was ''
Felis lunensis ''Felis lunensis,'' or the Martelli's cat is an extinct felid of the subfamily Felinae. Evolution and taxonomy Around 12 million years ago, the genus ''Felis'' appeared and eventually gave rise to many of the modern small cats. ''Felis lunen ...
'', which lived in Europe in the late
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Villafranchian Villafranchian age ( ) is a period of geologic time (3.5–1.0 Ma) spanning the Late Pliocene and Early Pleistocene used more specifically with European Land Mammal Ages. Named by Italian geologist Lorenzo Pareto for a sequence of terrestrial se ...
periods.
Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
remains indicate that the transition from ''lunensis'' to ''silvestris'' was completed by the Holstein interglacial about 340,000 to 325,000 years ago. Craniological differences between the European and African wildcats indicate that the wildcat probably migrated during the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
from Europe into the Middle East, giving rise to the steppe wildcat
phenotype In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology (physical form and structure), its developmental processes, its biochemical and physiological propert ...
.
Phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
research revealed that the ''lybica'' lineage probably diverged from the ''silvestris'' lineage about 173,000 years ago.


Characteristics

The wildcat has pointed ears, which are moderate in length and broad at the base. Its
whiskers Whiskers, also known as vibrissae (; vibrissa; ) are a type of stiff, functional hair used by most therian mammals to sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are coarser as ta ...
are white, number 7 to 16 on each side and reach in length on the muzzle. Whiskers are also present on the inner surface of the paw and measure . Its eyes are large, with vertical
pupil The pupil is a hole located in the center of the iris of the eye that allows light to strike the retina.Cassin, B. and Solomon, S. (1990) ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing Company. It appears black becau ...
s and yellowish-green irises. The
eyelashes An eyelash (also called lash) (Neo-Latin: ''cilium'', plural ''cilia'') is one of the hairs that grows at the edges of the top and bottom eyelids, spanning outwards and away from the eyes. The lashes grow in up to six layers on each of the upper ...
range from in length, and can number six to eight per side. The European wildcat has a greater
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
volume than the domestic cat, a ratio known as Schauenberg's index. Further, its skull is more spherical in shape than that of the
jungle cat The jungle cat (''Felis chaus''), also called reed cat and swamp cat, is a medium-sized cat native from the Eastern Mediterranean region and the Caucasus to parts of Central, South and Southeast Asia. It inhabits foremost wetlands like swamps, ...
(''F. chaus'') and
leopard cat The leopard cat (''Prionailurus bengalensis'') is a Felinae, small wild cat native to continental South Asia, South, Southeast Asia, Southeast, and East Asia. Since 2002 it has been listed as Least Concern on the IUCN Red List as it is widely di ...
(''Prionailurus bengalensis''). Its
dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiology ...
is relatively smaller and weaker than the jungle cat's. Both wildcat species are larger than the domestic cat. The European wildcat has relatively longer legs and a more robust build compared to the domestic cat. The tail is long, and usually slightly exceeds one-half of the animal's body length. The species size varies according to
Bergmann's rule Bergmann's rule is an ecogeographical rule that states that, within a broadly distributed taxonomic clade, populations and species of larger size are found in colder environments, while populations and species of smaller size are found in warmer ...
, with the largest specimens occurring in cool, northern areas of Europe and Asia such as
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
,
Manchuria Manchuria is a historical region in northeast Asia encompassing the entirety of present-day northeast China and parts of the modern-day Russian Far East south of the Uda (Khabarovsk Krai), Uda River and the Tukuringra-Dzhagdy Ranges. The exact ...
and
Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
. Males measure in head to body length, in tail length, and normally weigh . Females are slightly smaller, measuring in body length and in tail length, and weighing . Both sexes have two
thoracic The thorax (: thoraces or thoraxes) or chest is a part of the anatomy of mammals and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main ...
and two
abdominal The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
teat A teat is the projection from the mammary glands of mammals from which milk flows or is ejected for the purpose of feeding young. In many mammals, the teat projects from the udder. The number of teats varies by mammalian species and often corr ...
s. Both sexes have pre-
anal gland The anal glands or anal sacs are small glands near the anus in many mammals. They are situated in between the external anal sphincter muscle and internal anal sphincter muscle. In non-human mammals, the secretions of the anal glands contain most ...
s, consisting of moderately sized
sweat Perspiration, also known as sweat, is the fluid secreted by sweat glands in the skin of mammals. Two types of sweat glands can be found in humans: eccrine glands and Apocrine sweat gland, apocrine glands. The eccrine sweat glands are distribu ...
and
sebaceous gland A sebaceous gland or oil gland is a microscopic exocrine gland in the skin that opens into a hair follicle to secrete an oily or waxy matter, called sebum, which lubricates the hair and skin of mammals. In humans, sebaceous glands occur in ...
s around the anal opening. Large-sized sebaceous and
scent gland Scent gland are exocrine glands found in most mammals. They produce semi-viscous secretions which contain pheromones and other semiochemical compounds. These odor-messengers indicate information such as status, territorial marking, mood, and se ...
s extend along the full length of the tail on the dorsal side. Male wildcats have pre-anal pockets on the tail, activated upon reaching
sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized ...
, play a significant role in reproduction and
territorial marking In ethology, territory is the sociographical area that an animal consistently defends against conspecific competition (or, occasionally, against animals of other species) using agonistic behaviors or (less commonly) real physical aggression. ...
.


Distribution and habitat

The European wildcat inhabits
temperate broadleaf and mixed forest Temperate broadleaf and mixed forest is a temperate climate terrestrial habitat type defined by the World Wide Fund for Nature, with broadleaf tree ecoregions, and with conifer and broadleaf tree mixed coniferous forest ecoregions. Thes ...
s in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
,
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
and the Caucasus. In the
Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...
, it occurs from sea level to in the
Pyrenees The Pyrenees are a mountain range straddling the border of France and Spain. They extend nearly from their union with the Cantabrian Mountains to Cap de Creus on the Mediterranean coast, reaching a maximum elevation of at the peak of Aneto. ...
. Between the late 17th and mid 20th centuries, its European range became fragmented due to large-scale hunting and regional extirpation. It is possibly extinct in the
Czech Republic The Czech Republic, also known as Czechia, and historically known as Bohemia, is a landlocked country in Central Europe. The country is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the south ...
, and considered regionally extinct in
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
, though vagrants from
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
are spreading into Austria. It has never inhabited
Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and ; ), or the Fennoscandian Peninsula, is a peninsula in Europe which includes the Scandinavian Peninsula, Scandinavian and Kola Peninsula, Kola peninsulas, mainland ...
or
Estonia Estonia, officially the Republic of Estonia, is a country in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the Baltic Sea across from Sweden, to the south by Latvia, and to the east by Ru ...
.
Sicily Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...
is the only island in the
Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...
with a native wildcat population. The African wildcat lives in a wide range of habitats except
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
, but throughout the
savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
s of Africa from
Mauritania Mauritania, officially the Islamic Republic of Mauritania, is a sovereign country in Maghreb, Northwest Africa. It is bordered by the Atlantic Ocean to the west, Western Sahara to Mauritania–Western Sahara border, the north and northwest, ...
on the
Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...
coast eastward to the
Horn of Africa The Horn of Africa (HoA), also known as the Somali Peninsula, is a large peninsula and geopolitical region in East Africa.Robert Stock, ''Africa South of the Sahara, Second Edition: A Geographical Interpretation'', (The Guilford Press; 2004), ...
up to altitudes of . Small populations live in the
Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
and
Nubian Desert The Nubian Desert ( ) is in the eastern region of the Sahara, Sahara Desert, spanning approximately 400,000 km2 of northeastern Sudan and northern Eritrea, between the Nile and the Red Sea. The arid region is rugged and rocky and contains s ...
s,
Karoo The Karoo ( ; from the Afrikaans borrowing of the South Khoekhoe Khoemana (also known as !Orakobab or Korana) word is a semidesert natural region of South Africa. No exact definition of what constitutes the Karoo is available, so its extent is ...
region,
Kalahari The Kalahari Desert is a large semiarid sandy savanna in Southern Africa covering including much of Botswana as well as parts of Namibia and South Africa. It is not to be confused with the Angolan, Namibian, and South African Namib coastal d ...
and
Namib Desert The Namib ( ; ) is a coastal desert in Southern Africa. According to the broadest definition, the Namib stretches for more than along the Atlantic coasts of Angola, Namibia, and northwest South Africa, extending southward from the Carunjamba Ri ...
s. It occurs around the
Arabian Peninsula The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...
's periphery to the Caspian Sea, encompassing
Mesopotamia Mesopotamia is a historical region of West Asia situated within the Tigris–Euphrates river system, in the northern part of the Fertile Crescent. Today, Mesopotamia is known as present-day Iraq and forms the eastern geographic boundary of ...
, Israel and Palestine (region), Palestine region. In Central Asia, it ranges into Xinjiang and southern
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
, and in South Asia into the Thar Desert and arid regions in
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
.


Behaviour and ecology

Both wildcat species are largely Nocturnal animal, nocturnal and Solitary (animals), solitary, except during the breeding period and when females have young. The size of home ranges of females and males varies according to terrain, the availability of food, habitat quality and the age structure of the population. Male and female home ranges overlap, though core areas within Territory (animal), territories are avoided by other cats. Females tend to be more sedentary than males, as they require an exclusive hunting area when raising kittens. Wildcats usually spend the day in a hollow tree, a rock crevice or in dense thickets. It is also reported to shelter in abandoned burrows of other species such as of red fox (''Vulpes vulpes'') and in European badger (''Meles meles'') setts in Europe, and of fennec fox, fennec (''Vulpes zerda'') in Africa. When threatened, it retreats into a burrow, rather than climb trees. When taking residence in a tree hollow, it selects one low to the ground. Dens in rocks or burrows are lined with dry grasses and bird feathers. Dens in tree hollows usually contain enough sawdust to make lining unnecessary. If the den becomes infested with fleas, the wildcat shifts to another den. During winter, when snowfall prevents the European wildcat from travelling long distances, it remains within its den until travel conditions improve. Territorial marking consists of Urine marking#Scent marking, spraying urine on trees, vegetation and rocks, depositing faeces in conspicuous places, and leaving scent marks through glands in its paws. It also leaves visual marks by scratching trees.


Hunting and prey

Visual perception, Sight and hearing (sense), hearing are the wildcat's primary senses when hunting. It lies in wait for prey, then catches it by executing a few leaps, which can span three metres. When hunting near water courses, it waits on trees overhanging the water. It kills small prey by grabbing it in its claws, and piercing the neck or occiput with its fangs. When attacking large prey, it leaps upon the animal's back, and attempts to bite the neck or carotid. It does not persist in attacking if prey manages to escape. The European wildcat primarily preys on small mammals such as European rabbit (''Oryctolagus cuniculus'') and
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s. It also preys on dormouse, dormice, hares, nutria (''Myocastor coypus'') and birds, especially ducks and other waterfowl, galliformes, pigeons and passerines. It can consume large bone fragments. Although it kills insectivores such as mole (animal), moles and shrews, it rarely eats them. When living close to human settlements, it preys on poultry. In the wild, it consumes up to of food daily. The African wildcat preys foremost on Muridae, murids, to a lesser extent also on birds, small reptiles and invertebrates.


Reproduction and development

The wildcat has two estrus periods, one in December–February and another in May–July. Estrus lasts 5–9 days, with a gestation period lasting 60–68 days. Ovulation is induced ovulation (animals), induced through copulation. Spermatogenesis occurs throughout the year. During the mating season, males fight viciously, and may congregate around a single female. There are records of male and female wildcats becoming temporarily monogamous. Kittens are usually born between April and May, and up to August. Litter size ranges from 1–7 kittens. Kittens are born with closed eyes and are covered in a fuzzy coat. They weigh at birth, and kittens under usually do not survive. They are born with pink paw pads, which blacken at the age of three months, and blue eyes, which turn amber after five months. Their eyes open after 9–12 days, and their incisors erupt after 14–30 days. The kittens' milk teeth are replaced by their permanent dentition at the age of 160–240 days. The kittens start hunting with their mother at the age of 60 days, and start moving independently after 140–150 days. Lactation lasts 3–4 months, though the kittens eat meat as early as 1.5 months of age. Sexual maturity is attained at the age of 300 days. Similarly to the domestic cat, the physical development of African wildcat kittens over the first two weeks of their lives is much faster than that of European wildcats. The kittens are largely fully grown by 10 months, though skeletal growth continues for over 18–19 months. The family dissolves after roughly five months, and the kittens disperse to establish their own territories. Their maximum life span is 21 years, though they usually live up to 13–14 years. Generation time, Generation length of the wildcat is about eight years.


Predators and competitors

Because of its habit of living in areas with rocks and tall trees for refuge, dense thickets and abandoned burrows, wildcats have few natural predators. In Central Europe, many kittens are killed by European pine marten (''Martes martes''), and there is at least one account of an adult wildcat being killed and eaten. Competitors include the golden jackal (''Canis aureus''), red fox, marten, and other predators. In the steppe regions of Europe and Asia, village dogs constitute serious enemies of wildcats, along with the much larger Eurasian lynx, one of the rare habitual predators of healthy adult wildcats. In Tajikistan, the grey wolf (''Canis lupus'') is the most serious competitor, having been observed to destroy cat burrows. Birds of prey, including Eurasian eagle-owl (''Bubo bubo'') and saker falcon (''Falco cherrug''), have been recorded to kill wildcat kittens. Golden eagle (''Aquila chrysaetos'') are known to hunt both adults and kittens. Seton Gordon recorded an instance where a wildcat fought a golden eagle, resulting in the deaths of both combatants. In Africa, wildcats are occasionally killed and eaten by Central African rock python (''Python sebae'') and martial eagle (''Polemaetus bellicosus'').


Threats

Wildcat populations are foremost threatened by hybridization with the domestic cat. Mortality due to traffic accidents is a threat especially in Europe. The wildcat population in Scotland has declined since the turn of the 20th century due to habitat loss and persecution by landowners. In the former Soviet Union, wildcats were caught accidentally in traps set for European pine marten. In modern times, they are caught in unbaited traps on pathways or at abandoned trails of red fox, European badger, European hare or pheasant. One method of catching wildcats consists of using a modified muskrat trap with a spring placed in a concealed pit. A scent trail of pheasant viscera leads the cat to the pit. Wildcat skins were of little commercial value and sometimes converted into imitation Pinniped, sealskin; the fur usually fetched between 50 and 60 kopecks. Wildcat skins were almost solely used for making cheap scarfs, muff (handwarmer), muffs and coats for ladies.


Conservation

Wildcat species are protected in most range countries and listed in CITES Appendix II. The European wildcat is also listed in Appendix II of the Berne Convention on the Conservation of European Wildlife and Natural Habitats and in the European Union's Habitats Directive, Habitats and Species Directive. Conservation Action Plans have been developed in Germany and Scotland.


In culture


Domestication

An African wildcat skeleton Excavation (archaeology), excavated in a 9,500-year-old Neolithic grave in Cyprus is the earliest known indication for a close relationship between a human and a possibly tamed cat. As no cat species is native to Cyprus, this discovery indicates that Neolithic farmers may have brought cats to Cyprus from the Near East. Results of genetics and morphology (biology), morphological research corroborated that the African wildcat is the ancestor of the domestic cat. The first individuals were probably domesticated in the Fertile Crescent around the time of the introduction of agriculture. Murals and statuettes depicting cats as well Mummification in Ancient Egypt, mummified cats indicate that it was commonly kept by ancient Egyptians since at least the Twelfth Dynasty of Egypt.


In mythology

Celtic fables of the Cat Sìth, a fairy creature described as resembling a large white-chested black cat, are thought to have been inspired by the Kellas cat, itself thought to be a free-ranging crossbreed between a European wildcat and a domestic cat. In 1693, William Salmon mentioned how body parts of the wildcat were used for medicinal purposes; its flesh for treating gout, its fat for dissolving tumours and easing pain, its blood for curing "Epilepsy, falling sickness", and its excrement for treating baldness.


In heraldry

The Picts venerated wildcats, having probably named Caithness (Land of the Cats) after them. According to the foundation myth of the Catti tribe, their ancestors were attacked by wildcats upon landing in Scotland. Their ferocity impressed the Catti so much, that the wildcat became their symbol. The progenitors of Clan Sutherland use the wildcat as symbol on their family crest. The clan's chief bears the title ''Morair Chat'' (Great Man of the Cats). The wildcat is considered an Cultural icon, icon of Scottish wilderness, and has been used in clan heraldry since the 13th century. The Clan Chattan Association (also known as the Clan of Cats) comprises 12 clans, the majority of which display the wildcat on their badges.


In literature

Shakespeare referenced the wildcat three times:


References


Sources

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Further reading

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External links

* * * * * * * * Scottish wildcat ** ** {{Authority control Felis Mammals described in 1777 Mammals of Africa Mammals of Asia Mammals of Europe Mammals of Azerbaijan Mammals of Pakistan Mammals of the Middle East Mammals of Uganda