The wildcat is a

species complex In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...

comprising two small wild cat

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

: the European wildcat (''Felis silvestris'') and the African wildcat (''F. lybica''). The European wildcat inhabits

forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...

s in

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

Anatolia Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...

and the

Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...

, while the African wildcat inhabits semi- arid landscapes and

steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropica ...

s in

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

, the

Arabian Peninsula The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...

Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...

, into western

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

and western

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

. The wildcat species differ in fur pattern, tail, and size: the European wildcat has long fur and a bushy tail with a rounded tip; the smaller African wildcat is more faintly striped, has short sandy-gray fur and a tapering tail; the Asiatic wildcat (''F. lybica ornata'') is spotted. The wildcat and the other members of the cat family had a

common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. According to modern evolutionary biology, all living beings could be descendants of a unique ancestor commonl ...

about 10–15 million years ago. The European wildcat evolved during the Cromerian Stage about 866,000 to 478,000 years ago; its direct ancestor was '' Felis lunensis''. The ''silvestris'' and ''lybica'' lineages probably diverged about 173,000 years ago. The wildcat is categorized as

Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...

on the

IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...

since 2002, since it is widely distributed in a stable global population exceeding 20,000 mature individuals. Some local populations are threatened by introgressive hybridisation with the

domestic cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small Domestication, domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have sh ...

(''F. catus''), contagious disease, vehicle collisions and persecution. The association of African wildcats and humans appears to have developed along with the establishment of settlements during the

Neolithic Revolution The Neolithic Revolution, also known as the First Agricultural Revolution, was the wide-scale transition of many human cultures during the Neolithic period in Afro-Eurasia from a lifestyle of hunter-gatherer, hunting and gathering to one of a ...

, when

rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...

s in grain stores of early

farmer A farmer is a person engaged in agriculture, raising living organisms for food or raw materials. The term usually applies to people who do some combination of raising field crops, orchards, vineyards, poultry, or other livestock. A farmer ...

s attracted wildcats. This association ultimately led to it being

tamed A tame animal is an animal that is relatively tolerant of human presence. Tameness may arise naturally (as in the case, for example, of island tameness) or due to the deliberate, human-directed process of animal training, training an animal agai ...

and domesticated: the domestic cat is the direct descendant of the African wildcat. It was one of the revered

cats in ancient Egypt In ancient Egypt, cats were represented in social and religious scenes dating as early as 1980 BC. Several ancient Egyptian deities were depicted and sculptured with cat-like heads such as Mafdet, Bastet and Sekhmet, representing justice, fertili ...

. The European wildcat has been the subject of

mythology Myth is a genre of folklore consisting primarily of narratives that play a fundamental role in a society. For scholars, this is very different from the vernacular usage of the term "myth" that refers to a belief that is not true. Instead, the ...

and

literature Literature is any collection of Writing, written work, but it is also used more narrowly for writings specifically considered to be an art form, especially novels, Play (theatre), plays, and poetry, poems. It includes both print and Electroni ...

Taxonomy

''Felis (catus) silvestris'' was the

scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...

used in 1777 by Johann von Schreber when he described the European wildcat based on descriptions and names proposed by earlier naturalists such as

Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...

, Ulisse Aldrovandi and Conrad Gessner. ''Felis lybica'' was the name proposed in 1780 by

Georg Forster Johann George Adam Forster, also known as Georg Forster (; 27 November 1754 – 10 January 1794), was a German geography, geographer, natural history, naturalist, ethnology, ethnologist, travel literature, travel writer, journalist and revol ...

, who described an African wildcat from Gafsa on the Barbary Coast. In subsequent decades, several naturalists and explorers described 40 wildcat specimens collected in European, African and Asian range countries. In the 1940s, the taxonomist

Reginald Innes Pocock Reginald Innes Pocock, (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock and Edith Prichard. He began showing interest in natural history at St. Edward's ...

reviewed the collection of wildcat skins and skulls in the

Natural History Museum, London The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum (Lo ...

, and designated seven ''F. silvestris''

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

from Europe to

Asia Minor Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...

, and 25 ''F. lybica'' subspecies from

, and West to

. Pocock differentiated the: * Forest wildcat subspecies (''silvestris'' group) * Steppe wildcat subspecies (''ornata''-''caudata'' group): is distinguished from the forest wildcat by being smaller, with comparatively lighter fur colour, and longer and more sharply-pointed tails. The domestic cat is thought to have derived from this group. * Bush wildcat subspecies (''ornata''-''lybica'' group): is distinguished from the steppe wildcat by paler fur, well-developed spot patterns and bands. In 2005, 22 subspecies were recognized by the authors of ''

Mammal Species of the World ''Mammal Species of the World: A Taxonomic and Geographic Reference'' is a standard reference work in mammalogy giving descriptions and Bibliographic database, bibliographic data for the known species of mammals. It is now in its third edition, ...

'', who allocated subspecies largely in line with Pocock's assessment. In 2006, the Chinese mountain cat was placed within the wildcat lineage, being found to be more closely related to the Asiatic wildcat and the domestic cat than to the European wildcat through nuclear DNA, while being placed as an outgroup to the wildcat clade through mitochondrial DNA. In 2017, the Cat Classification Task Force revised the

taxonomy image:Hierarchical clustering diagram.png, 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme o ...

of the Felidae, and recognized the following as valid taxa:

Evolution

The wildcat is a member of the Felidae, a family that had a

about 10–15 million years ago. ''Felis'' species diverged from the Felidae around 6–7 million years ago. The European wildcat diverged from ''Felis'' about 1.09 to 1.4 million years ago. The European wildcat's direct ancestor was '' Felis lunensis'', which lived in Europe in the late

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Villafranchian periods.

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

remains indicate that the transition from ''lunensis'' to ''silvestris'' was completed by the Holstein interglacial about 340,000 to 325,000 years ago. Craniological differences between the European and African wildcats indicate that the wildcat probably migrated during the

Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...

from Europe into the Middle East, giving rise to the steppe wildcat

phenotype In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology (physical form and structure), its developmental processes, its biochemical and physiological propert ...

Phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

research revealed that the ''lybica'' lineage probably diverged from the ''silvestris'' lineage about 173,000 years ago.

Characteristics

The wildcat has pointed ears, which are moderate in length and broad at the base. Its whiskers are white, number 7 to 16 on each side and reach in length on the muzzle. Whiskers are also present on the inner surface of the paw and measure . Its eyes are large, with vertical pupils and yellowish-green irises. The eyelashes range from in length, and can number six to eight per side. The European wildcat has a greater skull volume than the domestic cat, a ratio known as Schauenberg's index. Further, its skull is more spherical in shape than that of the jungle cat (''F. chaus'') and leopard cat (''Prionailurus bengalensis''). Its

dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiology ...

is relatively smaller and weaker than the jungle cat's. Both wildcat species are larger than the domestic cat. The European wildcat has relatively longer legs and a more robust build compared to the domestic cat. The tail is long, and usually slightly exceeds one-half of the animal's body length. The species size varies according to Bergmann's rule, with the largest specimens occurring in cool, northern areas of Europe and Asia such as

Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...

Manchuria Manchuria is a historical region in northeast Asia encompassing the entirety of present-day northeast China and parts of the modern-day Russian Far East south of the Uda (Khabarovsk Krai), Uda River and the Tukuringra-Dzhagdy Ranges. The exact ...

and

Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...

. Males measure in head to body length, in tail length, and normally weigh . Females are slightly smaller, measuring in body length and in tail length, and weighing . Both sexes have two

thoracic The thorax (: thoraces or thoraxes) or chest is a part of the anatomy of mammals and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main ...

and two abdominal teats. Both sexes have pre- anal glands, consisting of moderately sized

sweat Perspiration, also known as sweat, is the fluid secreted by sweat glands in the skin of mammals. Two types of sweat glands can be found in humans: eccrine glands and Apocrine sweat gland, apocrine glands. The eccrine sweat glands are distribu ...

and

sebaceous gland A sebaceous gland or oil gland is a microscopic exocrine gland in the skin that opens into a hair follicle to secrete an oily or waxy matter, called sebum, which lubricates the hair and skin of mammals. In humans, sebaceous glands occur in ...

s around the anal opening. Large-sized sebaceous and scent glands extend along the full length of the tail on the dorsal side. Male wildcats have pre-anal pockets on the tail, activated upon reaching sexual maturity, play a significant role in reproduction and territorial marking.

Distribution and habitat

The European wildcat inhabits

temperate broadleaf and mixed forest Temperate broadleaf and mixed forest is a temperate climate terrestrial habitat type defined by the World Wide Fund for Nature, with broadleaf tree ecoregions, and with conifer and broadleaf tree mixed coniferous forest ecoregions. Thes ...

s in

Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...

and the Caucasus. In the

Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...

, it occurs from sea level to in the Pyrenees. Between the late 17th and mid 20th centuries, its European range became fragmented due to large-scale hunting and regional extirpation. It is possibly extinct in the

Czech Republic The Czech Republic, also known as Czechia, and historically known as Bohemia, is a landlocked country in Central Europe. The country is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the south ...

, and considered regionally extinct in

Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...

, though vagrants from

Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...

are spreading into Austria. It has never inhabited

Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and ; ), or the Fennoscandian Peninsula, is a peninsula in Europe which includes the Scandinavian Peninsula, Scandinavian and Kola Peninsula, Kola peninsulas, mainland ...

Estonia Estonia, officially the Republic of Estonia, is a country in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the Baltic Sea across from Sweden, to the south by Latvia, and to the east by Ru ...

Sicily Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...

is the only island in the

Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...

with a native wildcat population. The African wildcat lives in a wide range of habitats except

rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...

, but throughout the savannahs of Africa from

Mauritania Mauritania, officially the Islamic Republic of Mauritania, is a sovereign country in Maghreb, Northwest Africa. It is bordered by the Atlantic Ocean to the west, Western Sahara to Mauritania–Western Sahara border, the north and northwest, ...

on the

Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...

coast eastward to the

Horn of Africa The Horn of Africa (HoA), also known as the Somali Peninsula, is a large peninsula and geopolitical region in East Africa.Robert Stock, ''Africa South of the Sahara, Second Edition: A Geographical Interpretation'', (The Guilford Press; 2004), ...

up to altitudes of . Small populations live in the

Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...

and Nubian Deserts, Karoo region, Kalahari and Namib Deserts. It occurs around the

's periphery to the Caspian Sea, encompassing

Mesopotamia Mesopotamia is a historical region of West Asia situated within the Tigris–Euphrates river system, in the northern part of the Fertile Crescent. Today, Mesopotamia is known as present-day Iraq and forms the eastern geographic boundary of ...

Israel Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Isr ...

and Palestine region. In Central Asia, it ranges into

Xinjiang Xinjiang,; , SASM/GNC romanization, SASM/GNC: Chinese postal romanization, previously romanized as Sinkiang, officially the Xinjiang Uygur Autonomous Region (XUAR), is an Autonomous regions of China, autonomous region of the China, People' ...

and southern

, and in

South Asia South Asia is the southern Subregion#Asia, subregion of Asia that is defined in both geographical and Ethnicity, ethnic-Culture, cultural terms. South Asia, with a population of 2.04 billion, contains a quarter (25%) of the world's populatio ...

into the Thar Desert and arid regions in

Behaviour and ecology

Both wildcat species are largely

nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...

and solitary, except during the breeding period and when females have young. The size of

home range A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. ...

s of females and males varies according to terrain, the availability of food, habitat quality and the age structure of the population. Male and female home ranges overlap, though core areas within territories are avoided by other cats. Females tend to be more sedentary than males, as they require an exclusive hunting area when raising kittens. Wildcats usually spend the day in a hollow tree, a rock crevice or in dense thickets. It is also reported to shelter in abandoned

burrow file:Chipmunk-burrow (exits).jpg, An eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of Animal lo ...

s of other species such as of

red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe and Asia, plus ...

(''Vulpes vulpes'') and in European badger (''Meles meles'') setts in Europe, and of fennec (''Vulpes zerda'') in Africa. When threatened, it retreats into a burrow, rather than climb trees. When taking residence in a tree hollow, it selects one low to the ground. Dens in rocks or burrows are lined with dry grasses and bird

feather Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and an exa ...

s. Dens in tree hollows usually contain enough sawdust to make lining unnecessary. If the den becomes infested with

flea Flea, the common name for the order (biology), order Siphonaptera, includes 2,500 species of small flightless insects that live as external parasites of mammals and birds. Fleas live by hematophagy, ingesting the blood of their hosts. Adult f ...

s, the wildcat shifts to another den. During winter, when snowfall prevents the European wildcat from travelling long distances, it remains within its den until travel conditions improve. Territorial marking consists of spraying urine on trees, vegetation and rocks, depositing faeces in conspicuous places, and leaving scent marks through glands in its paws. It also leaves visual marks by scratching trees.

Hunting and prey

Sight Visual perception is the ability to detect light and use it to form an image of the surrounding Biophysical environment, environment. Photodetection without image formation is classified as ''light sensing''. In most vertebrates, visual percept ...

and

hearing Hearing, or auditory perception, is the ability to perceive sounds through an organ, such as an ear, by detecting vibrations as periodic changes in the pressure of a surrounding medium. The academic field concerned with hearing is auditory sci ...

are the wildcat's primary senses when hunting. It lies in wait for prey, then catches it by executing a few leaps, which can span three metres. When hunting near water courses, it waits on trees overhanging the water. It kills small prey by grabbing it in its claws, and piercing the neck or

occiput The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone lies over the occipital lobes of the ...

with its fangs. When attacking large prey, it leaps upon the animal's back, and attempts to bite the neck or carotid. It does not persist in attacking if prey manages to escape. The European wildcat primarily preys on small mammals such as European rabbit (''Oryctolagus cuniculus'') and

s. It also preys on

dormice A dormouse is a rodent of the family (biology), family Gliridae (this family is also variously called Myoxidae or Muscardinidae by different taxonomists). Dormice are nocturnal animals found in Africa, Asia, and Europe. They are named for their ...

hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores and live Solitary animal, solitarily or in pairs. They nest in slight depressions called forms, and their young are precociality, able to fend for themselves ...

nutria The nutria () or coypu () (''Myocastor coypus'') is a herbivore, herbivorous, semiaquatic rodent from South America. Classified for a long time as the only member of the family Myocastoridae, ''Myocastor'' has since been included within Echimy ...

(''Myocastor coypus'') and

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s, especially

duck Duck is the common name for numerous species of waterfowl in the family (biology), family Anatidae. Ducks are generally smaller and shorter-necked than swans and goose, geese, which are members of the same family. Divided among several subfam ...

s and other waterfowl,

galliformes Galliformes is an order (biology), order of heavy-bodied ground-feeding birds that includes turkey (bird), turkeys, chickens, Old World quail, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems ...

pigeon Columbidae is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with small heads, relatively short necks and slender bills that in some species feature fleshy ceres. ...

s and passerines. It can consume large

bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...

fragments. Although it kills

insectivore file:Common brown robberfly with prey.jpg, A Asilidae, robber fly eating a hoverfly An insectivore is a carnivore, carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the Entomophagy ...

s such as moles and

shrew Shrews ( family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to dif ...

s, it rarely eats them. When living close to human settlements, it preys on

poultry Poultry () are domesticated birds kept by humans for the purpose of harvesting animal products such as meat, Eggs as food, eggs or feathers. The practice of animal husbandry, raising poultry is known as poultry farming. These birds are most typ ...

. In the wild, it consumes up to of food daily. The African wildcat preys foremost on murids, to a lesser extent also on birds, small reptiles and

invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...

Reproduction and development

The wildcat has two

estrus The estrous cycle (, originally ) is a set of recurring physiological changes induced by reproductive hormones in females of mammalian subclass Theria. Estrous cycles start after sexual maturity in females and are interrupted by anestrous phas ...

periods, one in December–February and another in May–July. Estrus lasts 5–9 days, with a gestation period lasting 60–68 days.

Ovulation Ovulation is an important part of the menstrual cycle in female vertebrates where the egg cells are released from the ovaries as part of the ovarian cycle. In female humans ovulation typically occurs near the midpoint in the menstrual cycle and ...

is induced through copulation. Spermatogenesis occurs throughout the year. During the mating season, males fight viciously, and may congregate around a single female. There are records of male and female wildcats becoming temporarily monogamous. Kittens are usually born between April and May, and up to August. Litter size ranges from 1–7 kittens. Kittens are born with closed eyes and are covered in a fuzzy coat. They weigh at birth, and kittens under usually do not survive. They are born with pink paw pads, which blacken at the age of three months, and blue eyes, which turn amber after five months. Their eyes open after 9–12 days, and their incisors erupt after 14–30 days. The kittens' milk teeth are replaced by their permanent dentition at the age of 160–240 days. The kittens start hunting with their mother at the age of 60 days, and start moving independently after 140–150 days.

Lactation Lactation describes the secretion of milk from the mammary glands and the period of time that a mother lactates to feed her young. The process naturally occurs with all sexually mature female mammals, although it may predate mammals. The process ...

lasts 3–4 months, though the kittens eat meat as early as 1.5 months of age. Sexual maturity is attained at the age of 300 days. Similarly to the domestic cat, the physical development of African wildcat kittens over the first two weeks of their lives is much faster than that of European wildcats. The kittens are largely fully grown by 10 months, though skeletal growth continues for over 18–19 months. The family dissolves after roughly five months, and the kittens disperse to establish their own territories. Their maximum life span is 21 years, though they usually live up to 13–14 years. Generation length of the wildcat is about eight years.

Predators and competitors

Because of its habit of living in areas with rocks and tall trees for refuge, dense thickets and abandoned burrows, wildcats have few natural predators. In Central Europe, many kittens are killed by European pine marten (''Martes martes''), and there is at least one account of an adult wildcat being killed and eaten. Competitors include the golden jackal (''Canis aureus''), red fox, marten, and other predators. In the steppe regions of Europe and Asia, village dogs constitute serious enemies of wildcats, along with the much larger

Eurasian lynx The Eurasian lynx (''Lynx lynx'') is one of the four wikt:extant, extant species within the medium-sized wild Felidae, cat genus ''Lynx''. It is widely distributed from Northern Europe, Northern, Central Europe, Central and Eastern Europe to Cent ...

, one of the rare habitual predators of healthy adult wildcats. In Tajikistan, the

grey wolf The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, including the dog and dingo, though gr ...

(''Canis lupus'') is the most serious competitor, having been observed to destroy cat burrows.

Birds of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...

, including

Eurasian eagle-owl The Eurasian eagle-owl (''Bubo bubo'') is a species of eagle-owl, a type of bird that resides in much of Eurasia. It is often just called the eagle-owl in Europe and Asia. It is one of the largest species of owl. Females can grow to a total le ...

(''Bubo bubo'') and saker falcon (''Falco cherrug''), have been recorded to kill wildcat kittens.

Golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of pr ...

(''Aquila chrysaetos'') are known to hunt both adults and kittens. Seton Gordon recorded an instance where a wildcat fought a golden eagle, resulting in the deaths of both combatants. In Africa, wildcats are occasionally killed and eaten by Central African rock python (''Python sebae'') and martial eagle (''Polemaetus bellicosus'').

Threats

Wildcat populations are foremost threatened by hybridization with the domestic cat. Mortality due to traffic accidents is a threat especially in Europe. The wildcat population in Scotland has declined since the turn of the 20th century due to

habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...

and persecution by landowners. In the former Soviet Union, wildcats were caught accidentally in traps set for European pine marten. In modern times, they are caught in unbaited traps on pathways or at abandoned trails of red fox, European badger, European hare or pheasant. One method of catching wildcats consists of using a modified muskrat trap with a spring placed in a concealed pit. A scent trail of pheasant viscera leads the cat to the pit. Wildcat skins were of little commercial value and sometimes converted into imitation sealskin; the fur usually fetched between 50 and 60 kopecks. Wildcat skins were almost solely used for making cheap scarfs, muffs and coats for ladies.

Conservation

Wildcat species are protected in most range countries and listed in CITES Appendix II. The European wildcat is also listed in Appendix II of the Berne Convention on the Conservation of European Wildlife and Natural Habitats and in the

European Union The European Union (EU) is a supranational union, supranational political union, political and economic union of Member state of the European Union, member states that are Geography of the European Union, located primarily in Europe. The u ...

's Habitats and Species Directive. Conservation Action Plans have been developed in Germany and Scotland.

In culture

Domestication

An African wildcat skeleton excavated in a 9,500-year-old Neolithic grave in Cyprus is the earliest known indication for a close relationship between a human and a possibly tamed cat. As no cat species is native to Cyprus, this discovery indicates that Neolithic farmers may have brought cats to Cyprus from the Near East. Results of

genetics Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinians, Augustinian ...

and morphological research corroborated that the African wildcat is the ancestor of the domestic cat. The first individuals were probably domesticated in the

Fertile Crescent The Fertile Crescent () is a crescent-shaped region in the Middle East, spanning modern-day Iraq, Israel, Jordan, Lebanon, Palestine, and Syria, together with northern Kuwait, south-eastern Turkey, and western Iran. Some authors also include ...

around the time of the introduction of agriculture.

Mural A mural is any piece of Graphic arts, graphic artwork that is painted or applied directly to a wall, ceiling or other permanent substrate. Mural techniques include fresco, mosaic, graffiti and marouflage. Word mural in art The word ''mural'' ...

s and statuettes depicting cats as well mummified cats indicate that it was commonly kept by ancient Egyptians since at least the

Twelfth Dynasty of Egypt The Twelfth Dynasty of ancient Egypt (Dynasty XII) is a series of rulers reigning from 1991–1802 BC (190 years), at what is often considered to be the apex of the Middle Kingdom of Egypt, Middle Kingdom (Dynasties XI–XIV). The dynasty period ...

In mythology

Celtic fables of the Cat Sìth, a fairy creature described as resembling a large white-chested black cat, are thought to have been inspired by the Kellas cat, itself thought to be a free-ranging crossbreed between a European wildcat and a domestic cat. In 1693, William Salmon mentioned how body parts of the wildcat were used for medicinal purposes; its flesh for treating

gout Gout ( ) is a form of inflammatory arthritis characterized by recurrent attacks of pain in a red, tender, hot, and Joint effusion, swollen joint, caused by the deposition of needle-like crystals of uric acid known as monosodium urate crysta ...

, its

fat In nutrition science, nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such chemical compound, compounds, most commonly those that occur in living beings or in food. The term often refers specif ...

for dissolving

tumour A neoplasm () is a type of abnormal and excessive growth of tissue (biology), tissue. The process that occurs to form or produce a neoplasm is called neoplasia. The growth of a neoplasm is uncoordinated with that of the normal surrounding tiss ...

s and easing pain, its blood for curing " falling sickness", and its excrement for treating baldness.

In heraldry

The

Picts The Picts were a group of peoples in what is now Scotland north of the Firth of Forth, in the Scotland in the early Middle Ages, Early Middle Ages. Where they lived and details of their culture can be gleaned from early medieval texts and Pic ...

venerated wildcats, having probably named

Caithness Caithness (; ; ) is a Shires of Scotland, historic county, registration county and Lieutenancy areas of Scotland, lieutenancy area of Scotland. There are two towns, being Wick, Caithness, Wick, which was the county town, and Thurso. The count ...

(Land of the Cats) after them. According to the foundation myth of the Catti tribe, their ancestors were attacked by wildcats upon landing in Scotland. Their ferocity impressed the Catti so much, that the wildcat became their symbol. The progenitors of Clan Sutherland use the wildcat as symbol on their family crest. The clan's chief bears the title ''Morair Chat'' (Great Man of the Cats). The wildcat is considered an

icon An icon () is a religious work of art, most commonly a painting, in the cultures of the Eastern Orthodox, Oriental Orthodox, Catholic Church, Catholic, and Lutheranism, Lutheran churches. The most common subjects include Jesus, Mary, mother of ...

of Scottish wilderness, and has been used in clan heraldry since the 13th century. The Clan Chattan Association (also known as the Clan of Cats) comprises 12 clans, the majority of which display the wildcat on their badges.

In literature

Shakespeare William Shakespeare ( 23 April 1564 – 23 April 1616) was an English playwright, poet and actor. He is widely regarded as the greatest writer in the English language and the world's pre-eminent dramatist. He is often called England's natio ...

referenced the wildcat three times:

References

Sources

* * * * * *

External links

* * * * * * * * Scottish wildcat ** ** {{Authority control Felis Mammals described in 1777 Mammals of Africa Mammals of Asia Mammals of Europe Mammals of Azerbaijan Mammals of Pakistan Mammals of the Middle East Mammals of Uganda

Taxonomy

Evolution

Characteristics

Distribution and habitat

Behaviour and ecology

Hunting and prey

Reproduction and development

Predators and competitors

Threats

Conservation

In culture

Domestication

In mythology

In heraldry

In literature

References

Sources

Further reading

External links