
Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon
) is a term used by some
taxonomist
In biology, taxonomy () is the science, scientific study of naming, defining (Circumscription (taxonomy), circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxon, taxa (si ...
s to refer to a
taxon
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
that has the purpose of classifying organisms that do not fit anywhere else. They are typically defined by either their designated members' often superficial similarity to each other, or their ''lack'' of one or more distinct
character states or by their ''not'' belonging to one or more other taxa. Wastebasket taxa are by definition either
paraphyletic
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
or
polyphyletic
A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...
, and are therefore not considered valid taxa under strict
cladistic
Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...
rules of taxonomy. The name of a wastebasket taxon may in some cases be retained as the designation of an
evolutionary grade
A grade is a taxon united by a level of morphological or physiological complexity. The term was coined by British biologist Julian Huxley, to contrast with clade, a strictly phylogenetic unit.
Phylogenetics
The concept of evolutionary grades ...
, however.
Examples
There are many examples of paraphyletic groups, but true "wastebasket" taxa are those that are known not to, and perhaps not intended to, represent natural groups, but are nevertheless used as convenient groups of organisms. The
acritarchs
Acritarchs () are organic microfossils, known from the Precambrian to the present. The classification is a catch all term used to refer to any organic microfossils that cannot be assigned to other groups. Their diversity reflects major ecologi ...
are perhaps the most famous example. Wastebasket taxa are often old (and perhaps not described with the systematic rigour and precision that is possible in the light of accumulated knowledge of diversity) and populous.
* The
Flacourtiaceae
The Flacourtiaceae is a defunct family of flowering plants whose former members have been scattered to various families, mostly to the Achariaceae and Salicaceae. It was so vaguely defined that hardly anything seemed out of place there and it bec ...
, a now-defunct family of
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s
– the
Angiosperm Phylogeny Group
The Angiosperm Phylogeny Group (APG) is an informal international group of systematic botanists who collaborate to establish a consensus on the taxonomy of flowering plants (angiosperms) that reflects new knowledge about plant relationships disc ...
has placed its
tribe
The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
s and genera in various other families, especially the
Achariaceae
Achariaceae is a family of flowering plants consisting of 31 genera and about 155 species of tropical herbs, shrubs, and trees. The APG IV system has greatly expanded the scope of the family by including many genera previously classified in Flac ...
and
Salicaceae
The Salicaceae are the willow family of flowering plants. The traditional family (Salicaceae ''sensu stricto'') includes the willows, poplars. Genetic studies summarized by the Angiosperm Phylogeny Group (APG) have greatly expanded the circumsc ...
.
* The obsolete kingdom
Protista
A protist ( ) or protoctist is any Eukaryote, eukaryotic organism that is not an animal, Embryophyte, land plant, or fungus. Protists do not form a Clade, natural group, or clade, but are a Paraphyly, paraphyletic grouping of all descendants o ...
is composed of all
eukaryote
The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms ...
s that are not
animal
Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s,
plant
Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
s or
fungi
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
, leaving to the protists all single-celled eukaryotes.
* The
Tricholomataceae
The Tricholomataceae are a large family of fungi within the order Agaricales. Originally a classic "wastebasket taxon", the family included any white-, yellow-, or pink-spored genera in the Agaricales not already classified as belonging to e.g. t ...
is a fungal group, at one point composed of the white-, yellow-, or pink-spored genera in the
Agaricales
The Agaricales are an order (biology), order of fungi in the division (mycology), division Basidiomycota. As originally conceived, the order contained all the agarics (gilled mushrooms), but subsequent research has shown that not all agarics are ...
not already classified as belonging to the
Amanitaceae
The Amanitaceae is a family of mushroom-forming fungi. ''Amanita'' Pers. is one of the most speciose and best-known fungal genera. The family, also commonly called the amanita family, is in order Agaricales, the gilled mushrooms. The family con ...
,
Lepiotaceae
The Agaricaceae are a family of basidiomycete fungi and include the genus ''Agaricus'', as well as basidiomycetes previously classified in the families Tulostomataceae, Lepiotaceae, and Lycoperdaceae.
Taxonomy
The family Agaricaceae was publishe ...
,
Hygrophoraceae
The Hygrophoraceae are a family (biology), family of fungi in the order (biology), order Agaricales. Originally conceived as containing white-spored, thick-gilled agarics (gilled mushrooms), including ''Hygrophorus'' and ''Hygrocybe'' species (th ...
,
Pluteaceae
The Pluteaceae are a family of small to medium-sized mushrooms which have free gill attachment and pink spores. Members of Pluteaceae can be mistaken for members of Entolomataceae, but can be distinguished by the angled spores and attached gills ...
, or
Entolomataceae
The Entolomataceae are a family of fungi in the order Agaricales. The family contains eight genera and 2250 species, the majority of which are in ''Entoloma''. Basidiocarps (fruit bodies) are typically agaricoid (mushrooms with gills), but a mino ...
.
*
Carnosauria
Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods.
While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Ca ...
and
Thecodontia
Thecodontia (meaning 'socket-teeth'), now considered an obsolete taxonomic grouping, was formerly used to describe a diverse "order" of early archosaurian reptiles that first appeared in the latest Permian period and flourished until the end of t ...
are fossil groups, banded together back when the limited fossil record did not allow for a more detailed scheme.
*
Condylarthra
Condylarthra is an informal group – previously considered an order – of extinct placental mammals, known primarily from the Paleocene and Eocene epochs. They are considered early, primitive ungulates and is now largely considered to be a wast ...
is an artificial clade into which ungulate mammals not clearly within
Perissodactyla
Perissodactyla (, ), or odd-toed ungulates, is an order of ungulates. The order includes about 17 living species divided into three families: Equidae (horses, asses, and zebras), Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They t ...
or
Cetartiodactyla
Artiodactyls are placental mammals belonging to the order (biology), order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof ...
were traditionally shoved. Many of these groups, like
Meridiungulata
South American native ungulates, commonly abbreviated as SANUs, are extinct ungulate-like mammals that were indigenous to South America from the Paleocene (from at least 63 million years ago) until the end of the Late Pleistocene (~12,000 years a ...
or ''
Protungulatum
''Protungulatum'' ('first ungulate') is an extinct genus of eutherian mammals within extinct family Protungulatidae, and is possibly one of the earliest known placental mammals in the fossil record, that lived in North America from the Late Cre ...
'', may not represent
laurasitherian mammals, while others like
phenacodontids have been clearly established as early odd-toed ungulates.
* The order
Insectivora
The Order (biology), order Insectivora (from Latin ''insectum'' "insect" and ''vorare'' "to eat") is a now-abandoned biological grouping within the class of mammals. Some species have now been moved out, leaving the remaining ones in the order ...
has traditionally been used as a dumping ground for placental insectivorous mammals (and similar forms such as
colugo
Colugos (), flying lemurs, or cobegos (), are arboreal gliding euarchontogliran mammals that are native to Southeast Asia. Their closest evolutionary relatives are primates. There are just two living species of colugos: the Sunda flying lemur ...
s), usually aligned with
carnivoran
Carnivora ( ) is an order of placental mammals specialized primarily in eating flesh, whose members are formally referred to as carnivorans. The order Carnivora is the sixth largest order of mammals, comprising at least 279 species. Carnivor ...
s,
ungulate
Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
s and
bat
Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...
s. While the core components (
mole
Mole (or Molé) may refer to:
Animals
* Mole (animal) or "true mole"
* Golden mole, southern African mammals
* Marsupial mole
Marsupial moles, the Notoryctidae family, are two species of highly specialized marsupial mammals that are found i ...
s,
shrew
Shrews ( family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to dif ...
s,
hedgehog
A hedgehog is a spiny mammal of the subfamily Erinaceinae, in the eulipotyphlan family Erinaceidae. There are 17 species of hedgehog in five genera found throughout parts of Europe, Asia, and Africa, and in New Zealand by introduction. The ...
s and their close relations) do in fact form a consistent clade,
Eulipotyphla
Eulipotyphla (, from '' eu-'' + '' Lipotyphla'', meaning truly lacking blind gut; sometimes called true insectivores) is an order of mammals comprising the Erinaceidae ( hedgehogs and gymnures); Solenodontidae (solenodons); Talpidae ( mole ...
, that is part of
Laurasiatheria
Laurasiatheria (; "Laurasian beasts") is a superorder of Placentalia, placental mammals that groups together true insectivores (eulipotyphlans), bats (chiropterans), carnivorans, pangolins (Pholidota, pholidotes), even-toed ungulates (Artiodacty ...
with the aforementioned clades, other mammals historically placed in the order have been found to belong to other branches of the placental tree:
tree shrew
The treeshrews (also called tree shrews or banxrings) are small mammals native to the tropical forests of South and Southeast Asia. They make up the entire Order (biology), order Scandentia (from Latin ''scandere'', "to climb"), which split into ...
s and colugos are
euarchonta
The Euarchonta are a proposed grandorder of mammals: the order Scandentia (treeshrews), and its sister Primatomorpha mirorder, containing the Dermoptera (colugos) and the primates (Plesiadapiformes and descendants).
The term "Euarchonta" (mea ...
ns related to
Primates
Primates is an order of mammals, which is further divided into the strepsirrhines, which include lemurs, galagos, and lorisids; and the haplorhines, which include tarsiers and simians ( monkeys and apes). Primates arose 74–63 ...
and sometimes grouped in
Sundatheria, while
tenrec
A tenrec () is a mammal belonging to any species within the afrotherian family Tenrecidae, which is endemic to Madagascar. Tenrecs are a very diverse group, as a result of adaptive radiation, and exhibit convergent evolution, some resemble hed ...
s,
golden mole
Golden moles are small insectivorous burrowing mammals endemic to Sub-Saharan Africa. They comprise the family Chrysochloridae (the only family in the suborder Chrysochloridea) and as such they are taxonomically distinct from the true moles, f ...
s and
elephant shrew
Elephant shrews, also called jumping shrews or sengis, are small insectivore, insectivorous mammals native to Africa, belonging to the family Macroscelididae, in the order (biology), order Macroscelidea. Their traditional common English name "ele ...
s are all
afrotheres, probably forming the clade
Afroinsectiphilia
The Afroinsectiphilia (African insectivores) is a clade that has been proposed based on the results of recent molecular phylogenetic studies. Many of the taxa within it were once regarded as part of the order Insectivora, but Insectivora is now ...
. Both of these clades have at times been accused of being wastebasket taxa themselves, grouping superficially similar animals in Euarchonta and Afrotheria, respectively, but they have been more strongly supported by genetic studies.
*
Vermes
Vermes ("vermin/vermes") is an obsolete taxon used by Carl Linnaeus and Jean-Baptiste Lamarck for non-arthropod invertebrate animals.
Linnaeus
In Linnaeus's ''Systema Naturae'', the Vermes had the rank of class (biology), class, occupying th ...
is an obsolete taxon of worm-like animals. It was a catch-all term used by
Carl Linnaeus
Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
and
Jean-Baptiste Lamarck
Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biologi ...
for non-arthropod invertebrate animals.
*The genus ''
Mamenchisaurus
''Mamenchisaurus'' ( , or spelling pronunciation ) is an extinct genus of sauropod dinosaurs known for their remarkably long necks which made up nearly half the total body length. Numerous species have been assigned to the genus; however, the val ...
'' is sometimes considered a wastebasket taxon for large, long-necked
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s.
Wastebasket taxa in science
Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
groups that are poorly known due to fragmentary remains are sometimes grouped together on gross morphology or
stratigraphy
Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks.
Stratigraphy has three related subfields: lithost ...
, only later to be found to be wastebasket taxa, such as the crocodile-like
Triassic
The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
group
Rauisuchia
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians ...
.
One of the roles of taxonomists is to identify wastebasket taxa and reclassify the content into more natural units. Sometimes, during taxonomic revisions, a wastebasket taxon can be salvaged after doing thorough research on its members, and then imposing tighter restrictions on what continues to be included. Such techniques "saved" Carnosauria and ''
Megalosaurus
''Megalosaurus'' (meaning "great lizard", from Ancient Greek, Greek , ', meaning 'big', 'tall' or 'great' and , ', meaning 'lizard') is an extinct genus of large carnivorous theropod dinosaurs of the Middle Jurassic Epoch (Bathonian stage, 166 ...
''. Other times, the
taxonomic
280px, Generalized scheme of taxonomy
Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme of classes (a taxonomy) and the allocation ...
name contains too much unrelated "baggage" to be successfully salvaged. As such, it is usually dumped in favour of a new, more restrictive name (for example,
Rhynchocephalia
Rhynchocephalia (; ) is an order of lizard-like reptiles that includes only one living species, the tuatara (''Sphenodon punctatus'') of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a speciose g ...
), or abandoned altogether (for example, ''
Simia
In his ''Systema Naturae'' of 1758, Carl Linnaeus divided the Order Primates within ''Mammalia'' into four genera: ''Homo'', ''Simia'', ''Lemur'', and ''Vespertilio''. His ''Vespertilio'' included all bats, and has since been moved from Primates t ...
'').
Related concepts
A related concept is that of
form taxon
Form classification is the classification of organisms based on their morphology (biology), morphology, which does not necessarily reflect their biological relationships. Form classification, generally restricted to palaeontology, reflects uncer ...
, "wastebasket" groupings that are united by gross morphology. This is often result of a common mode of life, often one that is
generalist, leading to generally similar body shapes by
convergent evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
.
The term wastebasket taxon is sometimes employed in a derogatory fashion to refer to an
evolutionary grade
A grade is a taxon united by a level of morphological or physiological complexity. The term was coined by British biologist Julian Huxley, to contrast with clade, a strictly phylogenetic unit.
Phylogenetics
The concept of evolutionary grades ...
taxon.
See also
*
Lazarus taxon
In paleontology, a Lazarus taxon (plural ''taxa'') is a taxon that disappears for one or more periods from the fossil record, only to appear again either in later fossil records, or as actual living organisms, and often in isolated, obscure, ...
*
Elvis taxon
In paleontology, an Elvis taxon (plural ''Elvis taxa'') is a taxon that has been misidentified as having re-emerged in the fossil record after a period of presumed extinction, but is not actually a descendant of the original taxon, instead having d ...
* ''
Incertae sedis
or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...
''
*
Glossary of scientific naming
This is a list of terms and symbols used in scientific names for organisms, and in describing the names. For proper parts of the names themselves, see List of Latin and Greek words commonly used in systematic names. Many of the abbreviations are ...
*
Not Otherwise Specified
References
{{Reflist, 2
Obsolete taxa
Taxonomy (biology)
Paraphyletic groups
Polyphyletic groups