''Vorona berivotrensis'' ( ; Malagasy for "bird", ''berivotrensis'', "from Berivotra") is a

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of prehistoric

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s. It was described from

fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

found in a

Maevarano Formation The Maevarano Formation is a Late Cretaceous sedimentary formation (geology), rock formation found in the Mahajanga Province of northwestern Madagascar. It is most likely Maastrichtian in age, and records a seasonal, semiarid environment with rive ...

quarry A quarry is a type of open-pit mining, open-pit mine in which dimension stone, rock (geology), rock, construction aggregate, riprap, sand, gravel, or slate is excavated from the ground. The operation of quarries is regulated in some juri ...

near the

village A village is a human settlement or community, larger than a hamlet but smaller than a town with a population typically ranging from a few hundred to a few thousand. Although villages are often located in rural areas, the term urban v ...

Berivotra Berivotra is a rural municipality in Madagascar. It belongs to the district of Maevatanana, which is a part of Betsiboka Region. The population of the municipality was 14,675 in 2018. Infrastructure It is situated on the Route Nationale 4 fro ...

Mahajanga Province Mahajanga was a former provinces of Madagascar, province of Madagascar that had an area of 150,023 km2. It had a population of 1,896,000 (2004). Its capital was Mahajanga, the second largest city in Madagascar. Except for Fianarantsoa Provi ...

Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...

. The age of the fossilised

specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository f ...

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...

, probably

Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...

(72.1-66.0 mya). ''V. berivotrensis'' is known from scattered remains, possibly from a single individual ( UA 8651 and FMNH PA715). The phylogenic

affinity Affinity may refer to: Commerce, finance and law * Affinity (law), kinship by marriage * Affinity analysis, a market research and business management technique * Affinity Credit Union, a Saskatchewan-based credit union * Affinity Equity Pa ...

of ''Vorona'' is hard to determine due to the fragmentary nature of the remains, mainly because the fossil shows a mix of basal avian features as well as some that seem very derived. ''Vorona'' might be a primitive ornithuromorph. At least two studies recovered it as part of

Enantiornithes The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct Avialae, avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teet ...

, however. ''Vorona'' is sometimes confused with the dromaeosaur '' Rahonavis ostromi'', a fossil of which was found in the same location. This confusion has led to the common misconception that ''Vorona'' had a

deinonychosaur Deinonychosauria is a clade of paravian dinosaurs which lived from the Late Jurassic to the Late Cretaceous periods. Fossils have been found across the globe in North America, Europe, Africa, Asia, South America, and Antarctica,Case, J.A., Marti ...

-like sickle claw on each foot.

References

Further reading