The Volga pikeperch, or Volga zander (''Sander volgensis''), is a species of

fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...

in the perch family

Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains nearly 250 ...

. It is found in

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, and

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Description

The Volga pikeperch is considerably smaller than the

zander The zander (''Sander lucioperca''), sander or pikeperch, is a species of ray-finned fish from the Family (biology), family Percidae, which also includes perch, Gymnocephalus, ruffe and Darter (fish), darter. It is found in freshwater and brackis ...

or common pike-perch (''Sander lucioperca''). It grows to a maximum length of , weighing . It differs from ''Sander lucioperca'' by not having large "vampire" like canine teeth, also the colour is more silvery-grey than green, with much more distinguishable dark stripes on the side. Its second dorsal fin has nineteen to twenty-one branched soft rays and the number of scales along the

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...

is seventy to eighty-three.

Distribution and habitat

The Volga pikeperch is found in the northern

Black Sea The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...

basin from the

Danube The Danube ( ; see also #Names and etymology, other names) is the List of rivers of Europe#Longest rivers, second-longest river in Europe, after the Volga in Russia. It flows through Central and Southeastern Europe, from the Black Forest sou ...

, as far upstream as

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, to the

Kuban River The Kuban is a river in Russia that flows through the Western Caucasus and drains into the Sea of Azov. The Kuban runs mostly through Krasnodar Krai for , but also in the Karachay–Cherkess Republic, Stavropol Krai and the Republic of Adygea. ...

drainages. It is also present in the

Caspian Sea The Caspian Sea is the world's largest inland body of water, described as the List of lakes by area, world's largest lake and usually referred to as a full-fledged sea. An endorheic basin, it lies between Europe and Asia: east of the Caucasus, ...

basin in the

Volga River The Volga (, ) is the longest river in Europe and the longest endorheic basin river in the world. Situated in Russia, it flows through Central Russia to Southern Russia and into the Caspian Sea. The Volga has a length of , and a catchment ...

and

Ural River The Ural, also known as the Yaik , is a river flowing through Russia and Kazakhstan in the continental border between Europe and Asia. It originates in the southern Ural Mountains and discharges into the Caspian Sea. At , it is the third-longes ...

drainages. It inhabits large, turbid rivers and lakes, as well as brackish estuaries and may make short migrations to breed.

Biology

The Volga pikeperch is most active at dusk and dawn when it feeds on small fish and invertebrates. Males become mature by the age of three or four and females a year later, at a minimum length of . Breeding takes place during April and May in shallow water with a sandy or gravelly bottom. The male scrapes a shallow depression in the

substrate Substrate may refer to: Physical layers *Substrate (biology), the natural environment in which an organism lives, or the surface or medium on which an organism grows or is attached ** Substrate (aquatic environment), the earthy material that exi ...

or among the roots of vegetation in which the female deposits one of two or three small batches of eggs. The larvae and juvenile fish are

pelagic The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...

and feed on

zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...

Taxonomy

The Volga pikeperch was first formally described as ''Perca volgensis'' in 1789 by the

German German(s) may refer to: * Germany, the country of the Germans and German things **Germania (Roman era) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizenship in Germany, see also Ge ...

naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...

botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...

entomologist Entomology (from Ancient Greek ἔντομον (''éntomon''), meaning "insect", and -logy from λόγος (''lógos''), meaning "study") is the branch of zoology that focuses on insects. Those who study entomology are known as entomologists. In ...

herpetologist Herpetology (from Ancient Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is a branch of zoology concerned with the study of amphibians (including frogs, salamanders, and caecilians (Gymnophiona)) and reptiles (in ...

, and

malacologist Malacology, from Ancient Greek μαλακός (''malakós''), meaning "soft", and λόγος (''lógos''), meaning "study", is the branch of invertebrate zoology that deals with the study of the Mollusca (molluscs or mollusks), the second-largest ...

Johann Friedrich Gmelin (1748–1804) with the type locality given as the

and the

in Russia. The Volga pikeperch is part of the European

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

within the genus ''Sander'' which split from a common ancestor with the North American clade, which the

walleye The walleye (''Sander vitreus'', Synonym (taxonomy), synonym ''Stizostedion vitreum''), also called the walleyed pike, yellow pike, yellow pikeperch or yellow pickerel, is a freshwater perciform fish native to most of Canada and to the Northern ...

(''S. vitreus'') and the

sauger The sauger (''Sander canadensis'') is a freshwater perciform fish of the family Percidae that resembles its close relative, the walleye. The species is a member of the largest vertebrate order, the Perciformes.Jaeger, Matthew. 2004. Montana's ...

(''S. canadensis'') belong to, around 20.8 million years ago. Within the European clade the Volga pikeperch (''S. volgaensis'') is the most basal taxon and shares features with the North American clade, such as being a broadcast spawner. In contrast in the

(''S. lucioperca'') and the estuarine perch ('' S. marinus'') the males build nests and the female spawn into these nests and the males then guard the eggs and fry. The lineage leading to the zander is thought to have diverged from the common ancestor with the Volga pikeperch circa 13.8 million years ago.

Status

The

IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...

has listed the Volga pikeperch as being of "

Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...

". It has a wide range, and although the population trend is unknown, no particular threats have been identified.

References

{{Taxonbar, from=Q847041 Sander (fish) Freshwater fish of Europe Fish of the Black Sea Fish of the Caspian Sea Fish described in 1789 Taxa named by Johann Friedrich Gmelin Taxonomy articles created by Polbot