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''Vitis amurensis'', the Amur grape, is a species of
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began approximately 8,0 ...
native to the
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
n continent. Its name comes from the
Amur The Amur River () or Heilong River ( zh, s=黑龙江) is a perennial river in Northeast Asia, forming the natural border between the Russian Far East and Northeast China (historically the Outer Manchuria, Outer and Inner Manchuria). The Amur ...
Valley in
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
and
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. It is very resistant to frost, but is not tolerant to drought. Selections vary, but as a species it has strong resistance to
anthracnose A plant canker is a small area of dead tissue, which grows slowly, often over years. Some cankers are of only minor consequence, but others are ultimately lethal and therefore can have major economic implications for agriculture and horticultur ...
and ripe rot, and moderately strong resistance to downy mildew and powdery mildew.


Botanical description

A vine with stem 5–10 cm in diameter and typically spreading to 15–18 m, rarely up to 20–25 meters.
Tendril In botany, a tendril is a specialized Plant stem, stem, leaf or Petiole (botany), petiole with a thread-like shape used by climbing plants for support and attachment, as well as cellular invasion by parasitic plants such as ''Cuscuta''. There ar ...
s capable of wrapping around things will entwine the branches of neighbouring plants or anything else they can use for support. The bark is dark, scaly and with vertical stripes on old shoots. Young shoots are green, often with a reddish hue, reddish-brown in autumn. The leaves vary greatly in shape. They can be solid, three- or five-lobed, ovate or rounded, arched at the base. The size ranges from 9 to 25 cm with sharp edges and rounded-triangular serrate teeth. The surface of the leaves are
glabrous Glabrousness () is the technical term for a lack of hair, down, setae, trichomes, or other such covering. A glabrous surface may be a natural characteristic of all or part of a plant or animal, or be due to loss because of a physical condition, ...
above, densely covered with short bristles. In autumn the leaves turn bright colours - red, yellow, orange, brown. The flowers are small, and a nectar source for bees. They appear in the second week of May in its natural habitat. The Amur grape is usually
dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...
, although hermaphrodite vines do occur rarely. Fruits of the Amur grape are spherical black or purple, sometimes dark blue berries ranging from very acidic to sweet, with a diameter in the main to 12 mm, with a thick skin. The sugar content in the fruit reaches 22-23%. In their natural habitat they ripen in late September. The flesh is juicy, the berries are usually sour. The clusters can be large, comparable to the number of berries. In exceptional cases, the length of bunches up to 25 cm, and with a weight of 250 grams (usually 20-70 g).


Distribution and ecology

''Vitis amurensis'' is a relic of pre-glacial subtropical vegetation of the Far East, it reaches the continental latitude Lake Kizi ( about fifty-first parallel). Along the coast of the
Sea of Japan The Sea of Japan is the marginal sea between the Japanese archipelago, Sakhalin, the Korean Peninsula, and the mainland of the Russian Far East. The Japanese archipelago separates the sea from the Pacific Ocean. Like the Mediterranean Sea, it ...
to the north it reaches the mouth of the Muli river, and the Amur region to the west - to the river Zeya. It grows in the forests of
Manchuria Manchuria is a historical region in northeast Asia encompassing the entirety of present-day northeast China and parts of the modern-day Russian Far East south of the Uda (Khabarovsk Krai), Uda River and the Tukuringra-Dzhagdy Ranges. The exact ...
,
Amur Oblast Amur Oblast () is a federal subject of Russia (an oblast), located on the banks of the Amur and Zeya rivers in the Russian Far East. The oblast borders Heilongjiang province of the People's Republic of China (PRC) to the south. The administrati ...
,
Primorsky Krai Primorsky Krai, informally known as Primorye, is a federal subjects of Russia, federal subject (a krais of Russia, krai) of Russia, part of the Far Eastern Federal District in the Russian Far East. The types of inhabited localities in Russia, ...
, North East
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
(
Anhui Anhui is an inland Provinces of China, province located in East China. Its provincial capital and largest city is Hefei. The province is located across the basins of the Yangtze and Huai rivers, bordering Jiangsu and Zhejiang to the east, Jiang ...
,
Hebei Hebei is a Provinces of China, province in North China. It is China's List of Chinese administrative divisions by population, sixth-most populous province, with a population of over 75 million people. Shijiazhuang is the capital city. It bor ...
,
Heilongjiang Heilongjiang is a province in northeast China. It is the northernmost and easternmost province of the country and contains China's northernmost point (in Mohe City along the Amur) and easternmost point (at the confluence of the Amur and Us ...
,
Jilin ) , image_skyline = Changbaishan Tianchi from western rim.jpg , image_alt = , image_caption = View of Heaven Lake , image_map = Jilin in China (+all claims hatched).svg , mapsize = 275px , map_al ...
,
Liaoning ) , image_skyline = , image_alt = , image_caption = Clockwise: Mukden Palace in Shenyang, Xinghai Square in Dalian, Dalian coast, Yalu River at Dandong , image_map = Liaoning in China (+all claims hatched).svg , ...
,
Shandong Shandong is a coastal Provinces of China, province in East China. Shandong has played a major role in Chinese history since the beginning of Chinese civilization along the lower reaches of the Yellow River. It has served as a pivotal cultural ...
,
Shanxi Shanxi; Chinese postal romanization, formerly romanised as Shansi is a Provinces of China, province in North China. Its capital and largest city of the province is Taiyuan, while its next most populated prefecture-level cities are Changzhi a ...
,
Zhejiang ) , translit_lang1_type2 = , translit_lang1_info2 = ( Hangzhounese) ( Ningbonese) (Wenzhounese) , image_skyline = 玉甑峰全貌 - panoramio.jpg , image_caption = View of the Yandang Mountains , image_map = Zhejiang i ...
), and
Korea Korea is a peninsular region in East Asia consisting of the Korean Peninsula, Jeju Island, and smaller islands. Since the end of World War II in 1945, it has been politically Division of Korea, divided at or near the 38th parallel north, 3 ...
. Amur grapes are classified into three
varieties Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
: *''V. amurensis'' var. ''amurensis'' *''V. amurensis'' var. ''dissecta'' *''V. amurensis'' var. ''yanshanensis'' It is most common in the valleys of rivers and streams, in clearings, forest edges, forests, the lower and middle slopes of mountains, where they climb to trees and spread along the ground. It is the northernmost and most stable type of grape growing in the Far Eastern
taiga Taiga or tayga ( ; , ), also known as boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, and larches. The taiga, or boreal forest, is the world's largest land biome. In North A ...
. Most grapes species are found in much warmer climates. Only the Amur grape can tolerate winter temperatures down to −45 °C, and the root zone of the soil to −16 °C.
Anthropogenic Anthropogenic ("human" + "generating") is an adjective that may refer to: * Anthropogeny, the study of the origins of humanity Anthropogenic may also refer to things that have been generated by humans, as follows: * Human impact on the enviro ...
factors have adversely affected the number and status of the Amur grapes in their natural habitats, leading to a marked reduction in their range.


Significance and use

Amur grapes are widely used as an ornamental plant for vertical gardening. They are normally cultivated as far north as St. Petersburg in
European Russia European Russia is the western and most populated part of the Russia, Russian Federation. It is geographically situated in Europe, as opposed to the country's sparsely populated and vastly larger eastern part, Siberia, which is situated in Asia ...
where they are planted. They are also widely crossed with other grape species (usually '' V. vinifera'') to produce cold hardy, early ripening wine and dessert grapes for cold climates. They tolerate urban conditions (smoke, dust, gases). Amur grapes require about 700 mm of rain a year, so do not perform well in dry climates typical of many wine regions unless irrigated. They are well suited to wetter areas usually considered too cool and wet for grapes such as
North West Europe North West Europe may refer to: * Northwestern Europe, a loosely defined subregion of Europe, overlapping Northern and Western Europe * North-West Europe 1940, World War II campaign also known as the Battle of France * North West Europe campaign, ...
, Northern Russia and the
Pacific Northwest The Pacific Northwest (PNW; ) is a geographic region in Western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though no official boundary exists, the most common ...
. They are relatively resistant to disease, ripen early and are evolved to a short growing season and have some partial resistance to
phylloxera Grape phylloxera is an insect pest of grapevines worldwide, originally native to eastern North America. Grape phylloxera (''Daktulosphaira vitifoliae'' (Fitch 1855) belongs to the family Phylloxeridae, within the order Hemiptera, bugs); orig ...
. They can be eaten raw when ripe and are sweet (acidic and bitter when not fully ripe) but are often made into wine, juice, jellies and jams and the leaves can be used in a salad. Amur grapes have a mild resistance to a few forms of mildew. Ripening capacity and growth rate of vines is very high, the annual growth is about 2.5 meters. Amur grape seedlings bloom at about the fifth year. The plant prefers loose acidic soil and cannot tolerate excessive lime. Amur grapes respond positively to the introduction of highly acidic peat. They fruit best in full sun. Some grape varieties resulting from crossing other species with Amur grapes are: 'Arctic', 'Buytur', ' Zarya severa', 'Currants Michurin', 'Metal', 'Russian Concorde', 'North', 'Black North' and 'Kurinka Russkaja' etc. - cultivated at present in northern vineyards. Some notable hybrids with ''V. amurensis'' ancestry grown in Western Europe are " Zarya severa" and it's descendants Severny, Solaris and
Rondo The rondo or rondeau is a musical form that contains a principal theme (music), theme (sometimes called the "refrain") which alternates with one or more contrasting themes (generally called "episodes", but also referred to as "digressions" or "c ...
.


History of cultivation

Samples of ''V. amurensis'' were made in the former USSR for studies into resistance to cold. Hybrid varieties from the cross ''V. amurensis'' x ''V.vinifera'' or ''V. labrusca'' x ''V. amurensis'' were obtained. The first crop of ''V. amurensis'' was during the Japanese occupation of Manchuria (1931-1945). In
Jilin ) , image_skyline = Changbaishan Tianchi from western rim.jpg , image_alt = , image_caption = View of Heaven Lake , image_map = Jilin in China (+all claims hatched).svg , mapsize = 275px , map_al ...
in northeast China, the Japanese created the first wineries producing wine from wild grapes ''V. amurensis''. About 800 km to the south, the culture of the European vine (''V. vinifera'') is possible in the region of
Beijing Beijing, Chinese postal romanization, previously romanized as Peking, is the capital city of China. With more than 22 million residents, it is the world's List of national capitals by population, most populous national capital city as well as ...
only if the vines are buried under a thick layer of earth in winter to protect them from extreme cold. Faced with this difficulty, the Japanese tried to take advantage of high resistance to cold of the wild vine in the region to produce wine. In 1936 the Changbaishan Winery Ltd was created and Tonghua Winery Ltd 1938. in the region of
Jilin ) , image_skyline = Changbaishan Tianchi from western rim.jpg , image_alt = , image_caption = View of Heaven Lake , image_map = Jilin in China (+all claims hatched).svg , mapsize = 275px , map_al ...
, west of
Changbai Mountains The Changbai Mountains () are a major mountain range in East Asia that extends from the Northeast Chinese provinces of Heilongjiang, Jilin and Liaoning, across the China-North Korea border (41°41' to 42°51'N; 127°43' to 128°16'E), to the Nor ...
near the North Korean border. The climate of this region can be characterized by long, cold winters and short, warm summers, with average January temperatures ranging from -14C to -40C. At the time, the basement of Changbaishan produced 40 tons of wine. In 1954, the Beijing Botanical Garden tried hybridizations of ''V. amurensis'' grape with the European vine. But the decisive step was taken when, in 1965, a wild vine hermaphrodite was discovered in the mountains a hundred miles from Changbeishan winery. From there, the agricultural research institutes in the region of
Jilin ) , image_skyline = Changbaishan Tianchi from western rim.jpg , image_alt = , image_caption = View of Heaven Lake , image_map = Jilin in China (+all claims hatched).svg , mapsize = 275px , map_al ...
embarked on programs to improve the Amur grape, seeking to harness its cold resistance genes. There was an improvement of cultivars (intraspecific crosses) and hybridization with the best varieties of the European vine. The hybrids were found to be lower in sugar than the European vine and higher in acidity, but further crosses have improved on this. Many crosses were also made in the Russian research stations to obtain new varieties resistant to cold and mildew .


Chemistry

The oligostilbenes amurensin A, B, C, D, E, F,Four Novel Oligostilbenes from the Roots of Vitis amurensis. Kai-Sheng Huang, Mao Lin, Lin-Ning Yu and Man Kong, Tetrahedron, 3 March 2000, Volume 56, Issue 10, Pages 1321–1329, G, H, I, J, K, L and MAnti-inflammatory tetramers of resveratrol from the roots of Vitis amurensis and the conformations of the seven-membered ring in some oligostilbenes. Kai-Sheng Huang, Mao Lin, and Gui-Fang Cheng, Phytochemistry, September 2001, Volume 58, Issue 2, Pages 357–362, can be found in ''V. amurensis''.


Gallery

File:Амурский виноград в Хабаровске.JPG File:Синий дальневосточный виноград ф4.JPG


See also

*
Hybrid grapes Hybrid grapes are grape varieties that are the product of a Hybrid (biology), crossing of two or more ''Vitis'' species. This is in contrast to crossings between grape varieties of the same species, typically ''Vitis vinifera'', the European grape ...


References


External links


Russian distribution map
{{Taxonbar, from=Q134364 amurensis Flora of Anhui Flora of Hebei Flora of Heilongjiang Flora of Jilin Flora of Liaoning Flora of Shandong Flora of Shanxi Flora of Zhejiang Flora of Japan Flora of Korea Flora of the Russian Far East Plants described in 1857 Dioecious plants