The greater mouse-eared bat (''Myotis myotis'') is a

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

an species of

bat Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...

in the family

Vespertilionidae Vespertilionidae is a family of microbats, of the order Chiroptera, flying, insect-eating mammals variously described as the common, vesper, or simple nosed bats. The vespertilionid family is the most diverse and widely distributed of bat familie ...

Description

''Myotis myotis'' is a large bat with a long, broad muzzle and big, long ears. The body's dorsal side is brown to reddish-brown, while the ventral side is dirty white or beige. The tragus forms half of the ear, with a small black tip in most individuals. Wing membranes are brownish in colour. The Greater mouse-eared bat is relatively large for a member of the genus ''Myotis'', weighing up to and measuring 8 to 9 cm from head to tail (a little larger than a

house mouse The house mouse (''Mus musculus'') is a small mammal of the rodent family Muridae, characteristically having a pointed snout, large rounded ears, and a long and almost hairless tail. It is one of the most abundant species of the genus '' Mus''. A ...

, ''Mus musculus''), making it one of the largest European bats. It has a 40 cm wingspan, with a forearm length of 6 cm, and a 4 to 5 cm long tail. Greater mouse-eared bat are exceptionally long-lived for their size, and can have lifespans of more than 35 years.

Distribution

The greater mouse-eared bat can be found throughout Europe, with populations in most European countries except

Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...

Denmark Denmark is a Nordic countries, Nordic country in Northern Europe. It is the metropole and most populous constituent of the Kingdom of Denmark,, . also known as the Danish Realm, a constitutionally unitary state that includes the Autonomous a ...

Latvia Latvia, officially the Republic of Latvia, is a country in the Baltic region of Northern Europe. It is one of the three Baltic states, along with Estonia to the north and Lithuania to the south. It borders Russia to the east and Belarus to t ...

Estonia Estonia, officially the Republic of Estonia, is a country in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the Baltic Sea across from Sweden, to the south by Latvia, and to the east by Ru ...

Finland Finland, officially the Republic of Finland, is a Nordic country in Northern Europe. It borders Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bothnia to the west and the Gulf of Finland to the south, ...

and the

Scandinavian Peninsula The Scandinavian Peninsula is located in Northern Europe, and roughly comprises the mainlands of Sweden, Norway and the northwestern area of Finland. The name of the peninsula is derived from the term Scandinavia, the cultural region of Denm ...

. It is also found on many Mediterranean islands, such as

Sicily Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...

Malta Malta, officially the Republic of Malta, is an island country in Southern Europe located in the Mediterranean Sea, between Sicily and North Africa. It consists of an archipelago south of Italy, east of Tunisia, and north of Libya. The two ...

, and the

Gymnesian Islands The Gymnesians ( , ), or Gymnesic Islands (), is a collective name given to the two largest (and easternmost) Balearic Islands, Mallorca and Menorca, distinguishing them from the Pityuses (; ), or Pine Islands, Ibiza and Formentera. Etymology ...

. In the Middle East, ''Myotis myotis'' has been found in

Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...

Israel Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Isr ...

Syria Syria, officially the Syrian Arab Republic, is a country in West Asia located in the Eastern Mediterranean and the Levant. It borders the Mediterranean Sea to the west, Turkey to Syria–Turkey border, the north, Iraq to Iraq–Syria border, t ...

Jordan Jordan, officially the Hashemite Kingdom of Jordan, is a country in the Southern Levant region of West Asia. Jordan is bordered by Syria to the north, Iraq to the east, Saudi Arabia to the south, and Israel and the occupied Palestinian ter ...

and

Lebanon Lebanon, officially the Republic of Lebanon, is a country in the Levant region of West Asia. Situated at the crossroads of the Mediterranean Basin and the Arabian Peninsula, it is bordered by Syria to the north and east, Israel to the south ...

. It possibly lives in the

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...

. In the

Balearics The Balearic Islands are an archipelago in the western Mediterranean Sea, near the eastern coast of the Iberian Peninsula. The archipelago forms a province and autonomous community of Spain, with Palma de Mallorca being its capital and largest ...

, ''Myotis myotis'' can be found on the islands of

Mallorca Mallorca, or Majorca, is the largest of the Balearic Islands, which are part of Spain, and the List of islands in the Mediterranean#By area, seventh largest island in the Mediterranean Sea. The capital of the island, Palma, Majorca, Palma, i ...

and

Menorca Menorca or Minorca (from , later ''Minorica'') is one of the Balearic Islands located in the Mediterranean Sea belonging to Spain. Its name derives from its size, contrasting it with nearby Mallorca. Its capital is Maó, situated on the isl ...

. It is the most abundant species of bat on Mallorca, with several caves containing large colonies of 200, 400 and 500 individuals. It is also very common in the

Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...

and

France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...

. During the 20th century, this species was very rare in Great Britain, occurring only in southern

England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...

. However, the bats at the only known

hibernation Hibernation is a state of minimal activity and metabolic reduction entered by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It is mos ...

roost declined until only a few males were left, and when these disappeared the species was believed locally extinct. However, occasional individuals have been discovered in recent years, suggesting either that a colony survives or that further animals have colonized Great Britain from mainland Europe. In 2012, a LIFE-Nature project was initiated, aiming to protect the several thousands of greater mouse-eared bat in the Gola della Rossa and Frasassi Nature Park in the Marches.

Behaviour

Like its relatives, the greater mouse-eared bat is an

insectivore file:Common brown robberfly with prey.jpg, A Asilidae, robber fly eating a hoverfly An insectivore is a carnivore, carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the Entomophagy ...

, feeding on various

arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...

s; however, unlike many bats, it does not capture its prey by using echolocation in flight. Instead, it gleans it from the ground, locating prey passively by listening for the noises produced by insects such as carabid beetles, other

beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...

s and large

moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...

s, as well as

centipede Centipedes (from Neo-Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', "lip", and Neo-Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, ...

s and

spider Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...

s. As a result, it uses echolocation only for spatial orientation, even if it emits ultrasound calls when approaching prey. ''Myotis myotis'' forages in open deciduous woodlands, forest edges and pastures. It roosts underground throughout the year, although roosts in Northern Europe are also located in attics and lofts of buildings. More rarely, small tree-based colonies also occur. Colonies on Mediterranean islands such as Mallorca usually have up to 500 individuals, though colonies of over 4.500 bats have been found in continental Europe. It has been known to roost with other bats, such as ''Myotis capaccinii'' and ''Miniopterus schreibersi''. In mainland Europe, ''Myotis myotis'' performs annual dispersions of up to 200 km in spring, although most are only of 10 km or so. As with most

vespertilionid Vespertilionidae is a family of microbats, of the order Chiroptera, flying, insect-eating mammals variously described as the common, vesper, or simple nosed bats. The vespertilionid family is the most diverse and widely distributed of bat familie ...

bats, the greater mouse-eared bat exhibits slight

sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

, with females being slightly larger than males. Mating takes place in autumn, with females giving birth from May to June. Each female has 1–2 pups, which are carried around for about 45 days. During this period, females form nursery colonies from which males are excluded. The frequencies used by this bat species for echolocation lie between 22 and 86 kHz, have most energy at 37 kHz and have an average duration of 6.0 ms.

References

{{Authority control Mouse-eared bats Bats of Asia Mammals of Turkey Fauna of the Middle East Mammals of the Middle East Bats of Europe Mammals described in 1797 Habitats Directive species