Vervain Hummingbird (Mellisuga Minima)
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The vervain hummingbird (''Mellisuga minima'') is a species of
hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
in tribe
Mellisugini Mellisugini is one of the three Tribe (biology), tribes that make up the subfamily Trochilinae in the hummingbird family (biology), family Trochilidae. The other two tribes in the subfamily are Lampornithini (mountain gems) and Trochilini (emeral ...
of subfamily
Trochilinae Trochilinae is one of the six subfamily, subfamilies that make up the hummingbird family (biology), family Trochilidae. The subfamily is divided into three Tribe (biology), tribes: Lampornithini (mountain gems) containing 18 species, Mellisugini ...
, the "bee hummingbirds". It is found on
Hispaniola Hispaniola (, also ) is an island between Geography of Cuba, Cuba and Geography of Puerto Rico, Puerto Rico in the Greater Antilles of the Caribbean. Hispaniola is the most populous island in the West Indies, and the second-largest by List of C ...
(split between the
Dominican Republic The Dominican Republic is a country located on the island of Hispaniola in the Greater Antilles of the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean. It shares a Maritime boundary, maritime border with Puerto Rico to the east and ...
and
Haiti Haiti, officially the Republic of Haiti, is a country on the island of Hispaniola in the Caribbean Sea, east of Cuba and Jamaica, and south of the Bahamas. It occupies the western three-eighths of the island, which it shares with the Dominican ...
) and
Jamaica Jamaica is an island country in the Caribbean Sea and the West Indies. At , it is the third-largest island—after Cuba and Hispaniola—of the Greater Antilles and the Caribbean. Jamaica lies about south of Cuba, west of Hispaniola (the is ...
.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved 27 May 2021


Taxonomy and systematics

The vervain hummingbird was
formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...
by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1758 in the
tenth edition Tenth may refer to: Numbers * 10th, the ordinal form of the number ten * One tenth, , or 0.1, a fraction, one part of a unit divided equally into ten parts. ** the SI prefix deci- ** tithe, a one-tenth part of something * 1/10 of any unit of me ...
of his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Trochilus minimus''. Linnaeus based his description on a bird that had been described and illustrated in 1747 by the English naturalist George Edwards. The type locality is Jamaica. The vervain hummingbird is now placed together with the tiny Cuban
bee hummingbird The bee hummingbird, zunzuncito or Helena hummingbird (''Mellisuga helenae'') is a species of hummingbird, native to the island of Cuba in the Caribbean. It is the smallest known bird. The bee hummingbird feeds on nectar of flowers and bugs foun ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Mellisuga ''Mellisuga'' is a genus of hummingbirds in the family Trochilidae. They are notable for being the first and second smallest bird species in the world. The genus was introduced by the French zoologist Mathurin Jacques Brisson in 1760 with the ve ...
'' that was introduced by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...
in 1760. Two
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of vervain hummingbird are recognised, the nominate ''M. m. minima'' (Linnaeus, 1758) and ''M. m. vielloti'' ( Shaw, 1812).


Description

The vervain hummingbird is perhaps the second-smallest bird in the world after the smallest, the
bee hummingbird The bee hummingbird, zunzuncito or Helena hummingbird (''Mellisuga helenae'') is a species of hummingbird, native to the island of Cuba in the Caribbean. It is the smallest known bird. The bee hummingbird feeds on nectar of flowers and bugs foun ...
, though some other tiny ones are close to it in size. It is about long, including the bill, and weighs about . The sexes are nearly alike but for their tails and the subspecies have only subtle plumage differences. All have a short, straight, dull black bill. Males are unique among hummingbirds by not having any iridescent feathering.Clark, C. J. (2020). Vervain Hummingbird (''Mellisuga minima''), version 1.0. In Birds of the World (T. S. Schulenberg, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.verhum1.01 retrieved July 27, 2022 Males have dull metallic green uppersides that are almost black on the rump and uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
. The chin, throat, and chest are pale gray lightly spotted with darker gray; the belly and undertail coverts are dark metallic green. Its tail is entirely black and slightly forked. Females are dark metallic green to bluish green above and on the flanks. The throat is pale gray that darkens down the underparts to the vent area. The tail is rounded, not forked like the male's. The base of its central feathers are dark green and the rest of their length is black. The other tail feathers are black with varying amounts of white on their tips.


Distribution and habitat

The nominate subspecies of vervain hummingbird is found on Jamaica. ''M. m. vielloti'' is found in both the Dominican Republic and Haiti on the island of Hispaniola and also smaller nearby islands. There is one sight report from
Puerto Rico ; abbreviated PR), officially the Commonwealth of Puerto Rico, is a Government of Puerto Rico, self-governing Caribbean Geography of Puerto Rico, archipelago and island organized as an Territories of the United States, unincorporated territo ...
. The species inhabits almost every available landscape throughout its range except the interior of dense
montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...
. It occurs in
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
, dry forest, desert scrub-shrub, gardens, and even urban areas. In elevation it ranges from sea level to at least .


Behavior


Movement

The vervain hummingbird does not migrate in the conventional sense but appears to make elevational movements in response to food availability.


Feeding

The vervain hummingbird forages for nectar at a very wide variety of flowering plants though it typically feeds at small, low-yield flowers. It does not defend feeding territories. It typically cocks its tail up when feeding. In addition to nectar, it captures small insects by hawking from a perch or by hovering in a patch of flying insects.


Breeding

Male vervain hummingbirds make what appears to be courtship display flights, a flat horizontal circle by itself or with dives between two circles. The species' breeding season spans from December to August. The female makes a tiny cup nest of soft material such as hair and plant down covered with lichen or moss; it is held together and anchored to a branch with spiderweb. Nests are typically somewhat hidden and are often within of the ground. The clutch is two eggs that are among the smallest of all birds', with an average length of and weighing about . The female incubates them for 16 to 19 days and fledging occurs about 21 to 25 days after hatch.


Vocalization

Male vervain hummingbirds are highly vocal. The sing during most of the day from exposed perches in their territory and also during the courtship display and during encounters with other males. The song is complex, a mix of syllables some of which may be repeated during a song bout. The songs of the two subspecies are subtly different but the differences might be within the range of individual birds' repertoires. Both sexes make a "dry 'chittering' scolding sound" during
agonistic An agonist is a chemical that activates a receptor to produce a biological response. Receptors are cellular proteins whose activation causes the cell to modify what it is currently doing. In contrast, an antagonist blocks the action of the agoni ...
encounters.


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the vervain hummingbird as being of Least Concern. It has large range, and though its population size is not known it is believed to be stable. It is widespread and uses many habitats including human-altered areas both for food and breeding. No specific threats have been identified.


Gallery

Vervain hummingbirds (Mellisuga minima) courting.jpg, Courting, Jamaica Vervain Hummingbird.jpg, On nest Vervain hummingbird (Mellisuga minima) feeding.jpg, Feeding, Jamaica


References


Further reading

* Raffaele, Herbert; James Wiley, Orlando Garrido, Allan Keith & Janis Raffaele (2003) ''Birds of the West Indies'', Christopher Helm, London.


External links

* * {{Taxonbar, from=Q1273378 Vervain Hummingbird Endemic birds of the Caribbean Birds of the Dominican Republic Birds of Haiti Birds of Jamaica Vervain Hummingbird Vervain Hummingbird Taxonomy articles created by Polbot