Verdin
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The verdin (''Auriparus flaviceps'') is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
penduline tit The penduline tits constitute the family (biology), family Remizidae, of small perching bird, passerine birds related to the true tit (bird), tits. All but the verdin make elaborate bag nests hanging from trees (whence "penduline", hanging), usua ...
. It is the only species in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Auriparus'' and the only representative of the old world family Remizidae to be found in
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
.


Taxonomy

The verdin was
formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...
in 1850 by the Swedish zoologist
Carl Jakob Sundevall Carl Jakob Sundevall (22 October 1801 in Högestad – 2 February 1875) was a Sweden, Swedish zoologist. Sundevall studied at Lund University, where he received a Ph.D. in 1823. After traveling to East Asia, he studied medicine, graduating as a ...
under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Aegithalus flaviceps''. It is now the only species placed in the genus ''Auriparus'' that was introduced in 1864 by the American naturalist
Spencer Fullerton Baird Spencer Fullerton Baird (; February 3, 1823 – August 19, 1887) was an American naturalist, ornithologist, ichthyologist, Herpetology, herpetologist, and museum curator. Baird was the first curator to be named at the Smithsonian Institution. He ...
. The genus name combines the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
''aurum'', ''auri'' meaning "gold" with the genus name ''
Parus ''Parus'' is a genus of Old World birds in the tit family Paridae. It was formerly a large genus containing most of the 50 odd species in the family Paridae. The genus was split into several resurrected genera following the publication of a deta ...
'' that was introduced by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1758 for the tits. The specific epithet ''flaviceps'' combines the Latin ''flavis'' meaning "yellow" with ''-ceps'' meaning "-capped". Six
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised: * ''A. f. acaciarum'' Grinnell, 1931 – southwest USA to north Baja California, central
Sonora Sonora (), officially Estado Libre y Soberano de Sonora (), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is divided into Municipalities of Sonora, 72 ...
, central Chihuahua and central
Durango Durango, officially the Free and Sovereign State of Durango, is one of the 31 states which make up the Political divisions of Mexico, 32 Federal Entities of Mexico, situated in the northwest portion of the country. With a population of 1,832,650 ...
(northwest Mexico) * ''A. f. ornatus'' (
Lawrence Lawrence may refer to: Education Colleges and universities * Lawrence Technological University, a university in Southfield, Michigan, United States * Lawrence University, a liberal arts university in Appleton, Wisconsin, United States Preparator ...
, 1851) – south-central USA and northeast Mexico * ''A. f. flaviceps'' (
Sundevall Carl Jakob Sundevall (22 October 1801 in Högestad – 2 February 1875) was a Swedish zoologist. Sundevall studied at Lund University, where he received a Ph.D. in 1823. After traveling to East Asia, he studied medicine, graduating as a Doctor ...
, 1850) – central Baja California, south Sonora and north
Sinaloa Sinaloa (), officially the (), is one of the 31 states which, along with Mexico City, compose the Federal Entities of Mexico. It is divided into 18 municipalities, and its capital city is Culiacán Rosales. It is located in northwest Mexic ...
(northwest Mexico) * ''A. f. lamprocephalus'' Oberholser, 1897 – south Baja California (northwest Mexico) * ''A. f. sinaloae'' Phillips, AR, 1986 – northwest Sinaloa (northwest Mexico) * ''A. f. hidalgensis'' Phillips, AR, 1986 – north-central Mexico


Description

The verdin is a very small bird. At in length, it rivals the
American bushtit The American bushtit, or simply bushtit (''Psaltriparus minimus''), is a social songbird belonging to the genus ''Psaltriparus''. It is one of the smallest passerines in North America and it is the only species in the Family (taxonomy), family Ae ...
as one of the smallest
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
s in
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. It is gray overall, and adults have a bright yellow head and
rufous Rufous () is a color that may be described as reddish-brown or brownish- red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a d ...
shoulder patch (the lesser coverts). Unlike the tits, it has a sharply pointed
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Pl ...
. Juveniles have a light gray body without the rufous shoulder patch and yellow head. The base of the short bill is thick by the head, and draws to a sharp point at the tip.


Vocalizations

Little is known about geographic or individual variation in vocalizations. Verdins produce a variety of calls, loud for their size. The typical call note is a repetitive "tschep" sound repeated in rapid staccato. In Texas, rapid "chip-chip-chip" calls have been observed, all at the same pitch and at the same speed in each call series.” The day of hatching, nestlings can produce short peep notes, faint and high-pitched. Fledglings, during their post-juvenile molt, can make soft warbling notes, similar to more complex signatures of adult calls. The verdin gives rapid "tschep" calls frequently while foraging. Unmated males sound "tseet" calls repeatedly in early spring, which may function to attract females. "Tseet" calls are also given by either sex of a mated pair, by solitary foraging birds, and by older nestlings. This may be used as an identifier call to help keep members of the breeding pair together as they move through brush to forage.Dawson, W. L. (1923) The Birds of California: A Complete, Scientific and Popular Account of the 580 Species and Subspecies of Birds Found in the State. Volume 2. South Moulton, San Diego, CA, USA "Tweedle" warbling calls may reinforce a breeding pair bond. The verdin also gives a "gee-gee-gee-gee" alarm call in response to predators or when captured.


Distribution and habitat

Two subspecies nest in the United States. The subspecies ''A. f. acaciarum'' is resident from southern California, Nevada, and Utah southward into Mexico; it is associated mostly with the
Sonoran Desert The Sonoran Desert () is a hot desert and ecoregion in North America that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the Southwestern United States (in Arizona and California). It ...
. The subspecies ''A. f. ornatus'' is found from New Mexico and Oklahoma southward into Mexico and is associated with the Chihuahuan Desert and its scrublands. The remaining 4-5 subspecies are resident to Mexico, and aren't known to migrate into the United States. Verdins nest in shrubs and thorny thickets with few trees. Their nests are best described as spherical masses of bulky twigs and branches that are then lined with grasses, feathers, and hair. The entrance to the nest is about 1" in diameter, and is usually at the bottom of the sphere. They feed in shrub-land and scrubby outcrops where insects are hosted and berries and seeds are plentiful.


Behavior and ecology


Breeding

Verdins nest in the spring and tend to form monogamous pairs, but do not stay together year-round. Otherwise, they are usually solitary. Both males and females build nests for roosting and nesting, and both incubate the eggs and tend to young. Clutch sizes vary from 3-6 eggs, which are light green with irregular dark red-brown spots, concentrated at the bottom end of the egg. No formal courtship display is known for the species, but males sing more frequently during breeding and sometimes also build a "display nest" of sorts. This display nest may help to display partner suitability, but this correlation has not been confirmed. Once paired, the male and female claim a territory (as large as 20
acre The acre ( ) is a Unit of measurement, unit of land area used in the Imperial units, British imperial and the United States customary units#Area, United States customary systems. It is traditionally defined as the area of one Chain (unit), ch ...
s) and will defend the boundary starting early in the breeding season through
fledging Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
. After fledging, the young forage with both parents until winter, and then disperse short distances from their hatching site.


Food and feeding

Verdins are
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores we ...
, continuously foraging among the
desert A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the la ...
trees and scrubs. Verdins occasionally try to obtain sugar water from hummingbird feeders. They feed on many kinds of tiny insects, both adults and larvae, and also consume hard and soft
mast Mast, MAST or MASt may refer to: Engineering * Mast (sailing), a vertical spar on a sailing ship * Flagmast, a pole for flying a flag * Guyed mast, a structure supported by guy-wires * Mooring mast, a structure for docking an airship * Radio mas ...
of many trees and shrubs.


Conservation status

Although the verdin is listed as a species of least concern Cby the
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
, The North American
Breeding Bird Survey A breeding bird survey monitors the status and trends of bird populations. Data from the survey are an important source for the range maps found in field guides. The North American Breeding Bird Survey is a joint project of the United States Geo ...
suggests that verdin populations have declined by 1.9% per year between 1968 and 2015. This decline could mean that the species has experienced a cumulative decrease of 60% in that 47-year period. Threats to verdin populations include habitat loss as a result of
urbanization Urbanization (or urbanisation in British English) is the population shift from Rural area, rural to urban areas, the corresponding decrease in the proportion of people living in rural areas, and the ways in which societies adapt to this change. ...
, human structure collisions, and
predation Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
.


References


External links


Verdin profile
at State of Utah, Natural Resources, Division of Wildlife Resources * * * * {{Taxonbar, from=Q1300274 Auriparus Native birds of the Southwestern United States Birds of Mexico Endemic birds of Southwestern North America Birds described in 1850