Velafrons
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''Velafrons'' (meaning "sailed forehead") is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of lambeosaurine
hadrosaurid Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
. It is known from a mostly complete skull and partial skeleton of a juvenile individual, with a bony crest on the forehead. Its fossils were found in the late
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
-age Cerro del Pueblo Formation (about 72 million years oldLoewen, M.A., Sampson, S.D., Lund, E.K., Farke, A.A., Aguillón-Martínez, M.C., de Leon, C.A., Rodríguez-de la Rosa, R.A., Getty, M.A., Eberth, D.A., 2010, "Horned Dinosaurs (Ornithischia: Ceratopsidae) from the Upper Cretaceous (Campanian) Cerro del Pueblo Formation, Coahuila, Mexico", In: Michael J. Ryan, Brenda J. Chinnery-Allgeier, and David A. Eberth (eds), ''New Perspectives on Horned Dinosaurs: The Royal Tyrrell Museum Ceratopsian Symposium'', Indiana University Press, 656 pp.), near Rincon Colorado,
Coahuila Coahuila, formally Coahuila de Zaragoza, officially the Free and Sovereign State of Coahuila de Zaragoza, is one of the 31 states of Mexico. The largest city and State Capital is the city of Saltillo; the second largest is Torreón and the thi ...
, Mexico. The
type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
is CPC-59, and the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''V. coahuilensis''.


History of discovery

Largescale paleontological expeditions of the Cerro del Pueblo Formation in
Coahuila Coahuila, formally Coahuila de Zaragoza, officially the Free and Sovereign State of Coahuila de Zaragoza, is one of the 31 states of Mexico. The largest city and State Capital is the city of Saltillo; the second largest is Torreón and the thi ...
began in 1986 with an expedition by the
Royal Ontario Museum The Royal Ontario Museum (ROM) is a museum of art, world culture and natural history in Toronto, Ontario, Canada. It is one of the largest museums in North America and the largest in Canada. It attracts more than one million visitors every year ...
, which focused on a
hadrosaurid Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
bonebed near Presa de San Antonio. Subsequently, Dinamation International Society established multiple quarries around Rincon Colorado, including quarry 7A excavated from 1992 to 2001 where a mostly complete skeleton lacking a skull was found. In 2002 excavations of quarry 7A were restarted by the Utah Museum of Natural History,
Royal Tyrrell Museum of Palaeontology The Royal Tyrrell Museum of Palaeontology (RTMP; often referred to as the Royal Tyrrell Museum) is a palaeontology museum and research facility in Drumheller, Alberta, Canada. The museum was named in honour of Joseph Burr Tyrrell, and is situate ...
and Museo del Desierto, where a disarticulated skull and partial skeleton was found within a calcareous grey
mudstone Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from ''shale'' by its lack of fissility.Blatt, H., and R.J. Tracy, 1996, ''Petrology.'' New York, New York, ...
by Hernández-Rivera. From
magnetostratigraphy Magnetostratigraphy is a geophysical correlation technique used to date sedimentary and volcanic sequences. The method works by collecting oriented samples at measured intervals throughout the section. The samples are analyzed to determine their ' ...
, the Cerro de Pueblo Formation is around 73.5
million years Million years ago, abbreviated as Mya, Myr (megayear) or Ma (megaannum), is a unit of time equal to (i.e. years), or approximately 31.6 teraseconds. Usage Myr is in common use in fields such as Earth science and cosmology. Myr is also used ...
old, with the skeleton from around above the base of the formation. The skeleton and skull, CPC-59, was described in
2007 2007 was designated as the International Heliophysical Year and the International Polar Year. Events January * January 1 **Bulgaria and Romania 2007 enlargement of the European Union, join the European Union, while Slovenia joins the Eur ...
by American paleontologist Terry Gates and colleagues as the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
of the new taxon ''Velafrons coahuilensis'', the first new genus of lambeosaurine from North America in over 70 years. The
genus name Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial spec ...
combines the
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many countries in the Americas **Spanish cuisine **Spanish history **Spanish culture ...
word ''vela'' and
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
word ''frons'' as "sailed forehead" in reference to the cranial crest, while the
species name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
means "from Coahuila". Despite being one of the most complete hadrosaurs from Mexico, ''Velafrons'' remains only partially described, with its postcranial skeleton being undescribed. Gates and colleagues interpreted ''Velafrons'' to be a subadult individual based on absolute size and the crest anatomy, comparing it most favourably with the ''
Corythosaurus ''Corythosaurus'' (; ) is a genus of hadrosaurid "duck-billed" dinosaur from the Late Cretaceous Period (geology), period, about 77–75.7 million years ago, in what is now Laramidia, western North America. Its name is derived from the Anci ...
'' specimen CMN 34825. As that specimen is only about 60% of the full adult size of ''Corythosaurus'', that could suggest that ''Velafrons'' would be able to attain much larger sizes than its only known skeleton indicates.


Description

''Velafrons'' was most similar to young specimens of ''
Corythosaurus ''Corythosaurus'' (; ) is a genus of hadrosaurid "duck-billed" dinosaur from the Late Cretaceous Period (geology), period, about 77–75.7 million years ago, in what is now Laramidia, western North America. Its name is derived from the Anci ...
'' and ''
Hypacrosaurus ''Hypacrosaurus'' (meaning "near the highest lizard"
reek υπο-, ''hypo-'' = less + ακρος, ''akros'', high Reek may refer to: Places * Reek, Netherlands, a village in the Dutch province of North Brabant * Croagh Patrick, a mountain in the west of Ireland nicknamed "The Reek" People * Nikolai Reek (1890–1942), Estonian military commander * Salme Ree ...
because it was almost but not quite as large as ''Tyrannosaurus'') is an extinct genus of hadrosaurid, duckbill dinosaur simila ...
'', and was found to be a corythosaurin in the
phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...
performed by Gates and colleagues in their description of the genus. The skull was large in comparison to skulls from other genera at a similar growth stage, so the crest may have been small in adults or followed a different growth pattern, or it may be that adult ''Velafrons'' were also larger than adults of other lambeosaurine genera. Unusually large size is also seen in the Mexican hadrosaurids ''
Kritosaurus ''Kritosaurus'' is an incompletely known genus of hadrosaurid (duck-billed) dinosaur. It lived about 74.5-66 million years ago, in the Late Cretaceous of North America. The name means "separated lizard" (referring to the arrangement of the cheek ...
'' sp. and '' Magnapaulia laticaudus''. As a hadrosaurid, ''Velafrons'' would have been a
herbivore A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
. The animal is known from several disarticulated remains, those being certain cervical vertebrae and a disarticulated skull.


References

{{Taxonbar, from=Q134733 Lambeosaurinae Dinosaur genera Campanian dinosaurs Taxa named by Scott D. Sampson Taxa named by Lindsay Zanno Taxa named by James I. Kirkland Fossil taxa described in 2007 Dinosaurs of Mexico