Variable Squirrel
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Finlayson's squirrel or the variable squirrel (''Callosciurus finlaysonii'', sometimes misspelled ''C. finlaysoni'') is a species of rodent in the family Squirrel, Sciuridae. It is found in Cambodia, Laos, Myanmar, Thailand, and Vietnam. The species occurs in a wide range of wooded habitats, including gardens and parks in cities like Bangkok.Francis, C.M. (2008). A Guide to the Mammals of Southeast Asia. Pp. 144–145, 333–334. Princeton University Press. Ecology Asia
Variable or Finlayson's Squirrel
Retrieved 11 July 2018.
It was named in honour of the Scottish naturalist and traveller George Finlayson. It has numerous subspecies that vary greatly in appearance. One of these, ''C. f. bocourti'' (syn. ''C. f. floweri''), has been Introduced species, introduced to Singapore and two regions in Italy, probably a result of the species' popularity in the pet trade. It is possible that some of the ''Callosciurus'' squirrels introduced into Japan are also Finlayson's squirrels. Finlayson's squirrel has also been introduced in the Philippines, particularly in the Greater Manila area where it is considered as an Invasive species in the Philippines, invasive species.


Taxonomy and appearance

The Finlayson's squirrel has a head-and-body length of about and its tail is about long. There are currently 16 recognised subspecies. Additional subspecies are sometimes recognised. For example, some authorities recognise ''C. f. floweri'', while others consider it a Synonym (taxonomy), synonym of ''C. f. bocourti'' (as done in the following colour description). The subspecific name of ''C. f. boonsongi'' commemorates Thai zoologist and conservationist Boonsong Lekagul. The pelage colour in this species is extremely variable and the subspecies are often defined by this feature. For example, ''C. f. finlaysonii'' (Nominate subspecies, nominate) is overall whitish, ''C. f. albivexilli'', ''C. f. boonsongi'', ''C. f. germaini'' and ''C. f. nox'' are overall blackish (first with white tail-tip, second occasionally with white underside, face and feet), ''C. f. annellatus'' is overall rufous with a light band at the base of the tail, ''C. f. bocourti'' is whitish below with highly variable colour of the upperparts (whitish, grey, blackish, Olive (color), olive-brownish or reddish), ''C. f. cinnamomeus'' is overall reddish with a dark mid-back, ''C. f. ferrugineus'' is reddish-brown, ''C. f. harmandi'' has brownish upperparts, orange-red underparts and light grey tail, ''C. f. menamicus'' is reddish or orangish, often has greyish legs and flanks, and sometimes a white belly, and ''C. f. sinistralis'' has grizzled upperparts, reddish underparts and reddish tail with a pale band at the base.Boonkhaw, Phadet, Prayoon, Umphornpimon, Kanchanasaka, Budsabong, Hayashi, Fumio, Tamura, Noriko (2017). Colour polymorphism and genetic relationships among twelve subspecies of Callosciurus finlaysonii in Thailand. ''Mammalian Biology – Zeitschrift für Säugetierkunde 85'', 6–13. Undescribed taxon, Unnamed populations also remain (for example, a population in central Laos is shiny black with red tail and shoulder/chest region) and even within described subspecies there are often some individual variations. As currently defined, Finlayson's squirrel may comprise more than one species. ''C. f. ferrugineus'' has been treated as a separate species. A Genetics, genetic study of the 12 subspecies in Thailand, including 7 from the mainland and 5 from islands, found that they belonged in six clades, which often were separated by water (large rivers in the mainland and the sea for the islands). The study also revealed that the mitochondrial DNA of this species compared to the closely related Pallas's squirrel is not monophyletic. A subsequent study recommended that, to maintain monophyletic species, ''flavimanus'' and ''griseimanus'', which traditionally are recognized as being subspecies of Pallas's squirrel in southeastern Indochina (central and southern Vietnam, adjacent parts of Cambodia and southern Laos), should be considered part of Finlayson's squirrel. The study also found that ''flavimanus'', as traditionally defined, actually consists of at least two clearly separate Lineage (evolution), lineages, both part of Finlayson's squirrel. File:Callosciurus finlaysonii 85959681.jpg, ''C. f. bocourti'' mating pair in Nonthaburi province, Nonthaburi, Thailand (colour difference is individual, not related to male/female) File:Callosciurus finlaysonii 177035226.jpg, ''C. f. bocourti'' in Wang Nam Khiao district, Wang Nam Khiao, Thailand File:Callosciurus finlaysonii cinnamomeus 86117364.jpg, ''C. f. cinnamomeus'' in Khao Soi Dao Wildlife Sanctuary, Khao Soi Dao, Thailand File:Callosciurus finlaysonii 99592168.jpg, ''C. f. germaini'' on Côn Đảo, Vietnam


Behavior

Like other squirrels of its genus (the "beautiful squirrels", ''Callosciurus''), Finlayson's squirrel is normally a canopy-dweller, feeding mainly on fruit. Field evidence suggests that it has the usual form of dichromacy, dichromatic mammalian color vision, which may enable it to discriminate ripe from unripe fruits.Tamura, N., Fujii, Y., Boonkeow, P., & Kanchanasaka, B. (2015). Colour vision and food selection of ''Callosciurus finlaysonii'' (Sciuridae) in tropical seasonal forests. ''Journal of Tropical Ecology, 31'', 449–457.


References


External links


Wildlife Singapore Photos and description

Photo gallery showing some of the variations of Finlayson's Squirrel.
{{Taxonbar, from=Q2086835 Callosciurus Mammals of Southeast Asia Rodents of Myanmar Rodents of Thailand Rodents of Laos Fauna of Italy Mammals described in 1823 Taxa named by Thomas Horsfield Taxonomy articles created by Polbot