''Uteodon'' (meaning "
Ute
Ute or UTE may refer to:
* Ute (band), an Australian jazz group
* Ute (given name)
* ''Ute'' (sponge), a sponge genus
* Ute (vehicle), an Australian and New Zealand term for certain utility vehicles
* Ute, Iowa, a city in Monona County along the ...
tooth") is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
herbivorous
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
iguanodontian
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
. It is a
basal
Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''.
Science
* Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure
* Basal (medicine), a minimal level that is nec ...
iguanodontian which lived during the late
Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
period (
Tithonian
In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by t ...
age) in what is now
Uintah County,
Utah
Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its ...
. It is known from the middle of the
Brushy Basin Member,
Morrison Formation
The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone, ...
. The genus was named by Andrew T. McDonald in
2011 and the
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
is ''U. aphanoecetes''.
it has also been reported from portugal.
History
The holotype specimen, CM 11337 (a virtually complete skeleton minus the skull and tail), was assigned to ''
Camptosaurus medius'' (
Marsh
A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found a ...
, 1894) by
Charles W. Gilmore in 1925.
When ''C. medius'' was synonymised with ''Camptosaurus dispar'' in 1980,
[ G.S. Paul (2010) "'' The Princeton Field Guide to Dinosaurs''". (Page 284).] the holotype was seen to probably represent a new, then unnamed, species of ''Camptosaurus''. The species ''
Camptosaurus
''Camptosaurus'' ( ) is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America and possibly also Europe. The name means 'flexible lizard' (Greek (') meaning 'bent' and (') meaning 'lizar ...
aphanoecetes'' was first described in 2008 by Carpenter and Wilson. In 2011, it was assigned to the new genus ''Uteodon''.
In 2015, the ''Uteodon'' braincase was referred to ''
Dryosaurus
''Dryosaurus'' ( , meaning 'tree lizard', Greek ' () meaning 'tree, oak' and () meaning 'lizard'; the name reflects the forested habitat, not a vague oak-leaf shape of its cheek teeth as is sometimes assumed) is a genus of an ornithopod dinosaur ...
,'' and ''Uteodon'' and ''
Cumnoria'' were synonymized with ''Camptosaurus'', as ''C. aphanoectes'' and ''C. prestwichii,'' respectively.
Description

Based on the holotype and the related genus ''
Camptosaurus
''Camptosaurus'' ( ) is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America and possibly also Europe. The name means 'flexible lizard' (Greek (') meaning 'bent' and (') meaning 'lizar ...
'', when fully grown, ''Uteodon'' would have grown up to around long and would probably have weighed no more than around ,
although according to Hartman (2013), ''Uteodon'' could have been as small as around long.
Paleoecology
Provenance and occurrence
The single known specimen of ''Uteodon'', CM 11337, was found in the
Dry Mesa Quarry/
Douglass Quarry of the
Brushy Basin Member of the
Morrison Formation
The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone, ...
,
Utah
Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its ...
.
The rocks it was found in were medium-grained, coarse sandstone that was deposited during the
Tithonian
In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by t ...
and
Kimmeridgian
In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxford ...
stages of the
Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987.
In European lithostratigraphy, the ...
period, approximately 153 to 148 million years ago.
Fauna and habitat
Studies suggest that the
paleoenvironment of this section of the Morrison Formation included rivers that flowed from the west into a basin that contained a giant, saline alkaline lake and there were extensive wetlands in the vicinity. The Dry Mesa Dinosaur Quarry of western Colorado yields one of the most diverse Upper Jurassic vertebrate assemblages in the world.
[Richmond, D.R. and Morris, T.H., (1999), Stratigraphy and cataclysmic deposition of the Dry Mesa Dinosaur Quarry, Mesa County, Colorado, in Carpenter, K., Kirkland, J., and Chure, D., eds., The Upper Jurassic Morrison Formation: An Interdisciplinary Study, ''Modern Geology v. 22'', no. 1-4, pp. 121–143.] The Dry Mesa Quarry has produced the remains of the
sauropods ''
Apatosaurus
''Apatosaurus'' (; meaning "deceptive lizard") is a genus of herbivorous sauropod dinosaur that lived in North America during the Late Jurassic period. Othniel Charles Marsh described and named the first-known species, ''A. ajax'', in 187 ...
'', ''
Diplodocus
''Diplodocus'' (, , or ) was a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a neo-Latin term derived from Greek διπ� ...
'', ''
Barosaurus
''Barosaurus'' ( ) was a giant, long-tailed, long-necked, plant-eating sauropod dinosaur closely related to the more familiar '' Diplodocus''. Remains have been found in the Morrison Formation from the Upper Jurassic Period of Utah and Sou ...
'', ''
Supersaurus
''Supersaurus'' (meaning "super lizard") is a genus of diplodocid sauropod dinosaur that lived in North America during the Late Jurassic period. The type species, ''S. vivianae'', was first discovered by Vivian Jones of Delta, Colorado, in the ...
'', and ''
Camarasaurus
''Camarasaurus'' ( ) was a genus of quadrupedal, herbivorous dinosaurs and is the most common North American sauropod fossil. Its fossil remains have been found in the Morrison Formation, dating to the Late Jurassic epoch (Kimmeridgian to Titho ...
'', the
iguanodonts ''
Camptosaurus
''Camptosaurus'' ( ) is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America and possibly also Europe. The name means 'flexible lizard' (Greek (') meaning 'bent' and (') meaning 'lizar ...
'' and ''
Dryosaurus
''Dryosaurus'' ( , meaning 'tree lizard', Greek ' () meaning 'tree, oak' and () meaning 'lizard'; the name reflects the forested habitat, not a vague oak-leaf shape of its cheek teeth as is sometimes assumed) is a genus of an ornithopod dinosaur ...
'', and the
theropods ''
Allosaurus
''Allosaurus'' () is a genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic epoch ( Kimmeridgian to late Tithonian). The name "''Allosaurus''" means "different lizard" alludi ...
'', ''
Torvosaurus
''Torvosaurus'' () is a genus of carnivorous megalosaurid theropod dinosaur that lived approximately 165 to 148 million years ago during the late Middle and Late Jurassic period ( Callovian to Tithonian stages) in what is now Colorado, Portugal ...
''. ''
Tanycolagreus
''Tanycolagreus'' is a genus of coelurosaurian theropod from the Late Jurassic of North America.
Discovery and naming
In 1995 Western Paleontological Laboratories, Inc. uncovered the partial skeleton of a small theropod at the Bone Cabin Quarr ...
'', ''
Koparion'', ''
Stokesosaurus'', ''
Ceratosaurus
''Ceratosaurus'' (from Greek κέρας/κέρατος, ' meaning "horn" and σαῦρος ' meaning "lizard") was a carnivorous theropod dinosaur in the Late Jurassic period (Kimmeridgian to Tithonian). The genus was first described in 1884 ...
'', and ''
Ornitholestes
''Ornitholestes'' (meaning "bird robber") is a small theropod dinosaur of the late Jurassic ( Brushy Basin Member of the Morrison Formation, middle Kimmeridgian age, about 154 million years agoTurner, C.E. and Peterson, F., (1999). "Biostratigrap ...
'', as well as ''
Nanosaurus
''Nanosaurus'' ("small or dwarf lizard") is the name given to a genus of neornithischian dinosaur that lived about 155 to 148 million years ago, during the Late Jurassic-age. Its fossils are known from the Morrison Formation of the south-wester ...
'', ''
Gargoyleosaurus'', and ''
Stegosaurus
''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been fou ...
''.
The flora of the period has been revealed by fossils of
green algae
The green algae (singular: green alga) are a group consisting of the Prasinodermophyta and its unnamed sister which contains the Chlorophyta and Charophyta/ Streptophyta. The land plants ( Embryophytes) have emerged deep in the Charophyte alg ...
,
fungi
A fungus (plural, : fungi or funguses) is any member of the group of Eukaryote, eukaryotic organisms that includes microorganisms such as yeasts and Mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified ...
,
moss
Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta ('' sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and ...
es,
horsetails
''Equisetum'' (; horsetail, snake grass, puzzlegrass) is the only living genus in Equisetaceae, a family of ferns, which reproduce by spores rather than seeds.
''Equisetum'' is a "living fossil", the only living genus of the entire subclass ...
, ferns,
cycad
Cycads are seed plants that typically have a stout and woody (ligneous) trunk with a crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants of a species are either male or f ...
s,
ginkgo
''Ginkgo'' is a genus of non-flowering seed plants. The scientific name is also used as the English name. The order to which it belongs, Ginkgoales, first appeared in the Permian, 270 million years ago, and is now the only living genus withi ...
es, and several families of
conifer
Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ex ...
s. Animal fossils discovered include
bivalve
Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, biv ...
s,
snail
A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class G ...
s,
ray-finned fishes,
frog
A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" '' Triadobatrachus'' is ...
s,
salamander
Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten ...
s,
amphibians,
turtle
Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked ...
s,
sphenodonts,
lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia al ...
s, terrestrial (like ''
Hoplosuchus'') and aquatic
crocodylomorpha
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction.
During Mesozoic and early Cenozoic times, cr ...
ns, cotylosaurs, several species of
pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the Order (biology), order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cre ...
s like ''
Harpactognathus'', and early
mammals,
multituberculates
Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, ...
,
symmetrodonts, and
triconodonts
Eutriconodonta is an order of early mammals. Eutriconodonts existed in Asia, Africa, Europe, North and South America during the Jurassic and the Cretaceous periods. The order was named by Kermack ''et al.'' in 1973 as a replacement name for the ...
.
[
]
References
{{Taxonbar, from=Q420956
Iguanodonts
Late Jurassic dinosaurs of North America
Fossil taxa described in 2011
Dinosaurs of the Morrison Formation
Paleontology in Utah
Ornithischian genera