Urocoptidae
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Urocoptidae is a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of air-breathing
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial molluscs, terrestrial gastropod mollusks that have gastropod shell, shel ...
s,
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
pulmonate Pulmonata or pulmonates is an informal group (previously an order, and before that, a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group inclu ...
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
s in the superfamily
Urocoptoidea Urocoptoidea is a superfamily of land snails, gastropods in the suborder Helicina.MolluscaBase eds. (2020). MolluscaBase. Urocoptoidea Pilsbry, 1898 (1868). Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphi ...
.


Taxonomy


2005 taxonomy

The family Urocoptidae was classified in the superfamily
Orthalicoidea Orthalicoidea is a superfamily of air-breathing land snails, terrestrial gastropod mollusks in the infraorder Orthalicoidei of the suborder Helicina.MolluscaBase eds. (2021). MolluscaBase. Simpulopsidae Schileyko, 1999. Accessed through: Wor ...
(according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005): * Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868 * Apomatinae Paul, 1982 * Brachypodellinae H. B. Baker, 1956 * Eucalodiinae P. Fischer & Crosse, 1873 * Holospirinae Pilsbry, 1946 * Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972 * Tetrentodontinae Bartsch, 1943


2008 taxonomy

Uit de Weerd (2008) moved the Urocoptidae to the newly established superfamily
Urocoptoidea Urocoptoidea is a superfamily of land snails, gastropods in the suborder Helicina.MolluscaBase eds. (2020). MolluscaBase. Urocoptoidea Pilsbry, 1898 (1868). Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphi ...
based on
molecular phylogeny Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
research. The following
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
based on
Bayesian analysis Thomas Bayes ( ; c. 1701 – 1761) was an English statistician, philosopher, and Presbyterian Presbyterianism is a historically Reformed Protestant tradition named for its form of church government by representative assemblies of elde ...
of sequences of
28S ribosomal RNA 28S ribosomal RNA is the structural ribosomal RNA (rRNA) for the large subunit (LSU) of eukaryotic cytoplasmic ribosomes, and thus one of the basic components of all eukaryotic cells. It has a size of 25S in plants and 28S in mammals, hence th ...
shows the phylogenic relations within the Urocoptoidea. There are up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color in the cladogram.


2012 taxonomy

Fred G. Thompson established the family Epirobiidae in 2012 for five genera of snails ('' Epirobia'', '' Propilsbrya'', '' Pectinistemma'', '' Gyrocion'', '' Prionoloplax''), that were previously placed within Urocoptidae.Thompson F. G. (2012). "The land snail genus ''Epirobia'' and allied genera in México and Central America, with the description of a new family, the Epirobiidae (Gastropoda, Pulmonata, Urocoptoidea)". ''
Bulletin of the Florida Museum of Natural History The Florida Museum of Natural History (FLMNH) is Florida's official state-sponsored and chartered natural history museum. Its main facilities are located at 3215 Hull Road on the campus of the University of Florida in Gainesville. The main pub ...
'' 51: 167-215
PDF
Data from
molecular phylogeny Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
research as well as morphology supports monophyly of Holospiridae Pilsbry, 1946 and the Eucalodiidae Crosse & Fischer, 1868.


Genera

Genera in the family Urocoptidae include: Urocoptinae * '' Autocoptis'' Pilsbry, 1902 * '' Brachypodella'' L. Pfeiffer, 1840 (as ''Cylindrella'') * '' Urocoptis'' Beck, 1837 - type genus of the family Urocoptidae Apominae * '' Apoma'' Beck, 1837 - type genus of the subfamily Apomatinae * '' Simplicervix'' Pilsbry, 1903 Brachypodellinae * '' Brachypodella'' Beck, 1837 - type genus of the subfamily Brachypodellinae Eucalodiinae * '' Anisospira'' Strebel, 1880 * '' Archegocoptis'' Pilsbry, 1903 *†''
Condonella ''Condonella'' is an extinct genus of land snail in the family Urocoptidae known from the fossil species ''Condonella suciensis'' of Western North America. History and classification ''C. suciensis'' is known from a single internal mold fossil ...
'' McLellan, 1927 * '' Dissotropis'' Bartsch, 1906 * '' Eucalodium'' Crosse & P. Fischer, 1868 - type genus of the subfamily Eucalodiinae Holospirinae * '' Coelostemma'' Dall, 1895 * '' Hendersoniella'' Dall, 1905 * '' Holospira'' Martens, 1860 - type genus of the subfamily Holospirinae Microceraminae * '' Microceramus'' Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae * '' Macroceramus'' Guilding, 1828 Tetrentodontinae * '' Tetrentodon'' Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae * '' Torrecoptis'' Bartsch, 1943 Other genera include: * '' Allocoptis'' Thompson & Franz, 1976 * '' Amphicosmia'' Pilsbry & Vanatta, 1898 * '' Anoma'' * '' Arangia'' * '' Bactrocoptis'' * '' Badiofax'' * '' Bostrichocentrum'' * '' Bostrychocentrum'' * '' Callonia'' * '' Centralia'' * '' Cochlodinella'' * '' Coelocentrum'' * '' Geoscala'' * '' Gongylostoma'' * '' Gyraxis'' * '' Haplocion'' * '' Heterocoptis'' * '' Idiostemma'' * '' Liocallonia'' * ''
Metastoma The metastoma is a ventral single plate located in the opisthosoma of non-arachnid dekatriatan chelicerates such as eurypterids, chasmataspidids and the genus '' Houia''. The metastoma located between the base of 6th prosomal appendage pair and ...
'' * '' Mychostoma'' * '' Nesocoptis'' * '' Nodulia'' * '' Oligostylus'' * '' Organocoptis'' * '' Pfeiffericoptis'' * '' Pineria'' * '' Pleurostemma'' * '' Poeycoptis'' * '' Pseudopineria'' * '' Pycnoptychia'' Pilsbry & Vanatta, 1898 * '' Sagracoptis'' * '' Spiroceramus'' * '' Spirostemma'' * '' Strophina'' * '' Tomelasmus'' * '' Trilamellaxis'' * '' Uncinicoptis''


References


Further reading

* ; 7 figs. * Carlos de la Torre. 1912. ''New Cuban Urocoptis of the U. cinerea group''. The Nautilus
volume 26
number 4, page
55
58
plate 6
* Kabat A. R., Herschler R. & González G. A. (2012). "Resolution of taxonomic problems associated with the complex publication history of the seminal Torre and Bartsch monograph on Cuban Urocoptidae (Gastropoda, Pulmonata)". ''
Zootaxa ''Zootaxa'' is a peer-reviewed scientific mega journal for animal taxonomists. It is published by Magnolia Press (Auckland, New Zealand). The journal was established by Zhi-Qiang Zhang in 2001 and new issues are published multiple times a week ...
'' 3362: 43–53
PDF
* Pilsbry H. A. & Henderson J. B. jr. (February 1913). ''Two new Cuban Urocoptidae''
The Nautilus, volume 26
number 10, page
109
110. * Miguel L. Jaume & Alfredo de la Torre. 1976. ''Los Urocoptidae de Cuba (Mollusca-Pulmonata)''. Série 4: Ciencias biologicas; no. 53. La Habana, Cuba: Centro de Información Cientifica y Tecnica, 122 pp. * * Richardson C. L. 1991. ''Urocoptidae: catalogue of species''. Tryonia 22: 1–245. * Tryon G. W. (1903) '' Manual of Conchology. Second series: Pulmonata
Volume 15
Urocoptidae''. Continued by H. A. Pilsbry.


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