''Urera'' is a genus of

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s in the nettle family, Urticaceae. It has a

pantropical A pantropical ("all tropics") distribution is one which covers Tropics, tropical regions of both hemispheres. Examples of species include caecilians, modern sirenians and the plant genera ''Acacia'' and ''Bacopa''. ''Neotropical'' is a zoogeogra ...

distribution.Steinmann, V. W. (2005)
Four new neotropical species and a new combination of ''Urera'' (Urticaceae).
''Acta Botánica Mexicana'' (71), 19-43.

Description

''Urera'' are

liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ta ...

s,''Urera''.
Flora Zambesiaca.

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

s, and small

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s. Climbing species root along the stems and can reach the crowns of the trees they use for support. Some are spiny. They almost always have urticating hairs, often on bumps in the epidermis. These are stiff, stinging

trichome Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a pla ...

s with swollen bases. The leaves are variable in shape and venation, and help in the identification of species. Also useful is the varying appearance and arrangement of

cystolith Cystolith (Gr. "cavity" and "stone") is a botanical term for outgrowths of the epidermal cell wall, usually of calcium carbonate, formed in a cellulose matrix in special cells called lithocysts, generally in the leaf of plants. Cystoliths are pr ...

s on the leaf surface. The

inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...

is a divided, branching

panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...

of flowers. Most species are

dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...

, but a few are

monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...

. Most species have male flowers with either four or five

stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s and

tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...

s. Female flowers generally have four uneven, free to mostly fused tepals. The fruit is an

achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and indehiscent (they do not ope ...

covered in orange or red flesh.

Systematics and taxonomy

About 113 species epithets have been published for the genus. Many species have been transferred to other genera, including '' Boehmeria'', '' Dendrocnide'', ''

Girardinia ''Girardinia'' is a genus of flowering plants belonging to the family Urticaceae. Its native range is the tropical and subtropical Old World to Russian Far East. It is found in the countries of Angola, Assam, Bangladesh, Burundi, Cameroon, China, ...

'', ''

Gyrotaenia ''Gyrotaenia'' is a genus of plants in the family Urticaceae The Urticaceae are a family, the nettle family, of flowering plants. The family name comes from the genus ''Urtica''. The Urticaceae include a number of well-known and useful plant ...

'', '' Laportea'', and ''

Obetia ''Obetia'' is a genus of dioecious plants in the family Urticaceae The Urticaceae are a family, the nettle family, of flowering plants. The family name comes from the genus ''Urtica''. The Urticaceae include a number of well-known and usefu ...

''. It is not clear how many valid species currently belong in ''Urera'', with estimates ranging from 17''Urera''.
The Plant List. to 75. The genus is not well known, with current knowledge dependent mainly on outdated studies, and there is little agreement regarding its

taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...

. The genus ''Urera'' was first described by Charles Gaudichaud-Beaupré in 1826 in his review of the studies done on

Louis de Freycinet Louis Claude de Saulces de Freycinet (7 August 1779 – 18 August 1841) was a French Navy officer. He circumnavigated the earth, and in 1811 published the first map to show a full outline of the coastline of Australia. Biography He was born at ...

's voyage of the '' Uranie''. He noted its distinctness, proposing the

subtribe Subtribe is a taxonomic category ranking which is below the rank of tribe and above genus. The standard suffix for a subtribe is -ina (in animals) or -inae (in plants Plants are predominantly photosynthetic eukaryotes of the kingdom Plant ...

Urereae. This was later raised to the rank of

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...

, and subsequently merged into Urticeae.Friis, I. (1989): The Urticaceae: a systematic review. ''In'': Crane, P. E. & Blackmore, S. (eds.): ''Evolution, systematics, and fossil history of the Hamamelidae'' (Vol. 2): 285–308. ''Systematics Association Special Volume'' 40B. Oxford Science Publications, Oxford, UK. ''Urera'' was expanded to include the

monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

''Scepocarpus''. Analysis of '' trnL-F''

chloroplast A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, and stores it in ...

DNA

sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is calle ...

data suggests that ''Urera'' could be sister to ''

Poikilospermum ''Poikilospermum'' is a genus of shrubs or tall woody climbers, consisting of at least 33 accepted species. The plants are found from India and China (Yunnan), the Sino-Himalayan region to Sulawesi and Jawa in Indonesia. Some of the plants are u ...

'' within the Urticaceae.Monro, A. K. (2006)
The revision of species-rich genera: a phylogenetic framework for the strategic revision of ''Pilea'' (Urticaceae) based on CPDNA, NRDNA and morphology.
'' American Journal of Botany'' 93(3): 426-41. No botanical

monograph A monograph is a specialist work of writing (in contrast to reference works) or exhibition on a single subject or an aspect of a subject, often by a single author or artist, and usually on a scholarly subject. In library cataloging, ''monograph ...

s have been devoted to ''Urera'' since 1869. Knowledge of the systematics of the genus has accumulated mainly from new descriptions of species and floristic accounts. urera-caracasana-berries

Selected species

Species include: *''

Urera acuminata ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *'' Urera aurantiaca'' *'' Urera baccifera'' ( L.) Gaudich. ex Wedd., 1852 – guaritoto, scratchbush *''

Urera cameroonensis ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *'' Urera capitata'' *'' Urera caracasana'' (

Jacq. Nikolaus Joseph Freiherr von Jacquin (16 February 172726 October 1817) was a scientist who studied medicine, chemistry and botany. Biography Born in Leiden in the Netherlands, he studied medicine at Leiden University, then moved first to Par ...

) Gaudich. ex

Griseb. August Heinrich Rudolf Grisebach () was a German botany, botanist and phytogeography, phytogeographer. He was born in Hannover on 17 April 1814 and died in Göttingen on 9 May 1879. Biography Grisebach studied at the Lyceum in Hanover, the clo ...

, 1859 – flameberry *''

Urera elata ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *'' Urera expansa'' (

Sw. Olof Peter Swartz (21 September 1760 – 19 September 1818) was a Swedish botanist and taxonomist. He is best known for his taxonomic work and studies into pteridophytes. Biography Olof Swartz attended the University of Uppsala where he st ...

) Griseb. *'' Urera glabra'' (

Hook. Sir William Jackson Hooker (6 July 178512 August 1865) was an English botanist and botanical illustrator, who became the first director of Kew when in 1841 it was recommended to be placed under state ownership as a botanic garden. At Kew h ...

Arn. George Arnott Walker Arnott of Arlary (6 February 1799 – 17 April 1868) was a Scottish botanist. Early life George Arnott Walker Arnott was born in Edinburgh in 1799, the son of David Walker Arnott of Arlary. He attended Milnathort Parish ...

) Wedd. – hopue, ''Ōpuhe'' (

Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...

) *''

Urera henriquesii ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *''

Urera kaalae ''Urera kaalae'', opuhe, is a species of flowering plant in the nettle family, Urticaceae, that is endemic to the island of Oahu in Hawaii. It inhabits slopes and gulches in mesic forests at elevations of . Currently it is restricted to the sout ...

'' Wawra – ''Ōpuhe'' (

Oahu Oahu () (Hawaiian language, Hawaiian: ''Oʻahu'' ()), also known as "The Gathering place#Island of Oʻahu as The Gathering Place, Gathering Place", is the third-largest of the Hawaiian Islands. It is home to roughly one million people—over t ...

in Hawaii) *''

Urera laciniata ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *''

Urera mannii ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *''

Urera nitida ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *''

Urera repens ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *''

Urera simplex ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *''

Urera sinuata ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *''

Urera thonneri ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' *''

Urera trinervis ''Urera trinervis'' (Hochst.) Friis & Immelman is a softly woody dioecious liane, sometimes epiphytic, climbing to 20 m, often to the canopy and hanging in festoons. It is one of some 44 species of ''Urera'' belonging to the nettle family Urtic ...

'' (

Hochst. Christian Ferdinand Friedrich Hochstetter (16 February 1787 – 20 February 1860) was a German botanist and Protestant minister. Biography Hochstetter was born in Stuttgart in Baden-Württemberg. He was the father of geologist Ferdinand Hoc ...

) Friis & Immelman, 1987 *''

Urera verrucosa ''Urera'' is a genus of flowering plants in the nettle family, Urticaceae. It has a pantropical distribution.Steinmann, V. W. (2005)Four new neotropical species and a new combination of ''Urera'' (Urticaceae).''Acta Botánica Mexicana'' (71), 19- ...

'' – ''chichicastle'', ''mala mujer''

Description

Systematics and taxonomy

Selected species

References

Further reading