''Eusideroxylon'' is a genus of
evergreen
In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s of the family
Lauraceae
Lauraceae, or the laurels, is a plant Family (biology), family that includes the bay laurel, true laurel and its closest relatives. This family comprises about 2850 known species in about 45 genus (biology), genera worldwide. They are dicotyled ...
. The genus is
monotypic
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
, and includes one accepted species, ''Eusideroxylon zwageri''. It is known colloquially in English as Bornean ironwood,
billian, or ulin.
[
It is native to ]Borneo
Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...
and Sumatra
Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
,[ where it grows in lowland rain forests.][
''Eusideroxylon'' are hardwood trees reaching up to 50 metres in height with trunks over in diameter, producing commercially valuable timber. The wood of ''E. zwageri'' is impervious to termites, and can last up to 100 years after being cut. Due to extensive logging, it is listed as vulnerable in the ]IUCN Red List
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
.[
]
Description
''Eusideroxylon'' are canopy tree species with erect or spreading branches and extremely durable and decay-resistant wood.
''Eusideroxylon zwageri'' is a slow growing (0.5 metres per year) tall evergreen tree with a straight bole (usually host to '' Cassytha'', a parasitic vine with leaves reduced to scales, up to half of the tree's height). It is slightly fluted at the base, up to 150–220 cm in diameter. The trunk has many small, rounded buttress
A buttress is an architectural structure built against or projecting from a wall which serves to support or reinforce the wall. Buttresses are fairly common on more ancient (typically Gothic) buildings, as a means of providing support to act ...
es that give the base an ''elephant-foot'' like appearance. Individual trees may reach an age of 1,000 years or more. Common commercially exploitable trees attain a height of 30 or more metres (100 ft) with trunk diameters of exploitable trees up to 92 cm (36 inches). Protected trees are towering giants of the forest attaining a height of up to 50 metres and a diameter of 220 cm – though height is routinely reduced by lightning strikes. An Ulin tree discovered in 1993 in Kutai National Park, is one of the largest plants in Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
. It is an estimated 1,000 years old, and has increased its diameter from 2.41 to 2.47 metres in the 20 years since its discovery. Its height was however reduced from some 30 metres to only 20 after a lightning strike.[ Another at Sangkimah in the west of the park has a diameter of 2.25 metres and a height of some 45 metres.]
The color of the flowers is pale yellow to yellow. The flower is hermaphrodite, actinomorphic, with 6 tepal
A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s, distributed in two whorl
A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs).
In nature
File:Photograph and axial plane floral diagra ...
s that overlap. There are six staminode
In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co ...
s, three stamen
The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10
Morphology and terminology
A stamen typically consists of a stalk called the filament ...
s, and a simple pistil
Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
that consists of one carpel. Pollination
Pollination is the transfer of pollen from an anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds. Pollinating agents can be animals such as insects, for example bees, beetles or bu ...
is done by bees and other insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s. The fruits are drupe
In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
s, varying in size and shape from oblong to ovate or sub-cylindrical to asymmetric elongated or rounded. They are 8–13 cm long, 4–5 cm in diameter, and weigh 90g-170g. Seed dispersal
In spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant.
Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...
is by vertebrate animals such as birds, monkeys, bats and rodents, for which the fruits are an important food source.
The trees' leaves are dark green, simple, leathery, elliptical to ovate, 14–18 cm long (5.5–7.5 inches) and 5–11 cm wide (2–4 inches), and are alternate, rarely whorled or opposite, without stipules
In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). They are primarily found among dicots and rare among monocots. Stipules are considered part of the anatomy ...
and petiolate. The leaf blade is entire
Entire may refer to:
* Entire function, a function that is holomorphic on the whole complex plane
* Entire (animal), an indication that an animal is not neutered
* Entire (botany)
This glossary of botanical terms is a list of definitions o ...
(unlobed or lobed in '' Sassafras'') and occasionally with domatia
A domatium (plural: domatia, from the Latin "domus", meaning home) is a tiny chamber that houses arthropods, produced by a plant.
Ideally domatia differ from galls in that they are produced by the plant rather than being induced by their inhabi ...
(crevices or hollows serving as lodging for mite
Mites are small arachnids (eight-legged arthropods) of two large orders, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari. However, most recent genetic analyses do not recover the two as eac ...
s) in axils of main lateral veins (present in ''Cinnamomum
''Cinnamomum'' is a genus of evergreen aromatic trees and shrubs belonging to the laurel family, Lauraceae. The species of ''Cinnamomum'' have aromatic oils in their leaves and bark. The genus contains approximately 234 species, distributed in ...
''). Young leaves are reddish brown to yellowish red. They have a generous layer of wax, making them glossy in appearance, and are narrow, pointed oval in shape with an ''apical mucro'', or 'drip tip', which enables the leaves to shed excess water in a humid environment.
Timber segments collected from 15 logged stumps in Kubah National Park, Sarawak, Malaysia, found that ''E. zwageri'' can live more than 1000 years and that the growth rate of this species is very slow, with a mean radial growth rate of 0·058 cm y-1. The cutting of old trees of ''E. zwageri'' results in the species being replaced by faster growing species.[ The long life span and reduced growth rate of this species may be a result of its dense and durable wood containing abundant defensive compounds.][
]
Etymology
The name ''Eusideroxylon'' is Latinized Greek, derived from Greek ''sideros'' meaning iron, ''xylon'' meaning wood,[ with the prefix ''eu-'' meaning good, true, original.][ The name of the genus thus means "true iron wood".
]
Habitat
''Eusideroxylon zwageri'' seedlings require some shade, while older trees need plenty of light. It can be found in valleys and on hillsides and even on low ridges when soil moisture is sufficient at elevations between sea level and 625 m. The standing timber volume of trees with a diameter of over 50 cm may be as much as 90–112 m3.
Silviculture
''Eusideroxylon zwageri'' has a very slow growth rate of mean
A mean is a quantity representing the "center" of a collection of numbers and is intermediate to the extreme values of the set of numbers. There are several kinds of means (or "measures of central tendency") in mathematics, especially in statist ...
radial
Radial is a geometric term of location which may refer to:
Mathematics and Direction
* Vector (geometric), a line
* Radius, adjective form of
* Radial distance (geometry), a directional coordinate in a polar coordinate system
* Radial set
* A ...
growth of 0.058 cm per year. It is a canopy species in primary forests. The species is considered unsuitable for large-scale plantations due to slow growth and inadequate seed and seedling supply. Manual selection of trees in natural forests is common.
Properties
The heartwood
Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
when cut is coloured light brown to almost bright yellow. During the aging process the heartwood
Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
darkens to deep reddish brown, very dark brown or almost black. The sapwood is bright yellow when cut, and darkens slightly. The wood texture is fine and even, with a straight grain
A grain is a small, hard, dry fruit (caryopsis) – with or without an attached husk, hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and ...
or only slightly interlocked. The timber retains a pleasant lemon odour. This odour, along with the woods' natural high lustre, make it prized by cabinet-makers and fine furniture craftsmen.
The wood is dense, and texture is moderately fine to fine and even. Also attractive to users is the resistance to insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s, bacteria
Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr ...
, fungi
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
and marine borers. The wood has anti-bacterial properties (for local medicinal use)[Soerianegara, I. & Kartawinata, K. (1983) "Silvicultural management of the logged natural dipterocarp forest in South-east Asia", in J. Davidson, Tho Yow Pong & M. Bijleveld, eds. ''Future of tropical rainforests in South-east Asia'' Commission of Ecology Papers, 10. Gland, Switzerland, IUCN]
Vessels are diffuse-porous, medium-sized and generally evenly distributed, arranged in short radial rows (2–3 vessels). Moderate abundancy of aliform paratracheal parenchyma. Growth rings boundaries are indistinct or absent. Tyloses are often present.
The wood has a radial shrinkage rate of 2–4.5% and a tangential shrinkage rate of tangential 4.5–7.5%. The timber dries slowly, and care is needed to avoid checks and splits. The wood is famed for its easy working characteristics, despite high density. The wood planes, bores and turns cleanly, producing smooth and often lustrous surfaces. Nailing requires pre-bores prior to nailing. Saw
A saw is a tool consisting of a tough blade, Wire saw, wire, or Chainsaw, chain with a hard toothed edge used to cut through material. Various terms are used to describe toothed and abrasive saws.
Saws began as serrated materials, and when man ...
blades and other cutting instruments are moderately blunted during working the timber. The wood is apparently difficult to glue
Adhesive, also known as glue, cement, mucilage, or paste, is any non-metallic substance applied to one or both surfaces of two separate items that binds them together and resists their separation.
The use of adhesives offers certain advantage ...
with synthetic resin
Synthetic resins are industrially produced resins, typically viscous substances that convert into rigid polymers by the process of curing. In order to undergo curing, resins typically contain reactive end groups, such as acrylates or epoxides. ...
s.
Heartwood is rated as very durable – immune to termite
Termites are a group of detritivore, detritophagous Eusociality, eusocial cockroaches which consume a variety of Detritus, decaying plant material, generally in the form of wood, Plant litter, leaf litter, and Humus, soil humus. They are dist ...
attack; service life
A product's service life is its period of use in service. Several related terms describe more precisely a product's life, from the point of manufacture, storage, and distribution, and eventual use.
Service life has been defined as "a product' ...
of up to 100 years in direct soil contact and more than 20 years for marine work in tropical waters has been reported.
Usage
Due to the excellent resistance to bacterial
Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among the ...
, fungal
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as one of the tradit ...
, insect and marine borer attack the wood is highly prized for many outdoor uses, especially as decking. Additionally, the wood's high density and easy workability lend it to particularly desirability in maritime structures, dock
The word dock () in American English refers to one or a group of human-made structures that are involved in the handling of boats or ships (usually on or near a shore). In British English, the term is not used the same way as in American Engl ...
construction and ship building
Shipbuilding is the construction of ships and other floating vessels. In modern times, it normally takes place in a specialized facility known as a shipyard. Shipbuilders, also called shipwrights, follow a specialized occupation that traces i ...
, especially Indonesia's pinisi
Literally, the word pinisi refers to a type of rigging (the configuration of masts, sails and ropes ('lines')) of Indonesian Sailing ship, sailing vessels. A pinisi carries seven to eight sails on two masts, arranged like a gaff-ketch with what ...
sail-boat.
Common local uses include: House construction, door construction, water butts and troughs, boat building (Pinisi
Literally, the word pinisi refers to a type of rigging (the configuration of masts, sails and ropes ('lines')) of Indonesian Sailing ship, sailing vessels. A pinisi carries seven to eight sails on two masts, arranged like a gaff-ketch with what ...
), tools
A tool is an object that can extend an individual's ability to modify features of the surrounding environment or help them accomplish a particular task. Although many animals use simple tools, only human beings, whose use of stone tools dates ...
, tool handles, talisman
A talisman is any object ascribed with religious or magical powers intended to protect, heal, or harm individuals for whom they are made. Talismans are often portable objects carried on someone in a variety of ways, but can also be installed perm ...
, jewellery
Jewellery (or jewelry in American English) consists of decorative items worn for personal adornment such as brooches, ring (jewellery), rings, necklaces, earrings, pendants, bracelets, and cufflinks. Jewellery may be attached to the body or the ...
, medicinal slivers (for wounds, cuts, abrasions, bites and tooth-ache/infection), bridges, blowpipes and spear shafts.
Internationally, it is renowned for heavy construction such as a buffer between transportation trailers and heavy steel fabrications (such as boilers, pressure vessels
A pressure vessel is a container designed to hold gases or liquids at a pressure substantially different from the ambient pressure.
Construction methods and materials may be chosen to suit the pressure application, and will depend on the size o ...
, reactors and many others). It is also frequently found in dry docks
Dry or dryness most often refers to:
* Rain#Deserts, Lack of rainfall, which may refer to
**Arid regions
**Drought
* Dry or dry area, relating to legal prohibition of selling, serving, or imbibing alcoholic beverages
* Dry humor, deadpan
* Drynes ...
as a timber to separate the hull of ships
A ship is a large vessel that travels the world's oceans and other navigable waterways, carrying cargo or passengers, or in support of specialized missions, such as defense, research and fishing. Ships are generally distinguished from boats, ...
from the steel supporting stands. Other uses include use in boats and ships, industrial flooring, roofing (as shingles
Shingles, also known as herpes zoster or zona, is a viral disease characterized by a painful skin rash with blisters in a localized area. Typically the rash occurs in a single, wide mark either on the left or right side of the body or face. T ...
), fine indoor and outdoor furniture, coffin
A coffin or casket is a funerary box used for viewing or keeping a corpse, for burial, entombment or cremation. Coffins are sometimes referred to as caskets, particularly in American English.
A distinction is commonly drawn between "coffins" a ...
wood (esteemed by Chinese due to ability to withstand rot and insect attack) and tool handles (especially those exposed to continual high impact (the wood does not splinter and thus injure hands, eyes or endanger the operator on catastrophic failure) such as shovel
A shovel is a tool used for digging, lifting, and moving bulk materials, such as soil, coal, gravel, snow, sand, or ore. Most shovels are hand tools consisting of a broad blade fixed to a medium-length handle. Shovel blades are usually made ...
s, axe
An axe (; sometimes spelled ax in American English; American and British English spelling differences#Miscellaneous spelling differences, see spelling differences) is an implement that has been used for thousands of years to shape, split, a ...
s, block splitters, sledge hammer
A sledgehammer is a tool with a large, flat, massive, often metal head, attached to a long wooden or solid handle. The long handle is combined with a heavy head which allows the sledgehammer to pick up momentum during a swing and applying a large ...
s, heavy mallet
A mallet is a tool used for imparting force on another object, often made of rubber or sometimes wood, that is smaller than a maul or beetle, and usually has a relatively large head.
General overview
The term is descriptive of the ...
s, demolition hammer
Demolition Hammer is an American thrash metal band from the Bronx, New York (state), New York, active from 1986 to 1995 and again since 2016. To date, the band has released three full-length studio albums: ''Tortured Existence'' (1990), ''Epid ...
s, mattocks, picks, hoes and hammers
A hammer is a tool, most often a hand tool, consisting of a weighted "head" fixed to a long handle that is swung to deliver an impact to a small area of an object. This can be, for example, to drive nail (fastener), nails into wood, to sh ...
). Some expert cabinet-maker
A cabinet is a case or cupboard with shelves or drawers for storing or displaying items. Some cabinets are stand alone while others are built in to a wall or are attached to it like a Bathroom cabinet, medicine cabinet. Cabinets are typically mad ...
s treasure an ''ulin''-headed carpenter's mallet as an excellent intermediate density hammer face between the usual wood and a metal one and is able to quite easily tap or "whack" stubborn highly polished metal fixtures without damage to the face or the fixture.
Other sources indicate that ''ulin'' wood is often used for marine constructions such as piling
A pile or piling is a vertical structural element of a deep foundation, driven or drilled deep into the ground at the building site. A deep foundation is a type of foundation that transfers building loads to the earth farther down from th ...
s, wharf
A wharf ( or wharfs), quay ( , also ), staith, or staithe is a structure on the shore of a harbour or on the bank of a river or canal where ships may dock to load and unload cargo or passengers. Such a structure includes one or more Berth (mo ...
s, dock
The word dock () in American English refers to one or a group of human-made structures that are involved in the handling of boats or ships (usually on or near a shore). In British English, the term is not used the same way as in American Engl ...
s, sluices, dam
A dam is a barrier that stops or restricts the flow of surface water or underground streams. Reservoirs created by dams not only suppress floods but also provide water for activities such as irrigation, human consumption, industrial use, aqua ...
s, ships, bridges, but also used for power line poles, masts, roof shingles and house posts and to a minor extent as frame, board, heavy duty flooring, railway sleepers, fencing material, furniture etc.
Conservation
The government of Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
and the state government of Sarawak
Sarawak ( , ) is a States and federal territories of Malaysia, state of Malaysia. It is the largest among the 13 states, with an area almost equal to that of Peninsular Malaysia. Sarawak is located in East Malaysia in northwest Borneo, and is ...
have formally banned the export of this species. Illegal smuggling
Smuggling is the illegal transportation of objects, substances, information or people, such as out of a house or buildings, into a prison, or across an international border, in violation of applicable laws or other regulations. More broadly, soc ...
continues to be a major problem.
Conservation efforts are underway, with several countries banning imports.
Species
* ''Eusideroxylon zwageri''
Formerly placed here:
* '' Potoxylon melagangai'', as ''Eusideroxylon melagangai''
References
External links
Asian Forest Network
CITES website
* Indonesian Department of Forestry:
Department Perhutanan
': (English available onsite)
* at the IUCN Red List
{{Authority control
Lauraceae genera
Monotypic Laurales genera
Lauraceae
Lauraceae of Borneo
Trees of Sumatra
Flora of the Borneo lowland rain forests
Plants described in 1863
Taxonomy articles created by Polbot
Taxa named by Johannes Elias Teijsmann
Taxa named by Simon Binnendijk