Tympanic Bone
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The tympanic part of the
temporal bone The temporal bone is a paired bone situated at the sides and base of the skull, lateral to the temporal lobe of the cerebral cortex. The temporal bones are overlaid by the sides of the head known as the temples where four of the cranial bone ...
is a curved plate of bone lying below the squamous part of the temporal bone, in front of the mastoid process, and surrounding the external part of the
ear canal The ear canal (external acoustic meatus, external auditory meatus, EAM) is a pathway running from the outer ear to the middle ear. The adult human ear canal extends from the auricle to the eardrum and is about in length and in diameter. S ...
. It originates as a separate bone (tympanic bone), which in some
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s stays separate through life. Evolutionarily, a portion of it is derived from the angular bone of the reptilian
lower jaw In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
.


Surfaces

Its postero-superior surface is concave, and forms the anterior wall, the floor, and part of the posterior wall of the bony
ear canal The ear canal (external acoustic meatus, external auditory meatus, EAM) is a pathway running from the outer ear to the middle ear. The adult human ear canal extends from the auricle to the eardrum and is about in length and in diameter. S ...
. Medially, it presents a narrow furrow, the ''tympanic sulcus'', for the attachment of the
tympanic membrane In the anatomy of humans and various other tetrapods, the eardrum, also called the tympanic membrane or myringa, is a thin, cone-shaped membrane that separates the external ear from the middle ear. Its function is to transmit changes in pressur ...
. Its antero-inferior surface is quadrilateral and slightly concave; it constitutes the posterior boundary of the mandibular fossa, and is in contact with the retromandibular part of the
parotid gland The parotid gland is a major salivary gland in many animals. In humans, the two parotid glands are present on either side of the mouth and in front of both ears. They are the largest of the salivary glands. Each parotid is wrapped around the m ...
.


Borders

Its lateral border is free and rough, and gives attachment to the cartilaginous part of the ear canal. Internally, the tympanic part is fused with the petrous portion, and appears in the retreating angle between it and the squama, where it lies below and lateral to the orifice of the
auditory tube The Eustachian tube (), also called the auditory tube or pharyngotympanic tube, is a tube that links the nasopharynx to the middle ear, of which it is also a part. In adult humans, the Eustachian tube is approximately long and in diameter. It ...
. Posteriorly, it blends with the squama and mastoid part, and forms the anterior boundary of the tympanomastoid fissure. Its upper border fuses laterally with the back of the postglenoid process, while medially it bounds the
petrotympanic fissure The petrotympanic fissure (also known as the squamotympanic fissure or the glaserian fissure) is a fissure in the temporal bone that runs from the temporomandibular joint to the tympanic cavity. The mandibular fossa is bounded, in front, by the ...
. The medial part of the lower border is thin and sharp; its lateral part splits to enclose the root of the styloid process, and is therefore named the ''vaginal process''.


Ear canal

The central portion of the tympanic part is thin, as it gives rise to the bony inner two-thirds of the
ear canal The ear canal (external acoustic meatus, external auditory meatus, EAM) is a pathway running from the outer ear to the middle ear. The adult human ear canal extends from the auricle to the eardrum and is about in length and in diameter. S ...
, and in 5 - 20% of skulls the lower surface is perforated by a hole, the ''foramen of Huschke'' that opens onto the
temporomandibular joint In anatomy, the temporomandibular joints (TMJ) are the two joints connecting the jawbone to the skull. It is a bilateral Synovial joint, synovial articulation between the temporal bone of the skull above and the condylar process of mandible be ...
due to incomplete fusion of the anterior and posterior prominences during development. The bony portion of the ear canal is nearly 2 cm long and is directed inward and slightly forward: at the same time it forms a slight curve, so that the floor of the canal is convex upward. In sagittal section it presents an oval or elliptical shape with the long axis directed downward and slightly backward. Its anterior wall and floor and the lower part of its posterior wall are formed by the tympanic part; the roof and upper part of the posterior wall by the squama. Its inner end is closed, by the tympanic membrane which originates from the tympanic sulcus; the upper limit of its outer orifice is formed by the posterior root of the zygomatic process, immediately below which there is sometimes seen a small spine, the ''suprameatal spine'', situated at the upper and posterior part of the orifice. The auditory bulla (pl. bullae) is a hollow bony structure on the ventral, posterior portion of the skull that encloses parts of the middle and
inner ear The inner ear (internal ear, auris interna) is the innermost part of the vertebrate ear. In vertebrates, the inner ear is mainly responsible for sound detection and balance. In mammals, it consists of the bony labyrinth, a hollow cavity in the ...
. In most species, it is formed by the tympanic part of the temporal bone.


Other animals

In all extant and extinct
primate Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...
s, including humans, the auditory bulla is formed by the petrosal bone (the petrous part of the temporal bone). This is a diagnostic trait that can be used to distinguish primates, including
anthropoids The simians, anthropoids, or higher primates are an infraorder (Simiiformes ) of primates containing all animals traditionally called monkeys and apes. More precisely, they consist of the parvorders Platyrrhini (New World monkeys) and Catar ...
,
tarsier Tarsiers ( ) are haplorhine primates of the family Tarsiidae, which is the lone extant family within the infraorder Tarsiiformes. Although the group was prehistorically more globally widespread, all of the existing species are restricted to M ...
s,
lemur Lemurs ( ; from Latin ) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea ( ), divided into 8 Family (biology), families and consisting of 15 genera and around 100 existing species. They are Endemism, ...
s, and
loris Loris is the common name for the strepsirrhine mammals of the subfamily Lorinae (sometimes spelled Lorisinae) in the family Lorisidae. ''Loris'' is one genus in this subfamily and includes the slender lorises, ''Nycticebus'' is the genus cont ...
es, from all other mammals.


Additional images

File:Gray189.png, Left infratemporal fossa. File:Gray907.png, External and
middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles (malleus, incus, and stapes), which transfer the vibrations ...
, opened from the front. Right side. File:Bulles auditives sur crâne de renard roux.JPG, Auditory bullae on a
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe and Asia, plus ...
skull (').


References


External links

* - "Osteology of the Skull: Lateral Surface of Skull" {{Authority control Bones of the head and neck