The turtle-jawed moa-nalo (''Chelychelynechen quassus''), also formerly referred to as the large Kauai goose, is a species of

moa-nalo The moa-nalo are a group of extinct aberrant, goose-like ducks that lived on the larger Hawaiian Islands, except Hawaii itself, in the Pacific. They were the major herbivores on most of these islands until they became extinct after human settl ...

, one of a group of extinct,

flightless Flightless birds are birds that through evolution lost the ability to fly. There are over 60 extant species, including the well known ratites ( ostriches, emu, cassowaries, rheas, and kiwi) and penguins. The smallest flightless bird is ...

, large goose-like

duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a fo ...

s, which

evolved Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation te ...

in the

Hawaiian Islands The Hawaiian Islands ( haw, Nā Mokupuni o Hawai‘i) are an archipelago of eight major islands, several atolls, and numerous smaller islets in the North Pacific Ocean, extending some from the island of Hawaii in the south to northernmost ...

of the

North Pacific Ocean North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''no ...

. It was described in 1991 from

subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

material collected in 1976 by

Storrs Olson Storrs Lovejoy Olson (April 3, 1944 – January 20, 2021) was an American biologist and ornithologist who spent his career at the Smithsonian Institution, retiring in 2008. One of the world's foremost avian paleontologists, he was best known ...

from the

calcareous Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an a ...

Makawehi

dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, fl ...

s on the south-eastern coast of the island of Kauai. Its remains have also been found at the nearby Makauwahi Cave site.

Taxonomy

The generic name ''Chelychelynechen'' comes from the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

''chelys'' (“turtle”), ''chelyne'' (“lip” or “jaw”), with ''chen'' (“goose”), referring to the distinctly

turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked ...

-like appearance and structure of the

beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for fo ...

. The specific

epithet An epithet (, ), also byname, is a descriptive term (word or phrase) known for accompanying or occurring in place of a name and having entered common usage. It has various shades of meaning when applied to seemingly real or fictitious people, di ...

is from the

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...

''quassus'' (“broken” or “shattered”), alluding to the fragmentary condition of the type material.

Description

Apart from the head, the bird was similar to other moa-nalo species, with robust legs and reduced, non-functional wings. The main distinguishing feature was the remarkable beak which had a rostrum almost as high as it was long, with a broad

palatal The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly sepa ...

surface, and with

nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called t ...

s oriented almost vertically rather than horizontally. The structure of this feeding apparatus appears convergent on that of

tortoise Tortoises () are reptiles of the family Testudinidae of the order Testudines (Latin: ''tortoise''). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like ot ...

s and indicates adaptation to a diet of plants.

References

Anatidae Late Quaternary prehistoric birds Holocene extinctions Extinct birds of Hawaii Biota of Kauai Extinct flightless birds turtle-jawed moa-nalo turtle-jawed moa-nalo turtle-jawed moa-nalo {{Anseriformes-stub