Truncatella (gastropod)
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''Truncatella'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of very small
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial molluscs, terrestrial gastropod mollusks that have gastropod shell, shel ...
s with an operculum,
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
s in the family
Truncatellidae Truncatellidae, common name the "looping snails", is a Family (biology), family of small amphibious snails, with gills and an operculum (gastropod), operculum, semi-marine gastropod mollusks or micromollusks. Shell description This family of s ...
. These minute
snail A snail is a shelled gastropod. The name is most often applied to land snails, terrestrial molluscs, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gas ...
s live in land habitats close to
seawater Seawater, or sea water, is water from a sea or ocean. On average, seawater in the world's oceans has a salinity of about 3.5% (35 g/L, 35 ppt, 600 mM). This means that every kilogram (roughly one liter by volume) of seawater has approximat ...
. They walk with a looping action, and the adults have truncated shells. Many of the species are small enough to be considered
micromollusk A micromollusc is a shelled mollusc which is extremely small, even at full adult size. The word is usually, but not exclusively, applied to marine molluscs, although in addition, numerous species of land snails and freshwater molluscs also ...
s. ''Truncatella'' is the
type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...
of the family Truncatellidae.


Description

As implied by the name, these snails are notable for the way they modify their elongated shells as they mature, breaking off several of the apical
whorls A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). In nature File:Photograph and axial plane floral diagram ...
, and forming a relatively smooth seal for the remainder of the shell.


Locomotion

The method of locomotion used by ''Truncatella'' is unusual. Instead of gliding over a slime trail using minute waves in its foot, ''Truncatella'' uses a very large and muscular
proboscis A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular arthropod mouthparts, mouthparts used for feeding and sucking. In vertebrates, a pr ...
to reach ahead and grasp a surface, at which point the small foot releases its hold and the proboscis contracts to pull the animal forward, after which the entire procedure is usually repeated.


Distribution and habitat

This genus occurs in tropical and subtropical areas. The various
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of ''Truncatella'' are
pantropical A pantropical ("all tropics") distribution is one which covers tropical regions of both the Eastern and Western hemispheres. Examples of species include caecilians, modern sirenians and the plant genera ''Acacia'' and ''Bacopa''. ''Neotropical' ...
in distribution, with a few exceptions such as '' Truncatella truncatula'' and '' Truncatella subcylindrica''. Most of the species in this genus live in a
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
that is neither fully
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
nor fully marine: they live under plant debris near high tide level, where they are occasionally wetted with seawater by waves. A small minority of the species are fully terrestrial. These small snails are typically found associated with drifts of plant material, where their eggs are deposited. The adaptations of these land snails to the terrestrial environment are not so perfect as they are in the more usual
pulmonate Pulmonata or pulmonates is an informal group (previously an order, and before that, a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group inclu ...
land snails, and their terrestrial adaptations may in fact be comparatively recent.


Species

The genus ''Truncatella'' was erected by
Antoine Risso Giuseppe Antonio Risso (8 April 1777 – 25 August 1845), called Antoine Risso, was a naturalist from Nice. Risso was born in the city of Nice in the Duchy of Savoy, and studied under Giovanni Battista Balbis. He published ' (1810), ' (1826) an ...
(Risso, 1826) for ''T. costulata'' (now ''T. subcylindrica''), which is the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
for this genus. There are several genera within the Truncatellidae, but the eponymous genus ''Truncatella'' is the largest (in terms of number of species); other genera in the family are '' Geomelania'', ''
Taheitia ''Taheitia'' is a genus of very small land snails that have an Operculum (gastropod), operculum and live near saltwater, maritime terrestrial animal, terrestrial gastropod mollusks in the family Truncatellidae.
'', and '' Blandiella.'' Ross, Landon T. (1970). The anatomy of ''Truncatella'' Risso 1826 (Mollusca: Prosobranchia), and a revision of the genus in the Gulf of Mexico and the Caribbean Sea. unpublished Florida State Univ. dissertation. Retrieved on September 200

/ref> There are approximately 30 species in the genus ''Truncatella'', including: * † '' Truncatella arcicostata'' H.-J. Wang, 1982 * '' Truncatella avenacea'' Garrett, 1887 * '' Truncatella bahamensis'' Clench and Turner, 1948 * '' Truncatella bairdiana'' C. B. Adams, 1852 * '' Truncatella barbadensis'' L. Pfeiffer, 1857 * '' Truncatella brazieri'' Cox, 1868 * '' Truncatella californica'' Pfeiffer, 1857 - California truncatella * '' Truncatella caribaeensis'' Reeve, 1842 - Caribbean truncatella * '' Truncatella clathrus'' Lowe, 1832 * † '' Truncatella crassolabia'' H.-Z. Pan, 1982 * '' Truncatella diaphana'' Gassies, 1869 * † '' Truncatella directa'' Y.-T. Li, 1988 * '' Truncatella granum'' Garrett, 1872 * '' Truncatella guerinii'' Villa & Villa, 1841 * † '' Truncatella hermitei'' Bardin, 1879 * † '' Truncatella hubeiensis'' Y.-T. Li, 1987 * † '' Truncatella jiaozhouensis'' T. Yu, Salvador, H. Wang, Y. Fang, Neubauer, S. Li, H. Zang & X. Wan, 2021 * '' Truncatella marginata'' Küster, 1855 * '' Truncatella oagariensis'' (Kuroda, 1960) * † '' Truncatella obliqua'' H.-Z. Pan, 1982 * '' Truncatella obscura'' Morelet, 1882 * † '' Truncatella parva'' Y.-T. Li, 1988 * '' Truncatella pfeifferi'' Martens, 1860Truncatella
OBIS, accessed 13 June 2011.
* '' Truncatella quadrasi'' Möllendorff, 1893 * † '' Truncatella rabora'' H.-Z. Pan, 1982 * '' Truncatella reclusa'' (Guppy, 1871) * '' Truncatella rostrata'' Gould, 1848 * '' Truncatella rustica'' Mousson, 1865 * † '' Truncatella sanshuiensis'' W. Yü & X.-Q. Zhang, 1982 * '' Truncatella scalarina'' Cox, 1867 * '' Truncatella scalaris'' (Michaud, 1830) - ladder truncatella * '' Truncatella semperi'' Kobelt, 1884 * † '' Truncatella sinensis'' Y.-T. Li, 1988 * '' Truncatella stimpsonii'' Stearns, 1872: synonym of '' Truncatella californica'' L. Pfeiffer, 1857 * '' Truncatella subcylindrica'' (Linnaeus, 1767) * '' Truncatella teres'' (Pfeiffer, 1857): synonym of '' Truncatella marginata'' Küster, 1855 * '' Truncatella thaanumi'' Clench & Turner, 1948 * '' Truncatella truncatula'' J.P.R. Draparnaud, 1801: synonym of '' Truncatella subcylindrica'' (C. Linnaeus, 1767) * '' Truncatella vincentiana'' Cotton, 1942 * † '' Truncatella wattebledi'' Benoist, 1878 * † '' Truncatella xuanchengensis'' H.-Z. Pan, 1982 * '' Truncatella yorkensis'' (Cox, 1868): synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841


Synonyms

* † ''Truncatella antediluviana'' Deshayes, 1861: synonym of † '' Bouryia antediluviana'' (Deshayes, 1861) (superseded combination) * ''Truncatella arcasiana'' Crosse, 1868: synonym of '' Taheitia arcasiana'' (Crosse, 1868)(original combination) * ''Truncatella atomus'' Philippi, 1841: synonym of '' Omalogyra atomus'' (Philippi, 1841) (original combination) * ''Truncatella aurantia'' A. Gould, 1847: synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841 * † ''Truncatella bezanconi'' Cossmann, 1892: synonym of † '' Murchisonella bezanconi'' (Cossmann, 1892) (superseded combination) * ''Truncatella bilabiata'' L. Pfeiffer, 1840: synonym of '' Truncatella pulchella'' L. Pfeiffer, 1839 (junior subjective synonym) * ''Truncatella capillacea'' L. Pfeiffer, 1859: synonym of '' Truncatella pulchella'' L. Pfeiffer, 1839 * ''Truncatella caribaeensis'' Reeve, 1842, sensu Clench & Turner, 1948: synonym of Truncatella pulchella L. Pfeiffer, 1839 (misidentification) * ''Truncatella ceylanica'' Pfeiffer, 1857: synonym of '' Truncatella marginata'' Küster, 1855 (junior synonym) * ''Truncatella concinna'' Pease, 1871: synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841 * † ''Truncatella costata'' Cossmann, 1895: synonym of '' Truncatella wattebledi'' Benoist, 1878 (invalid; not L. Pfeiffer 1839) * ''Truncatella costata'' L. Pfeiffer, 1839: synonym of '' Truncatella scalaris'' (Michaud, 1830) * ''Truncatella costellifera'' Pease, 1871: synonym of '' Truncatella rustica'' Mousson, 1865 * ''Truncatella costulata'' Risso, 1826: synonym of '' Truncatella subcylindrica'' (Linnaeus, 1767) * ''Truncatella cristata'' Crosse, 1868: synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841 * ''Truncatella cumingii'' C. B. Adams, 1845: synonym of '' Truncatella scalaris'' (Michaud, 1830) * † ''Truncatella cylindrata'' Briart & Cornet, 1887: synonym of † '' Bouryia cylindrata'' (Briart & Cornet, 1887) (superseded combination) * ''Truncatella debilis'' Mousson, 1873: synonym of '' Truncatella subcylindrica'' (Linnaeus, 1767) * ''Truncatella desnoyersii'' (Payraudeau, 1826): synonym of '' Truncatella subcylindrica'' (Linnaeus, 1767) * † ''Truncatella distensa'' Cossmann, 1888: synonym of † '' Bouryia distensa'' (Cossmann, 1888) (superseded combination) * ''Truncatella elongata'' L. Pfeiffer, 1856: synonym of '' Geomelania elongata'' (L. Pfeiffer, 1856) * ''Truncatella fasciata'' Tapparone Canefri, 1886: synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841 * ''Truncatella ferruginea'' (Cox, 1868): synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841 * ''Truncatella filicosta'' Poey, 1858: synonym of '' Geomelania elongata'' (L. Pfeiffer, 1856) * ''Truncatella floridana'' Hubricht, 1983: synonym of '' Truncatella pulchella'' L. Pfeiffer, 1839 * ''Truncatella funiculus'' Mousson, 1870: synonym of '' Taheitia funiculus'' (Mousson, 1870) (original combination) * ''Truncatella fusca'' Philippi, 1841: synonym of '' Setia fusca'' (Philippi, 1841) * ''Truncatella futunaensis'' Mousson, 1871: synonym of '' Truncatella rustica'' Mousson, 1865 (junior subjective synonym) * ''Truncatella gracilenta'' W. G. Binney, 1858: synonym of '' Truncatella californica'' L. Pfeiffer, 1857 (nomen nudum) * ''Truncatella gracilenta'' E. A. Smith, 1897: synonym of '' Taheitia gracilenta'' (E. A. Smith, 1897) (original combination) * ''Truncatella haitensis'' Weinland, 1876: synonym of '' Geomelania haitensis'' (Weinland, 1876) (original combination) * ''Truncatella japonica'' Pilsbry & Y. Hirase, 1905: synonym of '' Truncatella pfeifferi'' E. von Martens, 1861 (junior synonym) * ''Truncatella juliae'' de Folin, 1872: synonym of '' Parthenina juliae'' (de Folin, 1872) (original combination) * ''Truncatella kiusiuensis'' Pilsbry, 1902: synonym of '' Truncatella pfeifferi'' E. von Martens, 1861 (junior synonym) * ''Truncatella labiosa'' Souverbie, 1862: synonym of ''Truncatella teres'' L. Pfeiffer, 1857: synonym of '' Truncatella marginata'' Küster, 1855 (junior subjective synonym) * ''Truncatella laevigata'' Risso, 1826: synonym of '' Truncatella subcylindrica'' (Linnaeus, 1767) * ''Truncatella laevissima'' Kuroda, 1956: synonym of '' Truncatella pfeifferi'' E. von Martens, 1861 * ''Truncatella lirata'' Poey, 1858: synonym of '' Geomelania lirata'' (Poey, 1858) * ''Truncatella lowei'' Shuttleworth, 1852: synonym of '' Truncatella subcylindrica'' (Linnaeus, 1767) * ''Truncatella lubrica'' Held, 1847: synonym of '' Platyla polita'' (W. Hartmann, 1840) (junior synonym) * ''Truncatella manchurica'' (A. Adams, 1861): synonym of '' Cecina manchurica'' A. Adams, 1861 (superseded combination) * ''Truncatella mariannarum'' Quadras & Möllendorff, 1894: synonym of ''
Taheitia mariannarum ''Taheitia mariannarum'' is a species of very small land snail that has an operculum and lives very near saltwater, a maritime terrestrial gastropod mollusc in the family Truncatellidae. This species is endemic to Guam Guam ( ; ) is an is ...
'' (Quadras & Möllendorff, 1894) (original combination) * ''Truncatella micra'' Tenison Woods, 1878: synonym of '' Truncatella scalarina'' Cox, 1867 (junior synonym) * ''Truncatella microlena'' Monterosato, 1878: synonym of '' Truncatella subcylindrica'' (Linnaeus, 1767) * † ''Truncatella minor'' Briart & Cornet, 1887: synonym of † '' Bouryia minor'' (Briart & Cornet, 1887) (superseded combination) * ''Truncatella minuscula'' de Folin, 1875: synonym of '' Graphis albida'' (Kanmacher, 1798) (synonym) * ''Truncatella modesta'' C. B. Adams, 1851: synonym of '' Turbonilla rixtae'' De Jong & Coomans, 1988 (junior homonym) * ''Truncatella montagui'' R. T. Lowe, 1832: synonym of '' Truncatella subcylindrica'' (Linnaeus, 1767) * ''Truncatella pacifica'' Pease, 1868: synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841 * † ''Truncatella parisiensis'' Deshayes, 1861: synonym of † '' Bouryia parisiensis'' (Deshayes, 1861) * ''Truncatella pulchella'' Pfeiffer, 1839"Mollusca"
. Diversidad Biológica Cubana, accessed 23 March 2011.
- beautiful truncatella - synonyms: ''Truncatella bilabiata'', ''Truncatella bilabiata bilabiata'', ''Truncatella bilabiata barbadensis''. * ''Truncatella pellucida'' Dohrn, 1860:Rosenberg, G. (2011). ''Truncatella pellucida'' Dohrn, 1860. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=557948 on 2011-06-13 synonym of ''Truncatella marginata f. pellucida'' Dohrn, 1860: synonym of '' Truncatella marginata'' Küster, 1855 * ''Truncatella porrecta'' A. Gould, 1847: synonym of '' Taheitia porrecta'' (A. Gould, 1847) (original combination) * ''Truncatella princeps'' Dohrn, 1866: synonym of '' Truncatella rostrata'' A. Gould, 1847 (junior synonym) * ''Truncatella punctata'' Monterosato, 1878: synonym of '' Truncatella subcylindrica'' (Linnaeus, 1767) * ''Truncatella regina'' Hubricht, 1983: synonym of '' Truncatella clathrus'' R. T. Lowe, 1832 * ''Truncatella scalariformis'' C. B. Adams, 1845: synonym of '' Truncatella scalaris'' (Michaud, 1830) * ''Truncatella scalariformis'' Reeve, 1842: synonym of '' Taheitia scalariformis'' (Reeve, 1842) (original combination) * ''Truncatella schneideri'' I. Rensch, 1937: synonym of '' Taheitia schneideri'' (Rensch, 1937) * ''Truncatella semicostulata'' Jickeli, 1874: synonym of ''Truncatella marginata f. pellucida'' Dohrn, 1860: synonym of '' Truncatella marginata'' Küster, 1855 * ''Truncatella stimpsonii'' Stearns, 1872: synonym of '' Truncatella californica'' L. Pfeiffer, 1857 * ''Truncatella striata'' Reeve, 1842: synonym of '' Coxiella striata'' (Reeve, 1842) (original combination) * ''Truncatella striatula'' Menke, 1843: synonym of '' Coxiella striatula'' (Menke, 1843) (original combination) * ''Truncatella subauriculata'' Quadras & Möllendorff, 1894: synonym of ''
Taheitia mariannarum ''Taheitia mariannarum'' is a species of very small land snail that has an operculum and lives very near saltwater, a maritime terrestrial gastropod mollusc in the family Truncatellidae. This species is endemic to Guam Guam ( ; ) is an is ...
'' (Quadras & Möllendorff, 1894) (junior synonym) * ''Truncatella succinea'' C. B. Adams, 1845: synonym of '' Truncatella caribaeensis'' Reeve, 1842 (junior subjective synonym) * ''Truncatella tatarica'' Schrenck, 1867: synonym of '' Cecina tatarica'' (Schrenck, 1867) (original rank) * ''Truncatella teres'' L. Pfeiffer, 1857: synonym of '' Truncatella marginata'' Küster, 1855 * Truncatella truncatula (Draparnaud, 1801): synonym of '' Truncatella subcylindrica'' (Linnaeus, 1767) (junior synonym) * ''Truncatella turricula'' Mousson, 1870: synonym of '' Taheitia turricula'' (Mousson, 1870) (original combination) * ''Truncatella ultima'' I. Rensch, 1937: synonym of '' Taheitia ultima'' (I. Rensch, 1937) (original combination) * ''Truncatella valida'' L. Pfeiffer, 1846: synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841 (junior synonym) * ''Truncatella ventricosa'' Reeve, 1842: synonym of '' Tomichia ventricosa'' (Reeve, 1842) (original combination) * ''Truncatella vitiana'' A. Gould, 1847: synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841 * ''Truncatella wallacei'' H. Adams, 1865: synonym of '' Taheitia wallacei'' (H. Adams, 1865) (original combination) * † ''Truncatella weixianensis'' Youluo, 1978 (unavailable name) * ''Truncatella wrighti'' L. Pfeiffer, 1862: synonym of '' Geomelania lirata'' (Poey, 1858) * ''Truncatella yorkensis'' Cox, 1868: synonym of '' Truncatella guerinii'' A. Villa & J. B. Villa, 1841


References


External links


Risso A. (1826). Histoire naturelle des principales productions de l'Europe méridionale et particulièrement de celles des environs de Nice et des Alpes Maritimes, vol. 4. Paris: Levrault. vii + 439 pp., pls 1-12

Issel, A. (1880). Crociera del Violante comandato dal capitano armatore Enrico d'Albertis dureante l'anno 1877. I. Parte narrativa (pp. 199-236). II. Risultati Scientifici. Cenni sulla geologia della Galita (pp. 237-258). Molluschi terrestri e d'acqua dolce, viventi e fossili (pp. 259-282). Annali del Museo civico di storia naturale di Genova. (1) 15: 199-282

Risso, A. (1826-1827). Histoire naturelle des principales productions de l'Europe Méridionale et particulièrement de celles des environs de Nice et des Alpes Maritimes. Paris, F.G. Levrault. 3(XVI): 1-480, 14 pls.

Leach, W. E. (1852). Molluscorum Britanniae Synopsis. A synopsis of the Mollusca of Great Britain arranged according to their natural affinities and anatomical structure. Van Voorst, London, viii + 376 pp. (edited posthumously by J. E. Gray)

Guppy, R. J. L. (1871). Notes on some new forms of terrestrial and fluviatile Mollusca found in Trinidad. American Journal of Conchology. 6(4): 306–311.

Clench, W. J. & Turner, R. D. (1948). A catalogue of the family Truncatellidae with notes and descriptions of new species. Occasional Papers on Mollusks. 1(13): 157–212, 4 plates.

Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca. in: Costello, M.J. et al. (eds), European Register of Marine Species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Patrimoines Naturels. 50: 180-213

Pilsbry, H. A. & Brown, A. P. (1914). The method of progression in Truncatella. Proceedings of the Academy of Natural Sciences of Philadelphia. 66: 426-428

Dall, W. H. (1905). Fossils of the Bahama Islands, with a list of the non-marine mollusks. In: G. B. Shattuck (ed.). The Bahama Islands. The Geographical Society of Baltimore, The Johns Hopkins Press, The MacMillan Co., New York, i-xxxii, 630 pp., 93 pls. pp. 21-48, pls. 11-13

van Hyning, T. (1942). Record find of Truncatella. The Nautilus. 56(1): 34

Pilsbry, H. A. (1926). Truncatella versus Acmea. The Nautilus. 40(1): 32-33

"Genus summary for ''Truncatella''"
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