Tropidoclonion
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''Tropidoclonion'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Natricinae The Natricinae are a subfamily of colubroid snakes, sometimes referred to as a family (Natricidae). The subfamily comprises 36 genera. Members include many very common snake species, such as the European grass snakes, and the North American wa ...
of the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Colubridae Colubridae (, commonly known as colubrids , from , 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest fossil species of the family date back to the Late Eocene epoch, with earlier origins suspected. C ...
. The genus is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
, containing the sole
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
''Tropidoclonion lineatum'', commonly known as the lined snake. The species is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
.


Common names

Additional common names for ''T. lineatum'' include common snake, dwarf garter snake, grass snake, line snake, ribbon snake, streaked snake, striped snake, and swamp snake.


Subspecies

Four
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognized as being valid, including the
nominotypical subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
. *''Tropidoclonion lineatum annectens'' – central lined snake *''Tropidoclonion lineatum lineatum'' – northern lined snake *''Tropidoclonion lineatum mertensi'' – Mertens' lined snake *''Tropidoclonion lineatum texanum'' – Texas lined snake ''
Nota bene ( ; plural: ) is the Latin language, Latin phrase meaning ''note well''. In manuscripts, ''nota bene'' is abbreviated in upper-case as NB and N.B., and in lower-case as n.b. and nb; the editorial usages of ''nota bene'' and ''notate bene'' fi ...
'': A trinomial authority in parentheses indicates that the subspecies was originally described in a genus other than ''Tropidoclonion''.


Etymology

The
subspecific name In biology, trinomial nomenclature is the system of names for taxa below the rank of species. These names have three parts. The usage is different in zoology and botany. In zoology In zoological nomenclature, a trinomen (), trinominal name, or ...
, ''mertensi'', is in honor of German
herpetologist Herpetology (from Ancient Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is a branch of zoology concerned with the study of amphibians (including frogs, salamanders, and caecilians (Gymnophiona)) and reptiles (in ...
Robert Mertens Robert Friedrich Wilhelm Mertens (1 December 1894 – 23 August 1975) was a German herpetologist. Several taxa of reptiles are named after him.Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore ...
.


Geographic range

The lined snake is found throughout the central
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
from
Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. It borders on Lake Michigan to its northeast, the Mississippi River to its west, and the Wabash River, Wabash and Ohio River, Ohio rivers to its ...
to
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
.


Habitat

The preferred
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
of ''T. lineatum'' is
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
areas with soft, moist soils.


Description

The lined snake is olive
green Green is the color between cyan and yellow on the visible spectrum. It is evoked by light which has a dominant wavelength of roughly 495570 nm. In subtractive color systems, used in painting and color printing, it is created by a com ...
to
brown Brown is a color. It can be considered a composite color, but it is mainly a darker shade of orange. In the CMYK color model used in printing and painting, brown is usually made by combining the colors Orange (colour), orange and black. In the ...
with a distinctive tan or
yellow Yellow is the color between green and orange on the spectrum of light. It is evoked by light with a dominant wavelength of roughly 575585 nm. It is a primary color in subtractive color systems, used in painting or color printing. In t ...
stripe running down the middle of the back from head to
tail The tail is the elongated section at the rear end of a bilaterian animal's body; in general, the term refers to a distinct, flexible appendage extending backwards from the midline of the torso. In vertebrate animals that evolution, evolved to los ...
. It has similar stripes, one down each side on scale rows 2 and 3. On the belly, it has a double row of clean-cut black half-moon spots running down the middle. Conant R (1975). ''A Field Guide to Reptiles and Amphibians of Eastern and Central North America, Second Edition''. Boston: Houghton Mifflin. xviii + 429 pp. + Plates 1-48. (hardcover), (paperback). (''Tropidoclonion lineatum'', pp. 166-167 + Plate 24 + Map 123). It has a narrow head and small
eye An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system. In higher organisms, the ey ...
s. Adult size is typically less than 35 cm (14 inches) in total length (including tail). However, maximum recorded total length is 53 cm (21 in). Smith HM, Brodie ED Jr (1982). ''Reptiles of North America: A Guide to Field Identification''. New York: Golden Press. 240 pp. (paperback). (''Tropidoclonion lineatum'', pp. 152-153). The keeled
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
are arranged in 19 rows at midbody. There are only 5 or 6
upper labials In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ( ...
.


Behavior

The lined snake is semi
fossorial A fossorial animal () is one that is adapted to digging and which lives primarily (but not solely) underground. Examples of fossorial vertebrates are Mole (animal), moles, badgers, naked mole-rats, meerkats, armadillos, wombats, and mole salamand ...
, spending most of its time hiding under rocks, leaf litter, logs, or buried in the soil.


Diet

The majority of the
diet Diet may refer to: Food * Diet (nutrition), the sum of the food consumed by an organism or group * Dieting, the deliberate selection of food to control body weight or nutrient intake ** Diet food, foods that aid in creating a diet for weight loss ...
of ''T. lineatum'' consists of
earthworm An earthworm is a soil-dwelling terrestrial invertebrate that belongs to the phylum Annelida. The term is the common name for the largest members of the class (or subclass, depending on the author) Oligochaeta. In classical systems, they we ...
s.


Reproduction

The lined snake is
ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a "bridging" form of reproduction between egg-laying oviparity, oviparous and live-bearing viviparity, viviparous reproduction. Ovoviviparous animals possess embryos that develo ...
, the young being born in August. The average brood is seven or eight.Force ER (1931). "Habits and Birth of the Lined Snake, ''Tropidoclonion lineatum'' (Hallowell)". ''Copeia'' 1931: 51-53. The newborn juveniles are 10–12 cm (4-4¾ in.) long at birth.


References


Further reading

* Behler JL, King FW (1979). ''The Audubon Society Field Guide to North American Reptiles and Amphibians''. New York: Alfred A. Knopf. 743 pp.657 color plates. . (''Tropidoclonion lineatum'', pp. 677-678 + Plate 507). * Conant R, Bridges W (1939). ''What Snake Is That? A Field Guide to the Snakes of the United States East of the Rocky Mountains''. (With 108 drawings by Edmond Malnate). New York and London: D. Appleton-Century. Frontispiece map + viii + 163 pp. + Plates A-C, 1-32. (''Tropidoclonion lineatum'', pp. 114–115 + Plate 21, Figure 63). * Hallowell E (1856). "Notice of a Collection of Reptiles from Kansas and Nebraska, presented to the Academy of Natural Sciences, by Dr. Hammond, U. S. A." ''Proc. Acad. Nat. Sci. Philadelphia'' 8: 238-253. (''Microps'', new genus, p. 240; ''Microps lineatus'', new species, p. 241). * Powell R, Conant R, Collins JT (2016). ''Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America, Fourth Edition''. Boston and New York: Houghton Mifflin Harcourt. xiv + 494 pp., 47 plates, 207 figures. . (''Tropidoclonion lineatum'', p. 433 + Plate 44). * Stebbins RC (2003). ''A Field Guide to Western Reptiles and Amphibians, Third Edition''. The Peterson Field Guide Series ®. Boston and New York: Houghton Mifflin. xiii + 533 pp. (paperback). (''Tropidoclonion lineatum'', p. 391 + Plate 50 + Map 168). * Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: Comstock Publishing Associates, a Division of Cornell University Press. 1,105 pp. (in 2 volumes) (Genus ''Tropidoclonion'' p. 879 + Figure 20, a-c, on p. 71; and species ''Tropidoclonion lineatum'', pp. 879–884, Figure 252, Map 62). * Zim HS, Smith HM (1956). ''Reptiles and Amphibians: A Guide to Familiar American Species: A Golden Nature Guide''. Revised Edition. New York: Simon and Schuster. 160 pp. ("Lined Snake", pp. 106, 156).


External links

*
''Tropidoclonion'' at Oklahoma SnakesLined Snake
Reptiles and Amphibians of Iowa {{Taxonbar, from=Q5082984 Natricinae Reptiles of the United States Extant Pleistocene first appearances Monotypic snake genera Taxa named by Edward Drinker Cope