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''Tritylodon'' (Greek for 3 cusped tooth) is an extinct genus of tritylodonts, one of the most advanced group of cynodont therapsids. They lived in the
Early Jurassic The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma  ...
and possibly Late Triassic periods along with
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s. They also shared many characteristics with mammals, and were once considered mammals because of overall skeleton construction. That was changed due to them retaining the vestigial reptilian jawbones and a different skull structure. Tritylodons are now regarded as non-mammalian
synapsid Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes rep ...
s.


Characteristics

If a living ''Tritylodon'' were to be seen today, it would look a lot like a large rodent. They were about long but there is no certainty about the exact weight. Their method of chewing food, a grinding motion with the bottom teeth sliding against the top teeth, resembled that of rodents as well. The bottom teeth were much like a set of cusps and the top teeth were a set of matching grooves that matched perfectly allowing this motion. There were large incisors at the very front of their mouth separated by a gap from the rest of the teeth. The incisors would stick out and remain slightly visible when the mouth was closed. The legs were directly beneath the body like mammals, unlike the earlier therapsids with sprawling limbs. These animals were burrowers; the structure of the shoulder, front limbs, and large front incisors show this. They used their incisors to help dig and unearth buried plants. The way they ate and the shape of their teeth demonstrate that ''Tritylodons'' were probably primarily
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
(though some tritylodontids show evidence of more omnivorous diets, and modern analogues like rodents tend to be more omnivorous than their dentitions lead on). Any of the Tritylodonts including ''Tritylodon'' were
warm-blooded Warm-blooded is an informal term referring to animal species which can maintain a body temperature higher than their environment. In particular, homeothermic species maintain a stable body temperature by regulating metabolic processes. The on ...
or endothermic. Like most non- placental mammalimorphs, it had epipubic bones, aiding in its erect gait but preventing the expansion of the abdomen, making it unable to go through prolonged pregnancy and instead give birth to larval young like modern
marsupials Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...
and
monotremes Monotremes () are prototherian mammals of the order Monotremata. They are one of the three groups of living mammals, along with placentals ( Eutheria), and marsupials (Metatheria). Monotremes are typified by structural differences in their bra ...
.


Habitat

The ''Tritylodons habitat was limited to the forests of South Africa, with other fossils found in the Hanson Formation of Antarctica. When the species originated, about 200 million years ago, the African area was drier and hotter. But for most of their existence the climate was tropical and wetter.


Fossils

The ''Tritylodon'' fossils in South Africa are found concentrated mainly in an area about 11,000 km² (4,250 mi²). They have been found in floodplain deposits of the Lower Jurassic Elliot Formation (upper
Karoo Supergroup The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a pe ...
). In this area there have been so many findings it has been named the Tritylodon Acme Zone. The fossil findings have all been in the Free State of South Africa. The genus ''Tritylodon'' of the Tritylodonts is restricted to the South African forms: ''Tritylodon longaevus'' and ''Tritylodon maximus''. It is suggested that ''T. maximus'' is either a large ''T. longeavus'' or a closely related species. If it is a closely related species it could possibly be ecological succession since the larger ''T. maximus'' fossils have been dated in the Sinemurian–Pliensbachian mainly less than 190 million years ago and the ''T. longaevus'' in the Hettangian–Sinemurian mainly more than 190 million years ago. With the fossil findings of each species overlapping in Sinemurian stage, the fossils show two differences, ''T. maximus'' being larger and having nine upper postcanines (neither species had canine teeth) instead of the seven teeth like ''T. longeavus''. All other structures of the two ''Tritylodon'' species were the same.http://home.arcor.de/ktdykes/jtherap.htm#tritylodon Below is a
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships:


References


External links

* * * * *https://archive.today/20130204222846/http://tritylodontidae.totallyexplained.com/ * {{Taxonbar, from=Q1441437 Prehistoric cynodont genera Jurassic synapsids of Africa Fossil taxa described in 1884 Taxa named by Richard Owen Tritylodontids