Triticum Durum
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Durum wheat (), also called pasta wheat or macaroni wheat (''Triticum durum'' or ''Triticum turgidum'' subsp. ''durum''), is a
tetraploid Polyploidy is a condition in which the cells of an organism have more than two paired sets of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two complete sets of chromosomes, one fro ...
species of
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
. It is the second most cultivated species of wheat after
common wheat Common wheat (''Triticum aestivum''), also known as bread wheat, is a cultivated wheat species. About 95% of wheat produced worldwide is common wheat; it is the most widely grown of all crops and the cereal with the highest monetary yield. Ta ...
, although it represents only 5% to 8% of global wheat production. It was developed by
artificial selection Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant ...
of the domesticated
emmer Emmer is a hybrid species of wheat, producing edible seeds that have been used as food since ancient times. The domesticated types are ''Triticum turgidum'' subsp. ''dicoccum'' and ''T. t. ''conv.'' durum''. The wild plant is called ''T. t.'' s ...
wheat strains formerly grown in Central Europe and the
Near East The Near East () is a transcontinental region around the Eastern Mediterranean encompassing the historical Fertile Crescent, the Levant, Anatolia, Egypt, Mesopotamia, and coastal areas of the Arabian Peninsula. The term was invented in the 20th ...
around 7000 BC, which developed a naked, free-threshing form. Like emmer, durum wheat is awned (with bristles). It is the predominant wheat grown in the Middle East. ''Durum'' in
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
means 'hard', and the species is the hardest of all wheats. This refers to the resistance of the grain to milling, in particular of the
starch Starch or amylum is a polymeric carbohydrate consisting of numerous glucose units joined by glycosidic bonds. This polysaccharide is produced by most green plants for energy storage. Worldwide, it is the most common carbohydrate in human diet ...
y
endosperm The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the Embryo#Pla ...
, causing
dough Dough is a malleable, sometimes elastic paste made from flour (which itself is made from grains or from leguminous or chestnut crops). Dough is typically made by mixing flour with a small amount of water or other liquid and sometimes includes ...
made from its
flour Flour is a powder made by Mill (grinding), grinding raw grains, List of root vegetables, roots, beans, Nut (fruit), nuts, or seeds. Flours are used to make many different foods. Cereal flour, particularly wheat flour, is the main ingredie ...
to be weak or "soft". This makes durum favorable for
semolina Semolina is the name given to roughly milled durum wheat mainly used in making pasta and sweet puddings. The term ''semolina'' is also used to designate coarse millings of other varieties of wheat, and sometimes other grains (such as rice or ma ...
and
pasta Pasta (, ; ) is a type of food typically made from an Leavening agent, unleavened dough of wheat flour mixed with water or Eggs as food, eggs, and formed into sheets or other shapes, then cooked by boiling or baking. Pasta was originally on ...
and less practical for flour, which requires more work than with
hexaploid Polyploidy is a condition in which the cells of an organism have more than two paired sets of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two complete sets of chromosomes, one fro ...
wheats such as common bread wheats. Despite its high
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
content, durum is not a strong wheat in the sense of giving strength to dough through the formation of a
gluten Gluten is a structural protein naturally found in certain Cereal, cereal grains. The term ''gluten'' usually refers to the elastic network of a wheat grain's proteins, gliadin and glutenin primarily, that forms readily with the addition of water ...
network. Durum contains 27% extractable wet gluten, about 3% higher than common wheat (''T. aestivum'' L.).


Taxonomy

Some authorities synonymize "durum" and ''
Triticum turgidum ''Triticum turgidum'' (with its various subspecies being known as ''pasta wheat'', ''macaroni wheat'' and ''durum wheat'') is a species of wheat. It is an annual and grows primarily in temperate areas and is native to countries around the eastern ...
''. Some reserve "durum" for ''Triticum turgidum'' subsp. ''durum''. The grain classification for durum is typically based on the
Hard Vitreous Kernel Vitreous kernel count is a way to characterize grain quality by observing the optical properties of the grains. The appearance of the grain correlates with the level of protein (at least up to protein content of 12.5%.) When the protein content i ...
(HVK) count.


Genetics

Durum wheat is a
tetraploid Polyploidy is a condition in which the cells of an organism have more than two paired sets of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two complete sets of chromosomes, one fro ...
wheat, having four sets of
chromosome A chromosome is a package of DNA containing part or all of the genetic material of an organism. In most chromosomes, the very long thin DNA fibers are coated with nucleosome-forming packaging proteins; in eukaryotic cells, the most import ...
s for a total of 28, unlike hard red winter and hard red spring wheats, which are
hexaploid Polyploidy is a condition in which the cells of an organism have more than two paired sets of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two complete sets of chromosomes, one fro ...
(six sets of chromosomes) for a total of 42. Durum wheat originated through intergeneric hybridization and
polyploid Polyploidy is a condition in which the biological cell, cells of an organism have more than two paired sets of (Homologous chromosome, homologous) chromosomes. Most species whose cells have Cell nucleus, nuclei (eukaryotes) are diploid, meaning ...
ization involving two
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, ...
(having two sets of chromosomes) grass species: '' T. urartu'' (2n=2x=14, AA genome) and a B-genome diploid related to '' Aegilops speltoides'' (2n=2x=14, SS genome) and is thus an allotetraploid (having four sets of chromosomes, from unlike parents) species. Durum—and indeed all tetraploids—lack '
allele An allele is a variant of the sequence of nucleotides at a particular location, or Locus (genetics), locus, on a DNA molecule. Alleles can differ at a single position through Single-nucleotide polymorphism, single nucleotide polymorphisms (SNP), ...
s. The only exception is found by Buerstmayr ''et al.'', 2012 on the . One of the predominant production areas of durum—Italy—has domesticated varieties with lower
genetic diversity Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species. It ranges widely, from the number of species to differences within species, and can be correlated to the span of survival for a species. It is d ...
than wild types, but ssp. ''turanicum'', ssp. ''polonicum'' and ssp. ''carthlicum'' have a level of diversity intermediate between those groups. There is evidence of an increase in the intensity of breeding after 1990.


Uses

Commercially produced dry
pasta Pasta (, ; ) is a type of food typically made from an Leavening agent, unleavened dough of wheat flour mixed with water or Eggs as food, eggs, and formed into sheets or other shapes, then cooked by boiling or baking. Pasta was originally on ...
, or , is made almost exclusively from durum
semolina Semolina is the name given to roughly milled durum wheat mainly used in making pasta and sweet puddings. The term ''semolina'' is also used to designate coarse millings of other varieties of wheat, and sometimes other grains (such as rice or ma ...
. Most home-made fresh pastas also use durum wheat or a combination of soft and hard wheats. Husked but unground, or coarsely ground, it is used to produce the
semolina Semolina is the name given to roughly milled durum wheat mainly used in making pasta and sweet puddings. The term ''semolina'' is also used to designate coarse millings of other varieties of wheat, and sometimes other grains (such as rice or ma ...
in the
couscous Couscous () is a traditional North African dish a quote: “Couscous or seksu (Image 1) in Berber language or kuskus in Arabic is by origin a Numidian (the Berber population of Numidia) dish…” of small steamed granules of rolled semolina ...
of North Africa and the
Levant The Levant ( ) is the subregion that borders the Eastern Mediterranean, Eastern Mediterranean sea to the west, and forms the core of West Asia and the political term, Middle East, ''Middle East''. In its narrowest sense, which is in use toda ...
. It is also used for Levantine dishes such as ''
tabbouleh Tabbouleh (), also transcribed tabouleh, tabbouli, tabouli, or taboulah, is a Levantine salad of finely chopped parsley, soaked bulgur, tomatoes, mint, and onion, seasoned with olive oil, lemon juice, salt and sweet pepper. Some variations a ...
'', ''
kashk Kashk, kishk, ( ''Kašk'', ), () qurut, kurut, qqet, jameed, shilanch ( Tuvan and , , , , , Tajik: қурут, , Turkish: ''kurut''), chortan ( ''chort’an''), aaruul or khuruud ( Mongolian: ''ааруул'' or ''хурууд'') is a range o ...
'', ''
kibbeh Kibbeh (, also kubba and other spellings; ; ) is a popular dish in the Arab world and the Levant in particular based on spiced lean ground meat and bulgur wheat. Kibbeh is considered to be a national dish of Lebanon and Syria. In Levantine ...
'', ''bitfun'' and the ''
bulgur Bulgur (; ; ; ), or Borghol (), is a cracked wheat foodstuff found in Egyptian cuisine, South Asian cuisine and West Asian cuisine. Characteristics Bulgur is distinct from cracked wheat, which is crushed wheat grain that, unlike bulgur, has ...
'' for
pilaf Pilaf (), pilav or pilau () is a rice dish, or in some regions, a wheat dish, whose recipe usually involves cooking in stock or broth, adding spices, and other ingredients such as vegetables or meat, and employing some technique for achieving ...
s. In
North African cuisine Maghreb cuisine is the cooking of the Maghreb region, the northwesternmost part of Africa along the Mediterranean Sea, consisting of the countries of Algeria, Libya, Mauritania, Morocco, and Tunisia. Well-known dishes from the region include '' co ...
and
Levantine cuisine Levantine cuisine is the traditional cuisine of the Levant, in the sense of the rough area of former Ottoman Syria. The cuisine has similarities with Egyptian cuisine, North African cuisine and Ottoman cuisine. It is particularly known for its ...
, it forms the basis of many
soup Soup is a primarily liquid food, generally served warm or hot – though it is sometimes served chilled – made by cooking or otherwise combining meat or vegetables with Stock (food), stock, milk, or water. According to ''The Oxford Compan ...
s, gruels, stuffings,
pudding Pudding is a type of food which can either be a dessert served after the main meal or a Savoury (dish), savoury (salty or sweet, and spicy) dish, served as part of the main meal. In the United States, ''pudding'' means a sweet, milk-based des ...
s and
pastries Pastry refers to a variety of doughs (often enriched with fat or eggs), as well as the sweet and savoury baked goods made from them. The dough may be accordingly called pastry dough for clarity. Sweetened pastries are often described as '' bak ...
. When ground as fine as flour, it is used for making
bread Bread is a baked food product made from water, flour, and often yeast. It is a staple food across the world, particularly in Europe and the Middle East. Throughout recorded history and around the world, it has been an important part of many cu ...
. In the
Middle East The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq. The term came into widespread usage by the United Kingdom and western Eur ...
, it is used for flat round breads, and in Europe and elsewhere, it can be used for
pizza Pizza is an Italian cuisine, Italian, specifically Neapolitan cuisine, Neapolitan, dish typically consisting of a flat base of Leavening agent, leavened wheat-based dough topped with tomato, cheese, and other ingredients, baked at a high t ...
or ''torte''. The use of wheat to produce pasta was described as early as the 10th century by Ibn Wahshīya of
Cairo Cairo ( ; , ) is the Capital city, capital and largest city of Egypt and the Cairo Governorate, being home to more than 10 million people. It is also part of the List of urban agglomerations in Africa, largest urban agglomeration in Africa, L ...
. The North Africans called the product ''itrīya'', from which Italian sources derived the term ''tria'' (or ''aletría'' in the case of Spanish sources) during the 15th century.


Production

''Durum wheat'' (''Triticum turgidum'' ssp. ''durum'') is the 10th most cultivated cereal worldwide, with a total production of about 38 million tons. Most of the durum grown today is amber durum, the grains of which are amber-colored due to the extra carotenoid pigments and are larger than those of other types of wheat. Durum has a yellow
endosperm The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the Embryo#Pla ...
, which gives pasta its color. When durum is milled, the endosperm is ground into a granular product called
semolina Semolina is the name given to roughly milled durum wheat mainly used in making pasta and sweet puddings. The term ''semolina'' is also used to designate coarse millings of other varieties of wheat, and sometimes other grains (such as rice or ma ...
. Semolina made from durum is used for premium
pasta Pasta (, ; ) is a type of food typically made from an Leavening agent, unleavened dough of wheat flour mixed with water or Eggs as food, eggs, and formed into sheets or other shapes, then cooked by boiling or baking. Pasta was originally on ...
s and
bread Bread is a baked food product made from water, flour, and often yeast. It is a staple food across the world, particularly in Europe and the Middle East. Throughout recorded history and around the world, it has been an important part of many cu ...
s. Notably semolina is also one of the only flours that is purposely oxidized for flavor and color. There is also a red durum, used mostly for
livestock feed Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food ...
. The cultivation of durum generates greater yield than other wheats in areas of low precipitation. Good yields can be obtained by
irrigation Irrigation (also referred to as watering of plants) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has bee ...
, but this is rarely done. In the first half of the 20th century, the crop was widely grown in
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
. Durum is one of the most important food crops in
West Asia West Asia (also called Western Asia or Southwest Asia) is the westernmost region of Asia. As defined by most academics, UN bodies and other institutions, the subregion consists of Anatolia, the Arabian Peninsula, Iran, Mesopotamia, the Armenian ...
. Although the variety of the wheat there is diverse, it is not extensively grown there, and thus must be imported. West amber durum produced in
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
is used mostly as semolina/pasta, but some is also exported to
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
for bread production. In the Middle East and North Africa, local bread-making accounts for half the consumption of durum. Some flour is even imported. On the other hand, many countries in Europe produce durum in commercially significant quantities. In India durum accounts for roughly 5% of total wheat production in the country, and is used to make products such as rava and sooji.


Processing and protein content

Durum wheat is subject to four processes: cleaning, tempering, milling and purifying. First, durum wheat is cleaned to remove foreign material and shrunken and broken kernels. Then it is tempered to a moisture content, toughening the seed coat for efficient separation of bran and endosperm. Durum milling is a complex procedure involving repetitive grinding and sieving. Proper purifying results in maximum semolina yield and the least amount of bran powder. To produce bread, durum wheat is ground into flour. The flour is mixed with water to produce dough. The quantities mixed vary, depending on the acidity of the mixture. To produce fluffy bread, the dough is mixed with
yeast Yeasts are eukaryotic, single-celled microorganisms classified as members of the fungus kingdom (biology), kingdom. The first yeast originated hundreds of millions of years ago, and at least 1,500 species are currently recognized. They are est ...
and lukewarm water, heavily kneaded to form a gas-retaining gluten network, and then
fermented Fermentation is a type of anaerobic metabolism which harnesses the redox potential of the reactants to make adenosine triphosphate (ATP) and organic end products. Organic compound, Organic molecules, such as glucose or other sugars, are Catabo ...
for hours, producing bubbles. The quality of the bread produced depends on the
viscoelastic In materials science and continuum mechanics, viscoelasticity is the property of materials that exhibit both Viscosity, viscous and Elasticity (physics), elastic characteristics when undergoing deformation (engineering), deformation. Viscous mate ...
properties of gluten, the
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
content and protein composition. Containing about 12% total protein in
defatted Degreasing, often called defatting or fat trimming, is the removal of fatty acids from an object. In culinary science, degreasing is done with the intention of reducing the fat content of a meal. Degreasing food Degreasing is often used by die ...
flour compared to 11% in common wheat, durum wheat yields 27% extractable, wet gluten compared to 24% in common wheat.


References


Sources

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Further reading

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External links


Durum Wheat Research, Grain Research Laboratory, Canadian Grain Commission
{{Authority control Middle Eastern cuisine Taxa named by René Louiche Desfontaines Wheat Wheat cultivars