Triphoridae
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Triphoridae is a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of very small
sea snail Sea snails are slow-moving marine (ocean), marine gastropod Mollusca, molluscs, usually with visible external shells, such as whelk or abalone. They share the Taxonomic classification, taxonomic class Gastropoda with slugs, which are distinguishe ...
s, marine
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
s in the informal group Ptenoglossa, within the clade
Hypsogastropoda Hypsogastropoda is a clade containing marine gastropods within the clade Caenogastropoda.MolluscaBase eds. (2021). MolluscaBase. Hypsogastropoda. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=tax ...
. Almost all the species in this family are
micromollusk A micromollusc is a shelled mollusc which is extremely small, even at full adult size. The word is usually, but not exclusively, applied to marine molluscs, although in addition, numerous species of land snails and freshwater molluscs also ...
s and almost all are left-handed in shell-coiling.Bouchet, P. (2012). Triphoridae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=131 on 2012-06-16


Distribution

Species of this family can be found worldwide, but the major part occurs in the Indo-Pacific Region. Most species live between the intertidal zone and offshore to a depth between 200 m to 500 m (exceptionally below 1,000 m).


Shell description

The shells of triphorids are small (between 2 mm and 10 mm - exceptionally 50 mm) and extremely high- spired, with numerous narrow whorls which often have distinctive sculpture. The majority of species in this family are
left-handed In human biology, handedness is an individual's preferential use of one hand, known as the dominant hand, due to and causing it to be stronger, faster or more dextrous. The other hand, comparatively often the weaker, less dextrous or simply l ...
or sinistral.


Taxonomy

Facts about their taxonomy are rather scant, complicated by the high diversity and the intra- and inter-specific variability of the species. Most triphorid collections available in Museums of Natural History are still undescribed. The following subfamilies were recognized in the taxonomy of Bouchet & Rocroi of 2005: * Iniforinae Kosuge, 1966 * Metaxiinae Marshall, 1977 * Triphorinae Gray, 1847 Gründel (1975), stated that the correct family name is Triforidae Jousseaume, 1884 and not Triphoridae, but this is only true when the genera ''Triforis'' and ''Triphora'' are placed in the same family (Kosuge, 1976). Marshall (1980) however stated that the two genera need to be placed in two different families: the Triforidae and the Triphoridae.
WoRMS The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive catalogue and list of names of marine organisms. Content The content of the registry is edited and maintained by scien ...
do not recognise these 3 subfamilies and consider them alternative representation of Triphoridae.


Genera

Genera within the family Triphoridae include: * '' Aclophora'' Laseron, 1958 * '' Aclophoropsis'' Marshall, 1983 * '' Bouchetriphora'' Marshall, 1983 * '' Cautor'' Finlay, 1927 * '' Cautotriphora'' Laws, 1940 * '' Cheirodonta'' Marshall, 1983 * '' Coriophora'' Laseron, 1958 * '' Cosmotriphora'' Olsson and Harbison, 1953 * '' Costatophora'' B. A. Marshall, 1994 * '' Differoforis'' Kosuge, 2008 * '' Euthymella'' Thiele, 1929 * '' Eutriphora'' Cotton & Godfrey, 1931 * '' Hedleytriphora'' Marshall, 1983 * '' Hypotriphora'' Cotton & Godfrey, 1931 * '' Inella'' Bayle, 1879 * '' Iniforis'' Jousseaume, 1884 * '' Isotriphora'' Cotton & Godfrey, 1931 * '' Latitriphora'' Marshall, 1983 * '' Liniphora'' Laseron, 1958 * '' Litharium'' Dall, 1924 * '' Magnosinister'' Laseron, 1954 * '' Marshallora'' Bouchet, 1985 * '' Mastonia'' Hinds, 1843 * '' Mastoniaeforis'' Jousseaume, 1884 * '' Metaxia'' Monterosato, 1884 * '' Monophorus'' Grillo, 1877 * '' Nanaphora'' Laseron, 1958 * '' Nototriphora'' Marshall, 1983 * '' Obesula'' Jousseaume, 1897 * '' Opimaphora'' Laseron, 1958 * '' Pogonodon'' Bouchet, 1997 * '' Sagenotriphora'' B.A. Marshall, 1983 * '' Seilarex'' Iredale, 1924 * '' Similiphora'' Bouchet, 1985 * '' Strobiligera'' Dall, 1924 * '' Subulophora'' Laseron, 1958 * '' Sychar'' Hinds, 1843 * '' Talophora'' Gründel, 1975 * '' Teretriphora'' Finlay, 1927 * '' Triphora'' Blainville, 1828Blainville (1828). ''Dict. Sci. Nat.'' 55: 344. * '' Viriola'' Jousseaume, 1884 * '' Viriolopsis'' Marshall, 1983 ; Genera brought into synonymy : * ''Biforina'' Bucquoy, Dautzenberg & Dollfus, 1884 : synonym of '' Monophorus'' Grillo, 1877 * ''Brucetriphora'' Beu, 2004: synonym of '' Costatophora'' B. A. Marshall, 1994 * ''Distophora'' Laseron, 1958:synonym of '' Cautor'' Finlay, 1927 * ''Epiforis'' Laseron, 1958:synonym of '' Mastoniaeforis'' Jousseaume, 1884 * ''Euthymia'' Jousseaume, 1884:synonym of '' Euthymella'' Thiele, 1929 * ''Ino'' Hinds, 1843:synonym of '' Inella'' Bayle, 1879 * ''Macrosinister'':synonym of '' Magnosinister'' Laseron, 1954 * ''Mastoniaeformis'' ic: synonym of '' Mastoniaeforis'' Jousseaume, 1884 * ''Mesophora'' Laseron, 1958: synonym of '' Coriophora'' Laseron, 1958 * ''Notosinister'' Finlay, 1926: synonym of '' Monophorus'' Grillo, 1877 * ''Orbitophora'' Laseron, 1958:synonym of '' Viriola'' Jousseaume, 1884 * ''Risbecia'' Kosuge, 1966:synonym of '' Differoforis'' Kosuge, 2008 * ''Sinistroseila'' W. R. B. Oliver, 1915: synonym of '' Viriola'' Jousseaume, 1884 * ''Solosinister'' Laseron, 1954:synonym of '' Viriola'' Jousseaume, 1884 * ''Tetraphora'' Laseron, 1958:synonym of '' Brucetriphora'' Beu, 2004 * ''Torresophora'' Laseron, 1958: synonym of '' Euthymella'' Thiele, 1929 * ''Triforis'' Deshayes, 1834:synonym of '' Triphora'' Blainville, 1828 * ''Triphoris'':synonym of '' Triphora'' Blainville, 1828


References

* Fernandes M..R., Pimenta A.D. & Leal J.H., 2013. ''Taxonomic review of Triphorinae (Gastropoda: Triphoridae) from the Vitoria-Trindade Seamount Chain, southeastern Brazil.'' The Nautilus 127(1): 1-18


External links


Serge GOFAS, Ángel A. LUQUE, Joan Daniel OLIVER, José TEMPLADO & Alberto SERRA (2021) - The Mollusca of Galicia Bank (NE Atlantic Ocean); European Journal of Taxonomy 785: 1–114
{{Taxonbar, from=Q3171656 Taxa named by John Edward Gray