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Nepomorpha is an
infraorder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classific ...
of
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s in the "true bug"
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
(
Hemiptera Hemiptera (; ) is an order of insects, commonly called true bugs, comprising more than 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from ...
). They belong to the "typical" bugs of the
suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...
Heteroptera The Heteroptera are a group of about 40,000 species of insects in the order Hemiptera. They are sometimes called "true bugs", though that name more commonly refers to the Hemiptera as a whole. "Typical bugs" might be used as a more unequivocal al ...
. Due to their
aquatic Aquatic means relating to water; living in or near water or taking place in water; does not include groundwater, as "aquatic" implies an environment where plants and animals live. Aquatic(s) may also refer to: * Aquatic animal, either vertebrate ...
habits, these
animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s are known as true water bugs. They occur all over the world outside the
polar region The polar regions, also called the frigid zones or polar zones, of Earth are Earth's polar ice caps, the regions of the planet that surround its geographical poles (the North and South Poles), lying within the polar circles. These high latitu ...
s, with about 2,000
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
altogether. The Nepomorpha can be distinguished from related Heteroptera by their missing or vestigial
ocelli A simple eye or ocellus (sometimes called a pigment pit) is a form of eye or an optical arrangement which has a single lens without the sort of elaborate retina that occurs in most vertebrates. These eyes are called "simple" to distinguish the ...
. Also, as referred to by the obsolete name Cryptocerata ("the hidden-horned ones"), their antennae are reduced, with weak muscles, and usually carried tucked against the head. Most of the
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
within this infraorder live in
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include non-salty mi ...
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s. The exceptions are members of the
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Ochteroidea Nepomorpha is an infraorder of insects in the "true bug" order (Hemiptera). They belong to the "typical" bugs of the suborder Heteroptera. Due to their aquatic habits, these animals are known as true water bugs. They occur all over the world ...
, which are found along the water's edge. Many of these insects are
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s of
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s and in some cases – like the large
water scorpion Nepidae is a family of exclusively aquatic Heteropteran insects in the order Hemiptera. They are commonly called water scorpions for their superficial resemblance to scorpions, due to their raptorial forelegs and the presence of a long slender p ...
s (Nepidae) and
giant water bug Belostomatidae is a family of freshwater hemipteran insects known as giant water bugs or colloquially as toe-biters, Indian toe-biters, electric-light bugs (because they fly to lights in large numbers), alligator ticks, or alligator fleas (in Flo ...
s (Belostomatidae) – even small
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
and
amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s. Others are
omnivore An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize t ...
s or feed on
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
s. Their mouthparts form a
rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit ** podium * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ...
as in all Heteroptera and most Hemiptera. With this, they pierce their food source to suck out fluids; some, like the
Corixidae Corixidae is a family of aquatic insects in the order Hemiptera. They are found worldwide in virtually any freshwater habitat and a few species live in saline water. There are about 500 known species worldwide, in 55 genera, including the genus ...
, are also able to chew their food to some extent, sucking up the resulting pulp. The rostrum can also be used to sting in defence; some, like the common backswimmer (''
Notonecta glauca ''Notonecta glauca'', also known as the greater water-boatman or common backswimmer, is a species of aquatic insect in the family Notonectidae. This species is found in large parts of Europe, North Africa, and east through Asia to Siberia and Chi ...
'') of the
Notonectidae Notonectidae is a Cosmopolitan distribution, cosmopolitan family of aquatic insects in the order Hemiptera, commonly called backswimmers because they swim "upside down" (inverted). They are all predators and typically range from in length. They ...
can easily pierce the skin of humans and deliver a wound often more painful than a
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...
's sting.


Systematics

The Nepomorpha probably originated around the start of the
Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between 251.9 Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which ...
, some . As evidenced by
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s such as the rather advanced Triassocoridae or the primitive water boatman '' Lufengnacta'', the radiation establishing today's superfamilies seems to have been largely complete by the end of the
Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
. There are a large number of fossil genera, but except those placed in Triassocoridae they can at least tentatively be assigned to the extant superfamilies. Though the
systematics Systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: phylogenetic trees, phylogenies). Phy ...
and
phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ...
of the higher
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
of Nepomorpha were long controversial,
cladistic analysis Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...
of
mitochondrial A mitochondrion () is an organelle found in the cells of most eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is used ...
16S and
nuclear Nuclear may refer to: Physics Relating to the nucleus of the atom: *Nuclear engineering *Nuclear physics *Nuclear power *Nuclear reactor *Nuclear weapon *Nuclear medicine *Radiation therapy *Nuclear warfare Mathematics * Nuclear space *Nuclear ...
28S rDNA 28S ribosomal RNA is the structural ribosomal RNA (rRNA) for the large subunit (LSU) of eukaryotic cytoplasmic ribosomes, and thus one of the basic components of all eukaryotic cells. It has a size of 25S in plants and 28S in mammals, hence th ...
sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is cal ...
data and
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
has more recently resolved to near-perfection. The long-accepted superfamilies are all
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
, with the exception of the
Naucoroidea Naucoridae is a small family of insects commonly known as the creeping water bugs and saucer bugs. They are similar in appearance and behavior to Belostomatidae (giant water bugs), but considerably smaller, at long. Naucoridae are found around t ...
, which is now
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
with the
Aphelocheiridae ''Aphelocheirus'' is a genus of true bugs, the only genus belonging to the family Aphelocheiridae. The genus was first described by John O. Westwood in 1833. The genus has almost cosmopolitan distribution In biogeography, a cosmopolitan ...
and Potamocoridae being split off in a new superfamily
Aphelocheiroidea Nepomorpha is an infraorder of insects in the "true bug" order (biology), order (Hemiptera). They belong to the "typical" bugs of the suborder Heteroptera. Due to their Aquatic insects, aquatic habits, these animals are known as true Heteroptera# ...
. The Cibariopectinata, a proposed
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
established on the presence of cibariopectine structures in the food-sucking pump of some of the most advanced true water bugs ( Tripartita), might indeed be monophyletic. Alternatively it might be
synonymous A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
with the Tripartita, the
Ochteroidea Nepomorpha is an infraorder of insects in the "true bug" order (Hemiptera). They belong to the "typical" bugs of the suborder Heteroptera. Due to their aquatic habits, these animals are known as true water bugs. They occur all over the world ...
having lost the cibariopectines again due to the different requirements of their (for Nepomorpha) unusual lifestyle. About seven superfamilies, in
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
ary sequence, from the most ancient to the most modern lineage, have been identified in the Infraorder Nepomorpha: * †'' Morrisonnepa'' (''
incertae sedis or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...
'':
Morrison Formation The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
,
Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
~ 151 Ma.)


Nepoidea Nepomorpha is an infraorder of insects in the "true bug" order (Hemiptera). They belong to the "typical" bugs of the suborder Heteroptera. Due to their aquatic habits, these animals are known as true water bugs. They occur all over the world ...

* Family
Belostomatidae Belostomatidae is a family of freshwater hemipteran insects known as giant water bugs or colloquially as toe-biters, Indian toe-biters, electric-light bugs (because they fly to lights in large numbers), alligator ticks, or alligator fleas (in Flo ...
– giant water bugs * Family
Nepidae Nepidae is a family of exclusively aquatic Heteropteran insects in the order Hemiptera. They are commonly called water scorpions for their superficial resemblance to scorpions, due to their raptorial forelegs and the presence of a long slender p ...
– water scorpions


Corixoidea Nepomorpha is an infraorder of insects in the "true bug" order (Hemiptera). They belong to the "typical" bugs of the suborder Heteroptera. Due to their aquatic habits, these animals are known as true water bugs. They occur all over the world ...

* Family
Corixidae Corixidae is a family of aquatic insects in the order Hemiptera. They are found worldwide in virtually any freshwater habitat and a few species live in saline water. There are about 500 known species worldwide, in 55 genera, including the genus ...
– water boatmen * Family
Micronectidae Micronectidae is a family of water boatmen often referred to as pygmy water boatmen. They were originally classified as a subfamily under Corixidae but were raised to family level by Nieser (2002). There are two subfamilies, Micronectinae with s ...
– pygmy water boatmen


Ochteroidea Nepomorpha is an infraorder of insects in the "true bug" order (Hemiptera). They belong to the "typical" bugs of the suborder Heteroptera. Due to their aquatic habits, these animals are known as true water bugs. They occur all over the world ...

;
Clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
Tripartita * Family Gelastocoridae – toad bugs * Family Ochteridae – velvety shore bugs ;Clade Cibariopectinata (disputed) * Family Triassocoridae (
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
, tentatively placed here)


Aphelocheiroidea Nepomorpha is an infraorder of insects in the "true bug" order (biology), order (Hemiptera). They belong to the "typical" bugs of the suborder Heteroptera. Due to their Aquatic insects, aquatic habits, these animals are known as true Heteroptera# ...

* Family
Aphelocheiridae ''Aphelocheirus'' is a genus of true bugs, the only genus belonging to the family Aphelocheiridae. The genus was first described by John O. Westwood in 1833. The genus has almost cosmopolitan distribution In biogeography, a cosmopolitan ...
* Family Potamocoridae


Naucoroidea Naucoridae is a small family of insects commonly known as the creeping water bugs and saucer bugs. They are similar in appearance and behavior to Belostomatidae (giant water bugs), but considerably smaller, at long. Naucoridae are found around t ...

* Family Naucoridae – creeping water bugs


Notonectoidea

* Family
Notonectidae Notonectidae is a Cosmopolitan distribution, cosmopolitan family of aquatic insects in the order Hemiptera, commonly called backswimmers because they swim "upside down" (inverted). They are all predators and typically range from in length. They ...
– backswimmers


Pleoidea Nepomorpha is an infraorder of insects in the "true bug" order (Hemiptera). They belong to the "typical" bugs of the suborder Heteroptera. Due to their aquatic habits, these animals are known as true water bugs. They occur all over the world ...

Note: sometimes included in Notonectoidea * Family
Helotrephidae Helotrephidae is a family of aquatic bugs found mainly in the tropical regions with many species in the Oriental Realm and a few from Africa, Madagascar and South America. These bugs are found swimming or walking amid submerged vegetation in stag ...
* Family
Pleidae Pleidae, the pygmy backswimmers, is a family of aquatic insects in the order Hemiptera (infraorder Nepomorpha, or "true water bugs"). There are 37 species in three genera, distributed across most of the world, except the polar regions and remote ...
– pygmy backswimmers


Footnotes


Further reading

{{Taxonbar, from=Q1079070 Insect infraorders