Trimeresurus Albolabris
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''Trimeresurus albolabris'', the white-lipped pit viper or white-lipped tree viper, is a
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
pit viper The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . or pit adders, are a subfamily (biology), subfamily of Viperidae, vipers found in Asia and the ...
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
endemic to
Southeast Asia Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
.


Taxonomy

Giannasi ''et al.'' (2001) raised '' insularis'' and '' septentrionalis'' to species level. Malhotra & Thorpe (2004) transferred this species (and a number of others) to the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Cryptelytrops''. David ''et al.'' (2011) returned it to the genus ''Trimeresurus'' and assigned it the
subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
''Trimeresurus'', creating the new combination ''Trimeresurus (Trimeresurus) albolabris''. Common names include green tree pit viper, white-lipped pit viper,Gumprecht A, Tillack F, Orlov NL,
Captain A Captain is a title, an appellative for the commanding officer of a military unit; the supreme leader or highest rank officer of a navy ship, merchant ship, aeroplane, spacecraft, or other vessel; or the commander of a port, fire or police depa ...
, Ryabov S. 2004. ''Asian Pitvipers''. GeitjeBooks Berlin. 1st Edition. 368 pp. .
white-lipped tree viper, white-lipped green pit viper and white-lipped bamboo pit viper.U.S. Navy. 1991. ''Poisonous Snakes of the World''. US Govt. New York: Dover Publications Inc. 203 pp. .


Description

Maximum total length males , females ; maximum tail length males , females . Image:Trimeresurus albolabris (adult, male).jpg, ''T. albolabris'', White-lipped pit viper (adult, male) – Kaeng Krachan National Park Image:Trimeresurus albolabris (adult, female).jpg, ''T. albolabris'', White-lipped pit viper (adult, female) – Kaeng Krachan National Park Image:Trimeresurus-albolabris-white-lipped-pit-viper-juvenile-male-kaeng-krachan-national-park.jpg, ''T. albolabris'' (juvenile/male) – Kaeng Krachan National Park Image:Trimeresurus-albolabris-white-lipped-pit-viper-juvenile-female-kaeng-krachan-national-park.jpg, ''T. albolabris'' (juvenile/female) – Kaeng Krachan National Park Image:White-lipped Pitviper.jpg, ''T. albolabris'' at the
Houston Zoo The Houston Zoo is a zoological park located within Hermann Park in Houston, Texas, United States. The zoo houses over 6,000 animals from more than 900 species. It receives around 2 million visitors each year and is the second most visited zoo ...
Head scalation consists of 10–11(12)
upper labials In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ( ...
, the first partially or completely fused to the nasal. Head scales small, subequal, feebly imbricate, smooth or weakly keeled. The
supraoculars In scaled reptiles, supraocular scales are (enlarged) scales on the crown immediately above the eye.Mallow D, Ludwig D, Nilson G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Malabar, Florida: Krieger Publishing Company. ...
are narrow (occasionally enlarged and undivided) with 8–12 interocular scales between them. Temporal scales smooth. Midbody has 29 (rarely 19) longitudinal dorsal scale rows. The ventral scales are 155–166 in males, 152–176 in females. The subcaudals are paired, 60–72 in males, 49–66 in females. The
hemipenes A hemipenis (: hemipenes) is one of a pair of intromittent organs of male squamates (snakes and lizards). Hemipenes are usually held inverted within the body, and are everted for reproduction via erectile tissue, much like that in the human peni ...
are without spines. Color pattern: green above, the side of the head below the eyes is yellow, white or pale green, much lighter than rest of head. The belly is green, yellowish or white below. A light ventrolateral stripe is present in all males, but absent in females. The end of tail is not mottled brown.


Distribution and habitat

Found in
Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
,
Bhutan Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...
,
Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
,
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
,
Cambodia Cambodia, officially the Kingdom of Cambodia, is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. It is bordered by Thailand to the northwest, Laos to the north, and Vietnam to the east, and has a coastline ...
,
Laos Laos, officially the Lao People's Democratic Republic (LPDR), is the only landlocked country in Southeast Asia. It is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and ...
,
Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
, southern
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
(
Fujian Fujian is a provinces of China, province in East China, southeastern China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capital is Fuzhou and its largest prefe ...
,
Hainan Hainan is an island provinces of China, province and the southernmost province of China. It consists of the eponymous Hainan Island and various smaller islands in the South China Sea under the province's administration. The name literally mean ...
,
Guangxi Guangxi,; officially the Guangxi Zhuang Autonomous Region, is an Autonomous regions of China, autonomous region of the China, People's Republic of China, located in South China and bordering Vietnam (Hà Giang Province, Hà Giang, Cao Bằn ...
,
Guangdong ) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...
),
Hong Kong Hong Kong)., Legally Hong Kong, China in international treaties and organizations. is a special administrative region of China. With 7.5 million residents in a territory, Hong Kong is the fourth most densely populated region in the wor ...
,
Macau Macau or Macao is a special administrative regions of China, special administrative region of the People's Republic of China (PRC). With a population of about people and a land area of , it is the most List of countries and dependencies by p ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
(
Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
,
Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...
,
Lombok Lombok, is an island in West Nusa Tenggara province, Indonesia. It forms part of the chain of the Lesser Sunda Islands, with the Lombok Strait separating it from Bali to the west and the Alas Strait between it and Sumbawa to the east. It is rou ...
,
Sumbawa Sumbawa, is an Indonesian island, located in the middle of the Lesser Sunda Islands chain, with Lombok to the west, Flores to the east, and Sumba further to the southeast. Along with Lombok, it forms the province of West Nusa Tenggara, but th ...
, Komodo,
Flores Flores is one of the Lesser Sunda Islands, a group of islands in the eastern half of Indonesia. Administratively, it forms the largest island in the East Nusa Tenggara Province. The area is 14,250 km2. Including Komodo and Rinca islands ...
,
Sumba Sumba (; ), natively also spelt as Humba, Hubba, Suba, or Zuba (in Sumba languages) is an Indonesian island (part of the Lesser Sunda Archipelago group) located in the Eastern Indonesia and administratively part of the East Nusa Tenggara pro ...
,
Roti Roti is a round flatbread originating from the Indian subcontinent. It is commonly consumed in many South Asian, Southeast Asian, Caribbean, East African, and Southeast African countries. It is made from stoneground whole-wheat flour, kno ...
,
Kisar Kisar, also known as ''Yotowawa'', is a small island in the Southwestern Moluccas in Indonesia, located to the northeast of Timor Island. The island now forms two districts ("North Kisar" and "South Kisar") within the Southwest Islands Regency o ...
,
Wetar Wetar is a tropical island which belongs to the Indonesian province of Maluku and is the largest island of the Maluku Barat Daya Islands (literally ''Southwest Islands'') Regency of the Maluku Islands. It lies east of the Lesser Sunda Islands, ...
). The type locality given is "China". The species is not found in India. Specimens from India have been recently re-identified as
Trimeresurus salazar ''Trimeresurus salazar'', also known as Salazar's pit viper, is a species of venomous, green pit viper first discovered in 2019 in the lowlands of the western part of Arunachal Pradesh, India; the fifth new reptile species to be discovered in the ...
(Vogel et al. 2022). Specimens from the Nicobar islands have been described as Trimeresurus davidi, a new species.


Diet

Its meals consist of birds, small frogs, and small
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s. This snake doesn't strike and release its prey; like many arboreal snakes, rather holds on to the prey item until it dies.


Venom

The venom is primarily hemotoxic. Results of bites from this species range from mild envenoming to death. The venom of white-lipped pitviper contains
procoagulant Coagulation, also known as clotting, is the process by which blood changes from a liquid to a gel, forming a blood clot. It results in hemostasis, the cessation of blood loss from a damaged vessel, followed by repair. The process of coagulation ...
properties. There have been numerous reported bites with few fatalities.O'Shea M. ''Venomous Snakes of the World'', p. 107.


References


Further reading

*Das I. 1999. ''Biogeography of the amphibians and reptiles of the Andaman and Nicobar Islands'', India. In: Ota, H. (ed) Tropical Island herpetofauna, Elsevier, pp. 43–77. *David P, Vogel V. 2000. "On the occurrence of ''Trimeresurus albolabris''" (Gray 1842) on Sumatra Island, Indonesia (Reptilia, Serpentes, Viperidae, Crotalinae). ''Senckenbergiana Biologica'' 80 (1/2): 225–232. *David P, Vogel G, Dubois A. 2011. "On the need to follow rigorously the Rules of the ''Code'' for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus ''Trimeresurus''" ''Lacépède'', 1804 (Reptilia: Squamata: Viperidae). Zootaxa 2992: 1–51. *Einfalt P. 2002. "''Haltung und Vermehrung von'' Trimeresurus albolabris" ''(Gray 1842)''. ''Elaphe'' 10 (4): 31–36. * Gray, JE. 1842. "Synopsis of the species of Rattle-Snakes, or Family of CROTALIDÆ." The Zoological Miscellany 2: 47–51. (''Trimeresurus albolabris'', p. 48.) *Gumprecht, A. 2001. "''Die Bambusottern der Gattung'' Trimeresurus ''Lacépède Teil IV: Checkliste der'' Trimeresurus''-Arten Thailands''". ''Sauria'' 23 (2): 25–32. *Parkinson CL. 1999. "Molecular systematics and biogeographical history of Pit Vipers as determined by mitochondrial ribosomal DNA sequences". ''Copeia'' 1999 (3): 576–586. *


External links

{{Taxonbar, from=Q263271 albolabris Snakes of Southeast Asia Reptiles of Cambodia Snakes of China Reptiles of Hong Kong Reptiles of Indonesia Reptiles of Malaysia Reptiles of Myanmar Reptiles of Thailand Snakes of Vietnam Reptiles described in 1842 Taxa named by John Edward Gray