''Lobatus raninus'',

common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...

the hawk-wing conch, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of medium to large

sea snail Sea snails are slow-moving marine (ocean), marine gastropod Mollusca, molluscs, usually with visible external shells, such as whelk or abalone. They share the Taxonomic classification, taxonomic class Gastropoda with slugs, which are distinguishe ...

, a marine

gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...

mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...

in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Strombidae Strombidae, common name, commonly known as the true conchs, is a Taxonomy (biology), taxonomic family (biology), family of medium-sized to very large sea snails in the superfamily (zoology), superfamily Stromboidea, and the Epifamily Neostromboid ...

, the true

conch Conch ( , , ) is a common name of a number of different medium-to-large-sized sea snails. Conch shells typically have a high Spire (mollusc), spire and a noticeable siphonal canal (in other words, the shell comes to a noticeable point on both ...

s.Bouchet, P. (2010). Lobatus raninus (Gmelin, 1791). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=531851 on 2011-03-21

Shell description

The maximum recorded

shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses Science Biology * Seashell, a hard outer layer of a marine ani ...

length is 121 mmWelch, J.J. (2010). "The "Island Rule" and Deep-Sea Gastropods: Re-Examining the Evidence". '' PLoS ONE'' 5(1): e8776. . or up to 130 mm.Cervigón, F. ''et al.'' (1993). ''"FAO species identification sheets for fishery purposes. Field guide to the commercial marine and brackish-water resources of the northern coast of South America"''. Rome, FAO. 513 p. Like other species in the same genus, ''Lobatus raninus'' has a robust, somewhat heavy and solid shell, with a distinct

stromboid notch The stromboid notch is an anatomical feature which is found in the gastropod shell, shell of one Taxonomy (biology), taxonomic family of medium-sized to large sea snails, the conches. Marine (ocean), Marine gastropods in the family Strombidae hav ...

. The

body whorl The body whorl is part of the morphology (biology), morphology of the gastropod shell, shell in those gastropod mollusks that possess a coiled shell. The term is also sometimes used in a similar way to describe the shell of a cephalopod mollusk ...

is dorsally ornamented by characteristic coarse spiral ridges. The posterior expansion of the flaring outer lip is always lower than the spire. The color is brownish, with several disperse white spots. Both inner and outer lips are cream or white.

Phylogeny

The phylogenetic relationships among the Strombidae have been mainly accessed on two occasions, using two methods. In 2005, Simone proposed a

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

(a tree of descent) based on an extensive

morpho Morpho may refer to: * ''Morpho'' (genus), a genus of butterflies * Morpho (VTOL), a Swiss prototype drone * Morpho (company) IDEMIA (formerly known as OT-Morpho) is a French multinational technology company headquartered in Courbevoie, Fra ...

anatomical Anatomy () is the branch of morphology concerned with the study of the internal structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old scien ...

analysis of representatives of

Aporrhaidae Aporrhaidae is a family of sea snails commonly called the "pelican's foot snails." The taxonomy of the Gastropoda by Bouchet & Rocroi, 2005Bouchet P. & Rocroi J.-P. (Ed.); Frýda J., Hausdorf B., Ponder W., Valdes A. & Warén A. 2005. ''Classif ...

, Strombidae,

Xenophoridae Xenophoridae, common name, commonly called carrier shells, is a family (biology), family of medium-sized to large sea snails, marine (ocean), marine gastropod mollusks in the clade Littorinimorpha. Distribution The Xenophorids live on sand and ...

and

Struthiolariidae Struthiolariidae is a family (biology), family of small to medium-sized sea snails, marine (ocean), marine gastropod molluscs in the superfamilia, superfamily Stromboidea.Bouchet, P.; Rosenberg, G. (2014). Struthiolariidae Gabb, 1868. Accessed th ...

, including ''L. raninus'' (there referred to as ''Tricornis raninus''). With the exception of ''

Lambis ''Lambis'' is a genus of large sea snails sometimes known as spider conchs, marine (ocean), marine gastropod mollusks in the family Strombidae, the true conch family.Bouchet, P.; Rosenberg, G. (2011)Lambis Röding, 1798 Accessed through: World Re ...

'' and '' Terebellum'', the remaining taxa were previously allocated within the genus ''

Strombus ''Strombus'' is a genus of medium to large sea snails, marine (ocean), marine gastropod molluscs in the family (biology), family Strombidae, which comprises the true conchs and their immediate relatives. The genus ''Strombus'' was named by Swedi ...

''. However, according to Simone, only '' Strombus gracilior'', ''

Strombus alatus ''Strombus alatus'', the Florida fighting conch, is a species of medium-sized, warm-water sea snail, a marine gastropod mollusk in the family Strombidae, the true conchs. Its name derives two Latin words. Strombus means, in Latin, a snail with s ...

'' and ''

Strombus pugilis ''Strombus pugilis'', common names the fighting conch and the West Indian fighting conch, is a species of medium to large sea snail, a marine gastropod mollusk in the family Strombidae, the true conchs. ''S. pugilis'' is similar in appearance t ...

'', the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

, remained within ''Strombus'', as they constituted a distinct group based on at least five

synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to ...

(traits that are shared by two or more

taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...

and their

most recent common ancestor A most recent common ancestor (MRCA), also known as a last common ancestor (LCA), is the most recent individual from which all organisms of a set are inferred to have descended. The most recent common ancestor of a higher taxon is generally assu ...

). The remaining taxa were previously considered as

subgenera In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...

, and were elevated to genus level by Simone in the end of his analysis. The genus '' Tricornis'' (now considered a synonym of ''

Lobatus ''Lobatus'' is a genus of very large sea snails, marine gastropod mollusks in the family Strombidae, the true conchs. Some of the species within this genus were previously placed in the genus '' Eustrombus''. Species Living and fossil species ...

''), in this case, only included ''T. raninus'' (now considered a synonym of ''Lobatus raninus''). A different approach, this time based on sequences of nuclear

histone H3 Histone H3 is one of the five main histones involved in the structure of chromatin in eukaryotic cells. Featuring a main globular domain and a long N-terminal end, N-terminal tail, H3 is involved with the structure of the nucleosomes of the 'b ...

and mitochondrial cytochrome-c oxidase I (COI) genes was proposed by Latiolais ''et al''. (2006). The phylogenic relations of (32 analyzed) species that used to belong or still belong in the genus ''Strombus'' and ''

'' are shown below:Latiolais J. M., Taylor M. S., Roy K. & Hellberg M. E. (2006). "A molecular phylogenetic analysis of strombid gastropod morphological diversity". ''

Molecular Phylogenetics and Evolution ''Molecular Phylogenetics and Evolution'' is a peer-reviewed scientific journal of evolutionary biology and phylogenetics. The journal is edited by E.A. Zimmer. Indexing The journal is indexed in: * EMBiology *Journal Citation Reports *Scopus ...

'' 41: 436-444.
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Distribution

This species is distributed in the

Caribbean Sea The Caribbean Sea is a sea of the Atlantic Ocean, North Atlantic Ocean in the tropics of the Western Hemisphere, located south of the Gulf of Mexico and southwest of the Sargasso Sea. It is bounded by the Greater Antilles to the north from Cuba ...

, the

Gulf of Mexico The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...

and the

Lesser Antilles The Lesser Antilles is a group of islands in the Caribbean Sea, forming part of the West Indies in Caribbean, Caribbean region of the Americas. They are distinguished from the larger islands of the Greater Antilles to the west. They form an arc w ...

Ecology

;Habitat ''S. raninus'' lives near seagrass beds, usually in shallow water. The minimum recorded depth is 0.3 m; the maximum recorded depth is 55 m. ;Feeding Like other species of the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

'', ''S. raninus'' is known to be a

herbivore A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...

References

* Rosenberg, G., F. Moretzsohn, and E. F. García. 2009.'' Gastropoda (Mollusca) of the Gulf of Mexico'', Pp. 579–699 in Felder, D.L. and D.K. Camp (eds.), Gulf of Mexico–Origins, Waters, and Biota. Biodiversity. Texas A&M Press, College Station, Texas {{DEFAULTSORT:Lobatusraninus Strombidae Gastropods described in 1791 Taxa named by Johann Friedrich Gmelin