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The Tremadocian is the lowest stage of
Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era, and the second of twelve periods of the Phanerozoic Eon (geology), Eon. The Ordovician spans 41.6 million years f ...
. Together with the later Floian Stage it forms the Lower Ordovician Epoch. The Tremadocian lasted from to million years ago. The base of the Tremadocian is defined as the
first appearance In comic books and other stories with a long history, first appearance refers to the first issue to feature a fictional character. These issues are often highly valued by collectors due to their rarity and iconic status. Reader interest in fir ...
of the
conodont Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard ...
species ''
Iapetognathus fluctivagus ''Iapetognathus fluctivagus'' is a species of denticulate cordylodan conodonts belonging to the genus '' Iapetognathus''. It existed during the Tremadocian Age ( million years ago) of the Ordovician. It is an important index fossil in biostrat ...
'' at the
Global Boundary Stratotype Section and Point A Global Boundary Stratotype Section and Point (GSSP), sometimes referred to as a golden spike, is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. ...
(GSSP) section on
Newfoundland Newfoundland and Labrador is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region of Labrador, having a total size of . As of 2025 the population ...
.


Naming

The Tremadocian is named after the village Tremadoc in
Wales Wales ( ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by the Irish Sea to the north and west, England to the England–Wales border, east, the Bristol Channel to the south, and the Celtic ...
. The name was proposed by
Adam Sedgwick Adam Sedgwick FRS (; 22 March 1785 – 27 January 1873) was a British geologist and Anglican priest, one of the founders of modern geology. He proposed the Cambrian and Devonian period of the geological timescale. Based on work which he did ...
in 1846 (as "Tremadoc group").


GSSP

The GSSP for the beginning of the Tremadocian is the Green Point section () in Gros Morne National Park, in western
Newfoundland Newfoundland and Labrador is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region of Labrador, having a total size of . As of 2025 the population ...
. It is defined as the
first appearance In comic books and other stories with a long history, first appearance refers to the first issue to feature a fictional character. These issues are often highly valued by collectors due to their rarity and iconic status. Reader interest in fir ...
of the
conodont Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard ...
species ''
Iapetognathus fluctivagus ''Iapetognathus fluctivagus'' is a species of denticulate cordylodan conodonts belonging to the genus '' Iapetognathus''. It existed during the Tremadocian Age ( million years ago) of the Ordovician. It is an important index fossil in biostrat ...
''. This horizon can be found 101.8 m above the Greenpoint section datum within bed number 23. The boundary lies within the Broom Point Member, of the Green Point Formation which is part of the
Cow Head Group The Cow Head Group is a geologic group in Newfoundland and Labrador. It preserves fossils dating back to the Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geolo ...
. The first planktonic
graptolites Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian (Miaolingian, Wuliuan) through th ...
appear 4.8 m above the first appearance of ''Iapetognathus fluctivagus'' at Greenpoint section. The Tremadocian ends with the beginning of the Floian which is defined as the first appearance of '' Tetragraptus approximatus'' at the GSSP in Diabasbrottet quarry,
Västergötland Västergötland (), also known as West Gothland or the Latinized version Westrogothia in older literature, is one of the 25 traditional non-administrative provinces of Sweden (''landskap'' in Swedish), situated in the southwest of Sweden. Vä ...
,
Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
. In 2015, the Lawson Cove section in Millard County,
Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
, was proposed as an Auxiliary boundary Stratotype Section and Point (ASSP) for the Tremadocian stage and Ordovician system. In addition to the
first appearance datum First appearance datum (FAD) is a term used by geologists and paleontologists to designate the first appearance of a species in the geologic record. FADs are determined by identifying the geologically oldest fossil discovered, to date, of a partic ...
of ''I. fluctivagus'', fossils of olenid trilobite '' Jujuyaspis'' and planktonic graptolite '' Anisograptus matanensis'' are present in a nearby section. In 2017, the Xiaoyangqiao section near the Dayangcha Village,
North China North China () is a list of regions of China, geographical region of the People's Republic of China, consisting of five province-level divisions of China, provincial-level administrative divisions, namely the direct-administered municipalities ...
, was proposed as the second ASSP for the base of Tremadocian/Lower Ordovician. The first planktonic graptolites can be found right below the ''Cordylodus lindstromi'' Conodont Zone in this section. Both ASSPs were approved through supermajority vote by the Subcommission on Ordovician Stratigraphy in 2016 and 2019, respectively. However, in 2021, the
International Union of Geological Sciences The International Union of Geological Sciences (IUGS) is an international non-governmental organization devoted to global cooperation in the field of geology. As of 2023, it represents more than 1 million geoscientists around the world. About Fo ...
(IUGS) proposed to deny the use of specific points and replace them by Standard Auxiliary Boundary Stratotypes (SABS) for more "flexible" correlations with GSSPs.


Regional stages

In North America the first stage of the Ordovician is the Gasconadian Stage. In the Baltic region, the stages corresponding to Tremadocian are the Pakerort stage (lower) and the Varangu stage (upper).


Major events

The Cambrian-Tremadocian boundary is marked by the Cambrian-Ordovician extinction event. Overall the amount of biodiversity of the
Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordov ...
was maintained. At the beginning of the Tremadocian, about 485.4 million years ago, biodiversity, which had been at a low level, began its long increase phase, known as the Great Ordovician Biodiversification Event. At the
Furongian The Furongian is the fourth and final epoch and series of the Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding ...
‒Tremadocian boundary, a
mantle plume A mantle plume is a proposed mechanism of convection within the Earth's mantle, hypothesized to explain anomalous volcanism. Because the plume head partially melts on reaching shallow depths, a plume is often invoked as the cause of volcanic ho ...
event occurred on the territory of the northwestern
Gondwana Gondwana ( ; ) was a large landmass, sometimes referred to as a supercontinent. The remnants of Gondwana make up around two-thirds of today's continental area, including South America, Africa, Antarctica, Australia (continent), Australia, Zea ...
, which is now the
Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...
. Ollo de Sapo magmatic event continued in this region further into Ordovician. Several global events are observed in sediments of the Tremadocian age: the Acerocare Regressive Event, Black Mountain Transgressive Event (both in the Early Tremadocian), Peltocare Regressive Event, Kelly Creek Regressive Event, and Ceratopyge Regressive Event (the last two in the Late Tremadocian). Lithological features of the Black Mountain event are observed in
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
and Gorny Altai,
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
. The Ceratopyge Regressive Event records in
Baltica Baltica is a paleocontinent that formed in the Paleoproterozoic and now constitutes northwestern Eurasia, or Europe north of the Trans-European Suture Zone and west of the Ural Mountains. The thick core of Baltica, the East European Craton, i ...
at the end of the ''Apatokephalus serratus'' zone. Above the disappearance of ''Ceratopyge'' fauna, sediments are presented in a more depleted form due to the decreased sea level in the Late Tremadocian. The middle of the Tremadocian witnessed an
extinction event An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp fall in the diversity and abundance of multicellular organisms. It occ ...
known as the Mid-Tremadocian Extinction Event or the Base Stairsian Mass Extinction Event, which is particularly known to have affected Baltican conodonts. This extinction event may have been caused by anoxia.


Tremadocian life

Planktonic graptolites, an important
index fossil Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. "Biostratigraphy." ''Oxford Reference: Dictionary of Biology ...
, appear during the Tremadocian. Tremadocian
cephalopods A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...
were not very different from their Cambrian predecessors. Specimens of '' Ellesmeroceras'' and possibly '' Bassleroceras'', found in Santa Rosita Formation, northwestern Argentina, show that cephalopods first migrated to the waters off western Gondwana already in the early Tremadocian. In the middle Tremadocian, cephalopods became more diverse and occupied new
ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of Resource (biology), resources an ...
s. During Tremadocian there was an exchange of fauna between
Avalonia Avalonia was a microcontinent in the Paleozoic era. Crustal fragments of this former microcontinent are terranes in parts of the eastern coast of North America: Atlantic Canada, and parts of the East Coast of the United States, East Coast of the ...
and Gondwana across the
Rheic Ocean The Rheic Ocean (; ) was an ocean which separated two major paleocontinents, Gondwana and Laurussia ( Laurentia- Baltica- Avalonia). One of the principal oceans of the Paleozoic, its sutures today stretch from Mexico to Turkey and its closure r ...
, as evidenced by the findings of morphologically similar
trilobites Trilobites (; meaning "three-lobed entities") are extinct marine arthropods that form the class Trilobita. One of the earliest groups of arthropods to appear in the fossil record, trilobites were among the most successful of all early animals, ...
of the genus '' Platypeltoides'' in Belgium, Wales (both were parts of Avalonia) and Morocco (Gondwana).


Ocean and climate

The
Early Ordovician The Early Ordovician is the first epoch (geology), epoch of the Ordovician period, corresponding to the Lower Ordovician series (stratigraphy), series of the Ordovician system. It began after the Cambrian Stage 10, Age 10 of the Furongian epoch o ...
in general was a time of transgression. The climate was slowly cooling throughout the Ordovician.


References


External links

* * * * {{Geological history, p, p, state=collapsed . Ordovician geochronology