Tree fern

Tree ferns are arborescent (tree-like) ferns that grow with a trunk elevating the fronds above ground level, making them trees. Many extant tree ferns are members of the order Cyatheales, to which belong the families Cyatheaceae (scaly tree ferns), Dicksoniaceae, Metaxyaceae, and Cibotiaceae. It is estimated that Cyatheales originated in the early Jurassic, and is the third group of ferns known to have given rise to tree-like forms. The others are the extinct '' Tempskya'' of uncertain position, and Osmundales where the extinct Guaireaceae and some members of Osmundaceae also grew into trees. In addition there were the Psaroniaceae including '' Tietea'' in the Marattiales, which is the sister group to all the leptosporangiate ferns.

Other ferns which are also tree ferns, are '' Leptopteris'' and '' Todea'' in the family Osmundaceae, which can achieve short trunks under a metre tall. Osmunda regalis is sometimes considered a tree fern. Fern species with short trunks in the genera '' Blechnum'', '' Cystodium'' and '' Sadleria'' from the order Polypodiales, and smaller members of Cyatheales like '' Calochlaena'', '' Cnemedaria'', '' Culcita'' (mountains only tree fern), ''Lophosoria'' and '' Thyrsopteris'' are also considered tree ferns. The species Ctenitis sloanei (The Florida Tree Fern) from Florida, Mexico, Bermuda and the Caribbean is sometimes called a tree fern.

Range

Tree ferns are found growing in tropical and subtropical areas worldwide, as well as cool to temperate rainforests in Australia, New Zealand and neighbouring regions (e.g. Lord Howe Island, etc.). Like all ferns, tree ferns reproduce by means of spores formed on the undersides of the fronds.

Description

The fronds of tree ferns are usually very large and multiple-pinnate. Their trunk is actually a vertical and modified rhizome, and woody tissue is absent. To add strength, there are deposits of lignin in the cell walls and the lower part of the stem is reinforced with thick, interlocking mats of tiny roots. If the crown of ''Dicksonia antarctica'' (the most common species in gardens) is damaged, it will inevitably die because that is where all the new growth occurs. But other clump-forming tree fern species, such as ''D. squarrosa'' and ''D. youngiae'', can regenerate from basal offsets or from "pups" emerging along the surviving trunk length. Tree ferns often fall over in the wild, yet manage to re-root from this new prostrate position and begin new vertical growth.

Uses

Tree-ferns have been cultivated for their beauty alone; a few, however, were of some economic application, chiefly as sources of starch. These include the '' Sphaeropteris excelsa'' of Norfolk Island that was threatened with extinction for the sake of its sago-like pith, which was eaten by pigs. It is now widely cultivated as an ornamental tree, although there is only one small wild population on Norfolk Island. '' Sphaeropteris medullaris'' (mamaku, black tree fern) also furnished a kind of sago to people living in New Zealand, Queensland and the Pacific islands. A Javanese species of ''Dicksonia'' (''D. chrysotricha'') furnishes silky hairs, which were once imported as a styptic, and the long silky or wooly hairs, abundant on the stem and frond-leaves in the various species of ''Cibotium'' have not only been put to a similar use, but in the Hawaiian Islands furnished wool for stuffing mattresses and cushions, which was formerly an article of export.

Species

250px, Transplanted '' Dicksonia antarctica'' tree ferns at Combe Martin Wildlife and Dinosaur Park, North Devon, England
image:Sphaeropteris lepifera crown.png, 250px, ''Sphaeropteris lepifera'' fern in Okinawa Prefecture, Japan
It is not certain the exact number of species of tree ferns there are, but it may be close to 600–700 species. Many species have become extinct in the last century as forest habitats have come under pressure from human intervention.
* ''Lophosoria'' (tropical America, 1 species)
* '' Metaxya'' (tropical America, 1 species)
* ''Sphaeropteris'' (tropical America, India, Southeast Asia to New Zealand, the Marquesas, and Pitcairn Island, about 120 species)
* '' Alsophila'' (pantropic area, about 230 species)
* '' Nephelea'' (tropical America, about 30 species)
* '' Trichipteris'' (tropical America, about 90 species)
* '' Cyathea'' (tropical America, Australasia, about 110 species)
* '' Cnemidaria'' (tropical America, about 40 species)
* '' Dicksonia'' (tropics and southern subtropics in Island Southeast Asia, Australasia, America, Hawaii, St. Helena, about 25 species)
* '' Cystodium'' (Island Southeast Asia, 1 species)
* '' Thyrsopteris'' (Juan Fernández Islands, 1 species)
* '' Culcita'' (tropical America, Macaronesia, Iberian Peninsula, 2 species)
* '' Cibotium'' (Southeast Asia, Hawaii, Central America, about 12 species)

References

{{reflist

External links

Flora Technical Note No. 5: Identification and management of tree ferns
from Tasmania Forest Practices Authority

Tree Fern
from the San Diego Zoo website

Ferns
Plant common names
Plants by habit