Traversodontidae is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

herbivorous A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...

cynodont Cynodontia () is a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 Megaannum, mya), and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extin ...

s. Traversodonts were primarily

Gondwana Gondwana ( ; ) was a large landmass, sometimes referred to as a supercontinent. The remnants of Gondwana make up around two-thirds of today's continental area, including South America, Africa, Antarctica, Australia (continent), Australia, Zea ...

n, with many species known from

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

and

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

. Recently, traversodonts have also been found from

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

and

North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...

. Traversodonts first appeared in the

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epoch (geology), epochs of the Triassic period (geology), period or the middle of three series (stratigraphy), series in which the Triassic system (stratigraphy), system is di ...

and diversified in the

Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...

before going extinct at the end of the epoch. The family Traversodontidae was erected by

Friedrich von Huene Baron Friedrich Richard von Hoyningen-Huene (22 March 1875 – 4 April 1969) was a German nobleman paleontologist who described a large number of dinosaurs, more than anyone else in 20th-century Europe. He studied a range of Permo-Carbonife ...

in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota,

Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...

Description

Traversodonts are members of

Gomphodontia Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent w ...

, a group of herbivorous

cynognathia Cynognathia ("dog jaw") is one of two major clades of cynodonts, the other being Probainognathia. Cynognathians included the large carnivorous genus '' Cynognathus'' and the herbivorous or omnivorous gomphodonts such as traversodontids. Cynogn ...

n cynodonts. As an adaptation toward eating plants, they have wide postcanine teeth behind large

canine Canine may refer to: Zoology and anatomy * Animals of the family Canidae, more specifically the subfamily Caninae, which includes dogs, wolves, foxes, jackals and coyotes ** ''Canis'', a genus that includes dogs, wolves, coyotes, and jackals ** Do ...

s. These postcanines are closely spaced with their crowns touching each other. Each is usually wider than it is long and is covered in several cusps. Because of their complexity, postcanine teeth are the primary means of identifying and distinguishing different species of traversodonts. Traversodonts have relatively short skulls. The snout is much narrower than the back of the skull. The tip of the snout is wider than the middle, as it constricts behind the large canines. The tooth rows of the upper jaws are inset while the

maxillae In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxillar ...

and

zygomatic arch In anatomy, the zygomatic arch (colloquially known as the cheek bone), is a part of the skull formed by the zygomatic process of temporal bone, zygomatic process of the temporal bone (a bone extending forward from the side of the skull, over the ...

es extend outward, suggesting that traversodonts may have had cheeks. Primitive features of traversodonts include costal plates around the vertebrae, which stabilize the spine. These plates were lost in more advanced traversodonts like ''

Exaeretodon ''Exaeretodon'' is an Extinction, extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are Knowledge, known, hailing from various geologic formation, formations. ''E. argentinus'' is fr ...

''.

Phylogeny

Traversodontidae belongs to a large clade of cynodonts called

Cynognathia Cynognathia ("dog jaw") is one of two major clades of cynodonts, the other being Probainognathia. Cynognathians included the large carnivorous genus '' Cynognathus'' and the herbivorous or omnivorous gomphodonts such as traversodontids. Cynogn ...

. Most

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

analyses place it as the

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to the family Trirachodontidae, and both groups are placed in the cynognathian clade

. The analysis of Kammerer ''et al.'' (2012) found strong support for two clades within Traversodontidae: the subfamily

Gomphodontosuchinae Gomphodontosuchinae is a subfamily (biology), subfamily of Triassic traversodontid cynodonts. It includes the genera ''Gomphodontosuchus'' (the type genus), ''Exaeretodon'', ''Santagnathus'', ''Siriusgnathus'', ''Menadon'', ''Proexaeretodon,'' '' ...

, which had been identified in 2008, and the newly named subfamily

Massetognathinae Massetognathinae is an extinct subfamily of cynodonts in the family Traversodontidae. It includes four species from the Middle and Late Triassic: '' Massetognathus pascuali'' from Argentina, '' Massetognathus ochagaviae'' and '' Santacruzodon hops ...

. Below is a

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

from that analysis:

Paleobiology

While most traversodonts were relatively large cynodonts, particularly Scalenodontoides with extrapolated 60cm skull making it potentially the largest nonmammalian cynodont, some Late Triassic European forms like '' Maubeugia'' and '' Habayia'' were very small. These traversodonts lived during the

Rhaetian The Rhaetian is the latest age (geology), age of the Triassic period (geology), Period (in geochronology) or the uppermost stage (stratigraphy), stage of the Triassic system (stratigraphy), System (in chronostratigraphy). It was preceded by the N ...

stage of the latest Triassic when

rising sea levels The sea level has been rising from the end of the last ice age, which was around 20,000 years ago. Between 1901 and 2018, the average sea level rose by , with an increase of per year since the 1970s. This was faster than the sea level had e ...

divided western Europe into an island

archipelago An archipelago ( ), sometimes called an island group or island chain, is a chain, cluster, or collection of islands. An archipelago may be in an ocean, a sea, or a smaller body of water. Example archipelagos include the Aegean Islands (the o ...

. Traversodontids on these islands likely became smaller as a result of

insular dwarfism Insular dwarfism, a form of phyletic dwarfism, is the process and condition of large animals evolving or having a reduced body size when their population's range is limited to a small environment, primarily islands. This natural process is disti ...

Dental microwear Dental microwear analysis is a method to infer diet and behavior in extinct animals, especially in fossil specimens. It has been used on a variety of taxa, including hominids, victoriapithecids, amphicyonids, canids, ursids, hyaenids, hyaenodont ...

evidence Evidence for a proposition is what supports the proposition. It is usually understood as an indication that the proposition is truth, true. The exact definition and role of evidence vary across different fields. In epistemology, evidence is what J ...

reveals that traversodontids appear to have fed on relatively abrasive plant material relative to basal

sauropodomorphs Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lo ...

, which fed on softer vegetation, thus effectively partitioning their environment's resources with them.

References

* Much of this information comes fro
Traversodontidae
{{Taxonbar, from=Q972010 Middle Triassic first appearances Late Triassic extinctions Prehistoric therapsid families

Description

Phylogeny

Paleobiology

See also

References