The tourmaline sunangel (''Heliangelus exortis'') is a species of

hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...

in the "coquettes", tribe

Lesbiini Lesbiini is one of the two Tribe (biology), tribes that make up the subfamily Lesbiinae in the hummingbird family (biology), family Trochilidae. The other tribe is Heliantheini (brilliants). The informal name "coquettes" has been proposed for th ...

of subfamily

Lesbiinae Lesbiinae is one of the six subfamilies that make up the hummingbird family Trochilidae. The subfamily is divided into two tribes: Heliantheini ("brilliants") containing 14 genera and Lesbiini ("coquettes") containing 18 genera. Phylogeny A mol ...

. It is found in

Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...

and

Ecuador Ecuador, officially the Republic of Ecuador, is a country in northwestern South America, bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Province which contain ...

.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved May 27, 2021

Taxonomy and systematics

The tourmaline sunangel is

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

. However, at times it has been considered

conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organism ...

with the flame-throated sunangel (''H. micraster'').Heynen, I. and P. F. D. Boesman (2020). Tourmaline Sunangel (''Heliangelus exortis''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.tousun1.01 retrieved January 21, 2022

Description

The tourmaline sunangel is long. Males weigh and females . Their bill is blackish, straight, and of medium length. Both sexes have dark shining green upperparts and dusky gray underparts with round green spots. Their tails' central feathers are dark bronze and the others blackish. Adult males have a glittering green frontlet just above the bill, a dark purple-blue chin, and a glittering rosy pink

gorget A gorget ( ; ) was a band of linen wrapped around a woman's neck and head in the English medieval clothing, medieval period or the lower part of a simple chaperon (headgear), chaperon hood. The term later described a steel or leather Collar (c ...

with a glittering emerald green border below it. Adult females lack the green frontlet, their chin is blackish, and their throat whitish with green to dusky speckles and sometimes a few glittering rosy feathers. Juveniles are like the adult female but the male has a white chin.

Distribution and habitat

The tourmaline sunangel is found in all three

Andes The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the List of longest mountain chains on Earth, longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range ...

ranges of Colombia and south on the eastern slope to Ecuador's

Morona-Santiago Province Morona Santiago () is a province in Ecuador. The province was established on February 24, 1954. The capital is Macas. Economy The provincial economy is industrially unexploited to its potential due to poor means of transportation. Its economy ...

. It inhabits the interior and edges of humid to wet premontane forest, especially

cloudforest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest, is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud cover, ...

and

elfin forest A "natural National Park in the Philippines, a UNESCO World Heritage Site file:Mount Kemiri (8187817161).jpg, An elfin forest in Sumatra's Gunung Leuser National Park Dwarf forest, elfin forest, or pygmy forest is an uncommon ecosystem feat ...

. It also occurs in bushy clearings and shrubby pastures. In elevation it mainly ranges between but is found as low as .

Behavior

Movement

The tourmaline sunangel is almost entirely sendentary but in Colombia makes occasional movements into the tropical zone.

Feeding

The tourmaline sunangel is aggressively territorial at feeding locations. It takes nectar from a variety of flowering plants, usually those low to the ground both in semi-open landscapes and within the forest. It usually clings to flowers to feed rather than hovering at them. It captures insects by hawking from a perch and by gleaning from vegetation.

Breeding

The tourmaline sunangel's breeding season appears to span from March to August. The clutch of two white eggs is incubated by the female. Nothing else is known about the species' breeding

phenology Phenology is the study of periodic events in biological life cycles and how these are influenced by seasonal and interannual variations in climate, as well as habitat factors (such as elevation). Examples include the date of emergence of leav ...

Vocalization

A vocalization believed to be the tourmaline sunangel's song is "a repeated series of single notes...typically a few high-pitched upslurred notes followed by a slightly lower-pitched downslurred note, 'suwee....suwee..tsew...suwee...tsew...'." Its call is "a short dry gravelly trill 'trrr', repeated at intervals" and it also makes a continuous dry chatter.

Status

The

IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...

has assessed the tourmaline sunangel as being of Least Concern. Its population size is not known but is believed to be stable. It is locally fairly common and occurs in several protected areas.

References

{{Taxonbar, from=Q1263086 tourmaline sunangel Birds of the Colombian Andes Birds of the Ecuadorian Andes tourmaline sunangel tourmaline sunangel Taxonomy articles created by Polbot