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''Toona sureni'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of tree in the mahogany family. It is native to
South Asia South Asia is the southern Subregion#Asia, subregion of Asia that is defined in both geographical and Ethnicity, ethnic-Culture, cultural terms. South Asia, with a population of 2.04 billion, contains a quarter (25%) of the world's populatio ...
,
Indochina Mainland Southeast Asia (historically known as Indochina and the Indochinese Peninsula) is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to th ...
,
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms. It is a phytogeographical floristic region in the Paleotropical kingdom. It was first recognized as a distinct region ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, and
Papua New Guinea Papua New Guinea, officially the Independent State of Papua New Guinea, is an island country in Oceania that comprises the eastern half of the island of New Guinea and offshore islands in Melanesia, a region of the southwestern Pacific Ocean n ...
. It is commonly known as the suren toon, surian, limpaga, iron redwood or the red cedar (a name also shared with various other trees). It is also known as the Indonesian mahogany or the Vietnamese mahogany. The species is a valuable timber tree.


Taxonomy

''Toona sureni'' belongs to the toon
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Toona'' of the
mahogany Mahogany is a straight- grained, reddish-brown timber of three tropical hardwood species of the genus ''Swietenia'', indigenous to the AmericasBridgewater, Samuel (2012). ''A Natural History of Belize: Inside the Maya Forest''. Austin: Universit ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Meliaceae Meliaceae, the mahogany family, is a flowering plant family of mostly trees and shrubs (and a few herbaceous plants, mangroves) in the order Sapindales. They are characterised by alternate, usually pinnate leaves without stipules, and by syncarp ...
. It was first described by the German-Dutch botanist
Carl Ludwig Blume Charles Ludwig de Blume or Karl Ludwig von Blume (9 June 1796 – 3 February 1862) was a German-Dutch botanist and entomologist who spent most of his professional life in the Netherlands and the Dutch East Indies. As deputy director of agricul ...
in 1823 as ''Swietenia sureni''. It was transferred to the genus ''Toona'' in 1917 by the American botanist
Elmer Drew Merrill Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
.


Description

''Toona sureni'' is a medium-sized to large tree, reaching a maximum height of around and a diameter of . The bole possesses
buttress root Buttress roots, also known as plank roots or stilt roots, are large, wide roots on all sides of a shallowly rooted tree. Typically, they are found in nutrient-poor tropical forest soils that may not be very deep. They may prevent the tree from fa ...
s up to a height of and remains unbranched and straight up to a height of . The bark is fibrous and flaky in texture with numerous vertical fissures. It is pale brown to whitish or grayish brown in color. When cut, the bark and sapwood produce a pleasant odor reminiscent of
cinnamon Cinnamon is a spice obtained from the inner bark of several tree species from the genus ''Cinnamomum''. Cinnamon is used mainly as an aromatic condiment and flavouring additive in a wide variety of cuisines, sweet and savoury dishes, biscuits, b ...
. The sap is colorless and does not change in color after being exposed to air. The sapwood of ''Toona sureni'' is white to pinkish or light red in color, while the
heartwood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
is light red or brown. The leaves are
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...
and large, with a length of about . They are arranged spirally, usually clustered at the ends of twigs. The leaflets are lanceolate to ovate-lanceolate in shape. They are arranged somewhat opposite each other, usually in pairs of 6 to 9, with a maximum of 12 pairs. The upper and lower surfaces of the leaflet midribs are characteristically hairy (pilose). The
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
are terminal, occurring at the tips of branchlets. They are paniculate and pendant, reaching a length of around . The individual flowers are small (around in length), and are sweetly fragrant. Flowers are individually unisexual, though both male and female flowers occur in the same plant. The fruits are leathery capsules, around long and brown in color. Each contains more than 100 seeds. The seeds are narrow and usually in length, with a maximum length of ; and in diameter. They are winged at both ends (subequal).


Ecology

''Toona sureni'' is
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
, shedding their leaves during the dry season (usually February to March or September to October). They produce flowers and fruits twice each year (usually during December to February and April to September).


Distribution and habitat

''Toona sureni'' is native to South Asia (
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
,
Bhutan Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...
, and
Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
); Indochina (
Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
,
Laos Laos, officially the Lao People's Democratic Republic (LPDR), is the only landlocked country in Southeast Asia. It is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and ...
,
Cambodia Cambodia, officially the Kingdom of Cambodia, is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. It is bordered by Thailand to the northwest, Laos to the north, and Vietnam to the east, and has a coastline ...
,
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
, and
Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
); China (
Guizhou ) , image_skyline = , image_caption = , image_map = Guizhou in China (+all claims hatched).svg , mapsize = 275px , map_alt = Map showing the location of Guizhou Province , map_caption = Map s ...
,
Hainan Hainan is an island provinces of China, province and the southernmost province of China. It consists of the eponymous Hainan Island and various smaller islands in the South China Sea under the province's administration. The name literally mean ...
,
Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
, and
Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
); Malesia (
Malaysia Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
, and the
Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...
); and
Papua New Guinea Papua New Guinea, officially the Independent State of Papua New Guinea, is an island country in Oceania that comprises the eastern half of the island of New Guinea and offshore islands in Melanesia, a region of the southwestern Pacific Ocean n ...
. They are usually found in
primary forest An old-growth forest or primary forest is a forest that has developed over a long period of time without Disturbance (ecology), disturbance. Due to this, old-growth forests exhibit unique ecological features. The Food and Agriculture Organizati ...
s on open hillsides, slopes, ravines, and riverbanks at an altitude of asl. They may sometimes also be found in secondary semi-
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, live oak, acacias, magnolia, and banksia in more temperate zo ...
s.


Names

The tree is commonly known in English as the "suren toon" (or "suren toona"), "surian", "limpaga", "iron redwood" or the "red cedar". It is also sometimes known as the "Indonesian mahogany" or the "Vietnamese mahogany", though it is not "genuine" mahogany (genus ''
Swietenia ''Swietenia'' is a genus of trees in the Melia azedarach, chinaberry family, Meliaceae. It occurs natively in the Neotropics, from southern Florida, the Caribbean, Mexico and Central America south to Bolivia. The genus is named for Dutch-Austria ...
''). Local names include ''suren'' in Indonesia, ''ye tama'' in Burma, ''danupra'' in the Philippines, ''surian'' (สุเหรียน) in Thailand, ''surian wangi'' in Malaysia, and ''zi chun'' (紫椿) in China.


Uses

''Toona sureni'', like other members of the mahogany family, are valuable timber trees. They are a source of high quality commercial
hardwood Hardwood is wood from Flowering plant, angiosperm trees. These are usually found in broad-leaved temperate and tropical forests. In temperate and boreal ecosystem, boreal latitudes they are mostly deciduous, but in tropics and subtropics mostl ...
s used for high-end furniture work, interior finishing, decorative paneling, musical instruments such as
Djembe A djembe or jembe ( ; from Maninka language, Malinke ''jembe'' , N'Ko script, N'Ko: ) is a rope-tuned skin-covered goblet drum played with bare hands, originally from West Africa. According to the Bambara people in Mali, the name of the djembe ...
, and other wood crafts. The bark extract is also used in
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) refers to the knowledge, skills, and practices rooted in the cultural beliefs of various societies, especially Indigenous groups, used for maintaining health and treatin ...
as an
astringent An astringent (sometimes called adstringent) is a chemical that shrinks or constricts body tissues. The word derives from the Latin '' adstringere'', which means "to bind fast". Astringency, the dry, puckering or numbing mouthfeel caused by t ...
,
purgative Laxatives, purgatives, or aperients are substances that loosen stools and increase bowel movements. They are used to treat and prevent constipation. Laxatives vary as to how they work and the side effects they may have. Certain stimulant, lubri ...
, antirheumatic, and for treating gastrointestinal ailments like
diarrhea Diarrhea (American English), also spelled diarrhoea or diarrhœa (British English), is the condition of having at least three loose, liquid, or watery bowel movements in a day. It often lasts for a few days and can result in dehydration d ...
and
dysentery Dysentery ( , ), historically known as the bloody flux, is a type of gastroenteritis that results in bloody diarrhea. Other symptoms may include fever, abdominal pain, and a feeling of incomplete defecation. Complications may include dehyd ...
. The leaf extracts are used as
antibacterial An antibiotic is a type of antimicrobial substance active against bacteria. It is the most important type of antibacterial agent for fighting bacterial infections, and antibiotic medications are widely used in the treatment and prevention ...
poultices. They are also planted as ornamentals and
shade tree A shade tree is a large tree whose primary role is to provide shade in the surrounding environment due to its spreading canopy and crown, where it may give shelter from sunlight in the heat of the summer for people who seek recreational needs i ...
s, as well as being used for
intercropping Intercropping is a multiple cropping practice that involves the cultivation of two or more crops simultaneously on the same field, a form of polyculture. The most common goal of intercropping is to produce a greater yield on a given piece of land ...
.


References

{{Taxonbar, from=Q15304020 sureni Trees of China Flora of tropical Asia Plants described in 1823