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The tomtit (''Petroica macrocephala'') is a small
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
bird in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Petroicidae The bird family Petroicidae includes 51 species in 19 genera. All are endemic to Australasia: New Guinea, Australia, New Zealand and numerous Pacific Islands as far east as Samoa. For want of an accurate common name, the family is often called th ...
, the Australasian robins. It is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the islands of
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
, ranging across the main islands as well as several of the outlying islands. In the
Māori language Māori (; endonym: 'the Māori language', commonly shortened to ) is an Eastern Polynesian languages, Eastern Polynesian language and the language of the Māori people, the indigenous population of mainland New Zealand. The southernmost membe ...
, the North Island tomtit is known as and the South Island tomtit is known as . This bird has several other
Māori Māori or Maori can refer to: Relating to the Māori people * Māori people of New Zealand, or members of that group * Māori language, the language of the Māori people of New Zealand * Māori culture * Cook Islanders, the Māori people of the Co ...
and English names as well. There are several
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
showing considerable variation in
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
and size. The species is not threatened and has adapted to the changes made to New Zealand's
biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...
.


Taxonomy

The tomtit was formally described in 1789 by the German naturalist
Johann Friedrich Gmelin Johann Friedrich Gmelin (8 August 1748 – 1 November 1804) was a German natural history, naturalist, chemist, botanist, entomologist, herpetologist, and malacologist. Education Johann Friedrich Gmelin was born as the eldest son of Philipp F ...
in his revised and expanded edition of
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
's ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
''. He placed it with the tits in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Parus ''Parus'' is a genus of Old World birds in the tit family Paridae. It was formerly a large genus containing most of the 50 odd species in the family Paridae. The genus was split into several resurrected genera following the publication of a deta ...
'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Parus macrocephalus''. Gmelin based his account on the "great-headed titmouse" that had been described and illustrated in 1783 by the English ornithologist John Latham in his book ''A General Synopsis of Birds''. The naturalist
Joseph Banks Sir Joseph Banks, 1st Baronet, (19 June 1820) was an English Natural history, naturalist, botanist, and patron of the natural sciences. Banks made his name on the European and American voyages of scientific exploration, 1766 natural-history ...
had provided Latham with either a specimen or a painting of the tomtit. The bird was seen at Queen Charlotte Sound on the north coast of New Zealand's
South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealand by surface area, the others being the smaller but more populous North Island and Stewart Island. It is bordered to the north by ...
during
James Cook Captain (Royal Navy), Captain James Cook (7 November 1728 – 14 February 1779) was a British Royal Navy officer, explorer, and cartographer famous for his three voyages of exploration to the Pacific and Southern Oceans, conducted between 176 ...
's second voyage to the Pacific Ocean. Banks owned a painting of the bird by
Georg Forster Johann George Adam Forster, also known as Georg Forster (; 27 November 1754 – 10 January 1794), was a German geography, geographer, natural history, naturalist, ethnology, ethnologist, travel literature, travel writer, journalist and revol ...
who had accompanied Cook on the voyage. This picture is now held by the
Natural History Museum A natural history museum or museum of natural history is a scientific institution with natural history scientific collection, collections that include current and historical records of animals, plants, Fungus, fungi, ecosystems, geology, paleo ...
in London. The tomtit is now placed with 13 other species in the genus ''
Petroica ''Petroica'' is a genus of Australasian robins, named for their red and pink markings. They are not closely related to the European robins nor the American robins. The genus was introduced by the English naturalist, William Swainson, in 1829, w ...
'' that was introduced in 1829 by the English naturalist,
William Swainson William Swainson Fellow of the Linnean Society, FLS, Fellow of the Royal Society, FRS (8 October 1789 – 6 December 1855), was an English ornithologist, Malacology, malacologist, Conchology, conchologist, entomologist and artist. Life Swains ...
. The generic name combines the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''petro-'' "rock" with ''oikos'' "home". The specific epithet ''macrocephala'' is from the Ancient Greek ''makrokephalos'' meaning "great-headed". The word combines ''makros'' meaning "long" with ''kephalē'' meaning "head". The tomtit is one of four species of the genus ''Petroica'' found in New Zealand, the ancestors of which colonised from Australia. The species was once thought to have been descended from the
scarlet robin The scarlet robin (''Petroica boodang'') is a common red-breasted Australasian robin in the passerine bird genus ''Petroica''. The species is found on continental Australia and its offshore islands, including Tasmania. The species was origina ...
, although more recent research has questioned this. It seems likely that there were two colonisation events, with the North Island robin and the South Island robin descended from one event, and the
black robin The black robin or Chatham Island robin (Moriori language, Moriori: , ; ''Petroica traversi'') is an endangered bird from the Chatham Islands off the east coast of New Zealand. It is closely related to the South Island robin (''P. australis'') ...
and tomtit from another. Other English names have been used for this species including New Zealand tomtit, black tomtit, pied tomtit and yellow-breasted tit or tomtit. The term ''tomtit'' was originally a shortened form of ''Tom titmouse''. Either form has been used to describe a number of small birds, but in England ''tomtit'' was most commonly used as an alternate name of the
blue tit The Eurasian blue tit (''Cyanistes caeruleus'') is a small passerine bird in the tit (bird), tit family, Paridae. It is easily recognizable by its blue and yellow plumage and small size. Eurasian blue tits, usually resident bird, resident a ...
. The word tit is today used for a number of small birds, especially of the family
Paridae The tits, chickadees, and titmice constitute the Paridae, a family of small passerine birds which occur mainly in the Northern Hemisphere and Africa. Many were formerly classified in the genus ''Parus''. Eurasian and African members of this f ...
. Originally, it was used for any small animal or object. Five
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised. Each subspecies is restricted to one of the islands or island groups. Four of these five subspecies have been elevated to full species in the past (the Chatham subspecies was retained with the South Island tomtit), but genetic studies have shown that these subspecies diverged relatively recently. * ''P. m. toitoi'' ( Lesson, RP & Garnot, 1828) –
North Island The North Island ( , 'the fish of Māui', historically New Ulster) is one of the two main islands of New Zealand, islands of New Zealand, separated from the larger but less populous South Island by Cook Strait. With an area of , it is the List ...
and satellites (New Zealand) * ''P. m. macrocephala'' ( Gmelin, JF, 1789) –
South South is one of the cardinal directions or compass points. The direction is the opposite of north and is perpendicular to both west and east. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Proto-Germanic ''*sunþa ...
and
Stewart Island Stewart Island (, ' glowing skies', officially Stewart Island / Rakiura, formerly New Leinster) is New Zealand's third-largest island, located south of the South Island, across Foveaux Strait. It is a roughly triangular island with a la ...
and satellites (New Zealand) * ''P. m. dannefaerdi'' (
Rothschild Rothschild () is a name derived from the German ''zum rothen Schild'' (with the old spelling "th"), meaning "to the red shield", in reference to the houses where these family members lived or had lived. At the time, houses were designated by signs ...
, 1894) – Snares Islands (south of South Island, New Zealand) * ''P. m. chathamensis'' Fleming, CA, 1950 –
Chatham Islands The Chatham Islands ( ; Moriori language, Moriori: , 'Misty Sun'; ) are an archipelago in the Pacific Ocean about east of New Zealand's South Island, administered as part of New Zealand, and consisting of about 10 islands within an approxima ...
(east of
South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealand by surface area, the others being the smaller but more populous North Island and Stewart Island. It is bordered to the north by ...
, New Zealand) * ''P. m. marrineri'' ( Mathews & Iredale, 1913) –
Auckland Islands The Auckland Islands ( Māori: ''Motu Maha'' "Many islands" or ''Maungahuka'' "Snowy mountains") are an archipelago of New Zealand, lying south of the South Island. The main Auckland Island, occupying , is surrounded by smaller Adams Island ...
(south of
South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealand by surface area, the others being the smaller but more populous North Island and Stewart Island. It is bordered to the north by ...
, New Zealand)


Description

The tomtit is a small (13 cm, 11 g) bird with a large head and a short bill. The male
North Island The North Island ( , 'the fish of Māui', historically New Ulster) is one of the two main islands of New Zealand, islands of New Zealand, separated from the larger but less populous South Island by Cook Strait. With an area of , it is the List ...
subspecies has black head, back, wings (with a white wing-bar), and a white belly. The subspecies from
South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealand by surface area, the others being the smaller but more populous North Island and Stewart Island. It is bordered to the north by ...
, the
Chatham Islands The Chatham Islands ( ; Moriori language, Moriori: , 'Misty Sun'; ) are an archipelago in the Pacific Ocean about east of New Zealand's South Island, administered as part of New Zealand, and consisting of about 10 islands within an approxima ...
, and
Auckland Islands The Auckland Islands ( Māori: ''Motu Maha'' "Many islands" or ''Maungahuka'' "Snowy mountains") are an archipelago of New Zealand, lying south of the South Island. The main Auckland Island, occupying , is surrounded by smaller Adams Island ...
are similar, but have a yellow band across the breast between the black head and white belly. The females are brown instead of black. The Snares Island subspecies is entirely black, and is known as the black tit. The island subspecies of tomtits show a striking variation in body size, being considerably larger than their mainland relatives, a tendency known as the Foster's rule or the island effect. Birds from the main islands weigh around 11g, compared with birds from Snares Island, which weigh in at 20 g.


Behaviour


Food and feeding

The tomtit is mostly an
insectivore file:Common brown robberfly with prey.jpg, A Asilidae, robber fly eating a hoverfly An insectivore is a carnivore, carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the Entomophagy ...
, feeding on small invertebrates, such as
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s, caterpillars,
spider Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...
s,
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
s,
wētā Wētā (also spelled weta in English) is the common name for a group of about 100 insect species in the families Anostostomatidae and Rhaphidophoridae endemism, endemic to New Zealand. They are giant wingless insect, flightless cricket (insect ...
,
earthworm An earthworm is a soil-dwelling terrestrial invertebrate that belongs to the phylum Annelida. The term is the common name for the largest members of the class (or subclass, depending on the author) Oligochaeta. In classical systems, they we ...
s, and flies. Fruit is taken during the winter and autumn. Most subspecies feed in vegetation, waiting on a perch and watching for
prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...
.
Insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s are also gleaned from branches and leaves. The Snares subspecies feeds on the ground as well, in a similar fashion to the
North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography. Etymology T ...
or South Island robins.


Breeding

Breeding takes place from September to February. The nest is placed in a hole in a tree, in a tangle of vines and sometimes on a sheltered ledge or bank. The bulky nest is built by the female but the male sometimes brings material. The clutch usually contains 3–4 creamy white eggs which are covered with light-brown or grey spots. The eggs are laid at daily intervals soon after sunrise, and measure around . They are incubated by the female and hatch after a period of 15–17 days. The
altricial Precocial species in birds and mammals are those in which the young are relatively mature and mobile from the moment of birth or hatching. They are normally nidifugous, meaning that they leave the nest shortly after birth or hatching. Altricial ...
nidicolous chicks are initially almost naked and are brooded by the female. Both parents feed the young and remove the faecal sacs. The chicks fledge when 17 to 20 days of age. Both parents continue feeding them for another ten days, although the male will feed them alone if the female re-nests. Pairs often raise two or three broods in a season.


Cultural depictions

The miromiro features in Māori traditional stories.
Te Arawa Te Arawa is a confederation of Māori people, Māori iwi and hapū (tribes and sub-tribes) of New Zealand who trace their ancestry to the ''Arawa (canoe), Arawa'' migration canoe (''waka''). The tribes are based in the Rotorua and Bay of Plent ...
tradition tells how Te Rongorere used a pet miromiro to help him search for his mother, Te Aotepairu, who had been carried away from the
Bay of Plenty The Bay of Plenty () is a large bight (geography), bight along the northern coast of New Zealand's North Island. It stretches from the Coromandel Peninsula in the west to Cape Runaway in the east. Called ''Te Moana-a-Toitehuatahi'' (the Ocean ...
to Whangaruru in Northland.


References


Sources

* *


External links

* {{Authority control Birds described in 1789 Taxa named by Johann Friedrich Gmelin Endemic birds of New Zealand Birds of the Chatham Islands Petroica