The tokay gecko (''Gekko gecko'')
is a
nocturnal
Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite.
Nocturnal creatur ...
arboreal
Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...
gecko
Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates. They range from .
Geckos are unique among lizards ...
in the genus ''
Gekko
''Gekko'' is a genus of Southeast Asian geckos, Common name, commonly known as true geckos or calling geckos, in the Family (biology), family Gekkonidae. Although species such as ''Gekko gecko'' (tokay gecko) are very widespread and common, some ...
'', the true geckos. It is native to
Asia
Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
and some
Pacific Islands
The Pacific islands are a group of islands in the Pacific Ocean. They are further categorized into three major island groups: Melanesia, Micronesia, and Polynesia. Depending on the context, the term ''Pacific Islands'' may refer to one of several ...
.
Etymology
The word "tokay" is an
onomatopoeia
Onomatopoeia (or rarely echoism) is a type of word, or the process of creating a word, that phonetics, phonetically imitates, resembles, or suggests the sound that it describes. Common onomatopoeias in English include animal noises such as Oin ...
of the sound made by males of this species.
The common and scientific names, as well as the family name Gekkonidae and the generic term "
gecko
Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates. They range from .
Geckos are unique among lizards ...
" come from this species, too, from ''ge'kok'' in
Javanese,
corresponding to ''tokek'' in
Malay.
Subspecies
Two subspecies are currently recognized:
*''G. g. gecko'' (
Linnaeus
Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1758) occurs in tropical
Asia
Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
from northeastern
India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
to eastern
Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
.
*''G. g. azhari'' (
Mertens
__NOTOC__
Mertens () is a surname of Flanders, Flemish origin, meaning "son of Merten" (Martin (name), Martin). It is the fifth most common name in Belgium with 18,518 people in 2008.
Geographical distribution
As of 2014, 43.4% of all known bearer ...
, 1955) is found only in
Bangladesh
Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...
.
Distribution and habitat
This species is found in northeast
India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
,
Bhutan
Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...
,
Nepal
Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
, and
Bangladesh
Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...
; throughout
Southeast Asia
Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
, including
Cambodia
Cambodia, officially the Kingdom of Cambodia, is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. It is bordered by Thailand to the northwest, Laos to the north, and Vietnam to the east, and has a coastline ...
,
Thailand
Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
, the
Philippines
The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...
,
Malaysia
Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...
,
Vietnam
Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
and
Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
; and toward western
New Guinea
New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
. Its native habitat is
rainforest
Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s, where it lives on trees and cliffs, and it frequently adapts to rural human habitations, roaming walls and ceilings at night in search of insect prey. This is an
introduced species
An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived ther ...
in some areas outside its native range. It is established in
Florida
Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
in the
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
,
Martinique
Martinique ( ; or ; Kalinago language, Kalinago: or ) is an island in the Lesser Antilles of the West Indies, in the eastern Caribbean Sea. It was previously known as Iguanacaera which translates to iguana island in Carib language, Kariʼn ...
, the islands of
Belize
Belize is a country on the north-eastern coast of Central America. It is bordered by Mexico to the north, the Caribbean Sea to the east, and Guatemala to the west and south. It also shares a maritime boundary with Honduras to the southeast. P ...
, and possibly
Hawaii
Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...
. Increasing urbanization is reducing its range.
Whether the species is native but very uncommon in
Taiwan
Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...
or whether the rare reports of individuals since the 1920s are based on repeated anthropogenic translocations that may or may not have resulted in established populations by now is unclear.
Physical characteristics
The tokay gecko is a large
nocturnal
Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite.
Nocturnal creatur ...
gecko, reaching a total length (including tail) of on average, but some grow as large as long. It is believed to be the third-largest species of gecko, after the
giant leaf-tail gecko (''Uroplatus giganteus'') and
New Caledonian giant gecko (''Rhacodactylus leachianus''). It is cylindrical, but somewhat flattened in body shape. The eyes have vertical pupils.
The skin is soft to the touch and is generally blue-gray with red or orange spots and speckles, but the animal can change the color of its skin to blend into the environment. The species is
sexually dimorphic
Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
, with the males being more brightly colored and slightly larger than females.
[Corl, J. 1999]
''Gekko gecko''.
Animal Diversity Web. University of Michigan. Retrieved 19 February 2016.
They have two visual pigments: a "green" with λmax at 521 nm and a "blue" at 467 nm.
It is a strong climber with foot pads that can support the entire weight of its body on a vertical surface for a long period of time. Compared to other gecko species, the tokay gecko has a robust build, with a semiprehensile tail, a large head, and muscular jaws.
Behaviour
Tokay geckos are generally aggressive and territorial, and can inflict a strong bite. Though common in the pet trade, the strong bite of the tokay gecko makes it ill-suited for inexperienced keepers. In addition, the strength of the bite depends on the gecko's size; larger (usually male) tokay geckos are capable of piercing skin, which often results in immediate
bleeding
Bleeding, hemorrhage, haemorrhage or blood loss, is blood escaping from the circulatory system from damaged blood vessels. Bleeding can occur internally, or externally either through a natural opening such as the mouth, nose, ear, urethr ...
.
Females lay clutches of one or two hard-shelled eggs and guard them until they hatch. Communal nesting has been observed in Cambodia with as many as four adults guarding the clutch of about eleven eggs.
Diet
The tokay gecko feeds on
insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s and small
vertebrate
Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain.
The vertebrates make up the subphylum Vertebra ...
s.
[ In captivity, they usually feed on ]springtail
Springtails (class Collembola) form the largest of the three lineages of modern Hexapoda, hexapods that are no longer considered insects. Although the three lineages are sometimes grouped together in a class called Entognatha because they have in ...
s, mealworm
Mealworms are the larval form of the yellow mealworm beetle, ''Tenebrio molitor'', a species of darkling beetle.
The yellow mealworm beetle prefers a warmer climate and higher humidity. Male mealworm beetles release a sex pheromone to attract ...
s, cockroach
Cockroaches (or roaches) are insects belonging to the Order (biology), order Blattodea (Blattaria). About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known Pest (organism), pests.
Modern cockro ...
es, cricket
Cricket is a Bat-and-ball games, bat-and-ball game played between two Sports team, teams of eleven players on a cricket field, field, at the centre of which is a cricket pitch, pitch with a wicket at each end, each comprising two Bail (cr ...
s, grasshopper
Grasshoppers are a group of insects belonging to the suborder Caelifera. They are amongst what are possibly the most ancient living groups of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago.
Grassh ...
s, locust
Locusts (derived from the Latin ''locusta'', locust or lobster) are various species of short-horned grasshoppers in the family Acrididae that have a swarming phase. These insects are usually solitary, but under certain circumstances they b ...
s, and pink mice
A mouse (: mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
. In a study conducted in Thailand, researchers noticed little variation in the diets of males, females, and juveniles, which was likely due to low insect availability in this area. In Southern Florida, roaches, caterpillars, spiders, and beetles were the most numerous taxa present and common to the greatest number of stomachs.
Call
The male's mating call, a loud croak, is variously described as sounding like ''token'', ''gekk-gekk'', ''tuck-too'', ''túc-key'', ''tou-kay'' or ''tokay''. Most of the time, the call is often preceded by a quick "cackling", similar to the chirping sounds made by house gecko
''Hemidactylus'' is a genus of the common gecko family, Gekkonidae. It has 194 described species, newfound ones being described every few years. These geckos are found in all the tropical regions of the world, extending into the subtropical parts ...
s albeit much lower in pitch. When threatened or alarmed, tokay geckos usually "bark" while opening their mouth in a defensive posture.
The tokay gecko's call is also responsible for the name given to it by Filipino residents: "Tuko," and by U.S. soldiers during the Vietnam War
The Vietnam War (1 November 1955 – 30 April 1975) was an armed conflict in Vietnam, Laos, and Cambodia fought between North Vietnam (Democratic Republic of Vietnam) and South Vietnam (Republic of Vietnam) and their allies. North Vietnam w ...
, the "Fuck-you lizard".
Light and temperature can affect its vocalizations. The most frequent calling occurs in May at dusk, and the second peak of call frequency occurs in May at dawn. Vocalizations and associated behavior were strongly affected by ambient temperature in both the lab and field and could thus play a role in regulating animal energetic metabolism
Conservation and relationship with humans
The tokay gecko is culturally significant in many East Asian countries. Regional folklore has attributed supernatural powers to the gecko. In Southeast Asia
Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
it is a symbol of good luck and fertility.[ It is believed to be descended from ]dragon
A dragon is a Magic (supernatural), magical legendary creature that appears in the folklore of multiple cultures worldwide. Beliefs about dragons vary considerably through regions, but European dragon, dragons in Western cultures since the Hi ...
s.[Naish, D]
People are modifying monitors to make gargantuan geckos.
''Scientific American'' Blog 16 April 2015.
This species is poached for the medicinal trades in parts of Asia. The tokay gecko is an ingredient in traditional Chinese medicine
Traditional Chinese medicine (TCM) is an alternative medicine, alternative medical practice drawn from traditional medicine in China. A large share of its claims are pseudoscientific, with the majority of treatments having no robust evidence ...
, known as ''Ge Jie'' (蛤蚧). It is believed to nourish the kidneys and lungs, beliefs that are not substantiated by medical science. The animal remains highly sought after in China, Hong Kong, Taiwan, Vietnam, Malaysia, Singapore and other parts of Asia with Chinese communities, to the point where unscrupulous merchants have taken to disfiguring monitor lizard
Monitor lizards are lizards in the genus ''Varanus,'' the only extant genus in the family Varanidae. They are native to Africa, Asia, and Oceania, and West African Nile monitor, one species is also found in south America as an invasive species. A ...
s with prosthetics to pass them off as colossal tokay gecko specimens.[
From 2009 to 2011, the poaching of tokay geckos intensified because of a short-lived belief that they were an effective ]HIV
The human immunodeficiency viruses (HIV) are two species of '' Lentivirus'' (a subgroup of retrovirus) that infect humans. Over time, they cause acquired immunodeficiency syndrome (AIDS), a condition in which progressive failure of the im ...
cure.
The tokay gecko is quickly becoming a threatened species in the Philippines because of indiscriminate hunting. Collecting, transporting and trading in geckos without a license can be punishable by up to 12 years in jail and a fine of up to ₱
The Philippine peso sign (₱) is the currency symbol used for the Philippine peso, the official currency of the Philippines. The symbol resembles a Latin script, Latin letter P with two horizontal strokes. It differs from the currency symbol u ...
1 million under Republic Act 9147, in addition to other applicable international laws. However, the trade runs unchecked because of the sheer number of illegal traders and reports of lucrative deals. Chinese
Chinese may refer to:
* Something related to China
* Chinese people, people identified with China, through nationality, citizenship, and/or ethnicity
**Han Chinese, East Asian ethnic group native to China.
**'' Zhonghua minzu'', the supra-ethnic ...
buyers and other foreign nationals are rumored to pay thousands of dollars for large specimens, because of their alleged medicinal value or as commodities in the illegal wildlife trade.
The species is protected under Appendix II of the Convention on International Trade in Endangered Species
Convention may refer to:
* Convention (norm), a custom or tradition, a standard of presentation or conduct
** Treaty, an agreement in international law
** Convention (political norm), uncodified legal or political tradition
* Convention (meeting ...
(CITES) meaning international trade (including in parts and derivatives) is subject to the CITES permitting system.
Captivity
Tokay geckos are becoming more popular as pets because of their striking colors and large size. Most of them are wild-caught imports, but captive-bred ones are becoming more common. Wild-caught adults are simple to keep in captivity, as the husbandry requirements can be fulfilled with standard hardware that is widely available from pet-supply vendors. Wild-caught tokay geckos can be difficult to handle because of their aggressive nature and powerful bite, but it has been commonly reported among hobbyists that captive-bred (those hatched in captivity from eggs laid by either wild-caught adults or by captive-raised adult progeny) tokay geckos can be less aggressive.
When well cared for, tokay geckos can live up to 15–20 years.
References
External links
Tokay Gecko
– Reptile Parkreptilepark.com.au.
– EcologyAsia.
Tokay Gecko Care.
TokayGecko.org
The sound of a Balinese geko in Sanur. Recorded October 2004.
SoundCloud.
Nonnatives – Tokay Gecko.
Florida Fish and Wildlife Conservation Commission.
Australian Reptile Park.
{{Authority control
Geckos
Gekko
Reptiles of Southeast Asia
Reptiles of Bangladesh
Reptiles of China
Reptiles of Taiwan
Reptiles described in 1758
Taxa named by Carl Linnaeus
Geckos of Thailand
Reptiles as pets