Todus
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''Todus'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of birds found in the
Caribbean The Caribbean ( , ; ; ; ) is a region in the middle of the Americas centered around the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean, mostly overlapping with the West Indies. Bordered by North America to the north, Central America ...
. It is the only genus within the todies family Todidae. The five
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
are small birds of the forests of the
Greater Antilles The Greater Antilles is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, and Jamaica, together with Navassa Island and the Cayman Islands. Seven island states share the region of the Greater Antille ...
: Puerto Rico, Jamaica, and Cuba, with adjacent islands, have one species each, and
Hispaniola Hispaniola (, also ) is an island between Geography of Cuba, Cuba and Geography of Puerto Rico, Puerto Rico in the Greater Antilles of the Caribbean. Hispaniola is the most populous island in the West Indies, and the second-largest by List of C ...
has two, the
broad-billed tody The broad-billed tody (''Todus subulatus'') is a species of bird in the family Todidae, and one of two Todus species found on Hispaniola, along with the narrow-billed tody. They are small insectivorous birds, characterized by their bright green ...
in the lowlands (including
Gonâve Island Gonâve Island or Zile Lagonav (, ; also ''La Gonâve'') is an island of Haiti located west-northwest of Port-au-Prince in the Gulf of Gonâve. It is the largest of the Hispaniolan satellite islands. The island is an arrondissement (''Arrondis ...
) and the
narrow-billed tody The narrow-billed tody (''Todus angustirostris'') is a species of bird in the family Todidae. It is one of two Todus species endemic to Hispaniola, a Caribbean island shared by the Dominican Republic and Haiti. Taxonomy and systematics The ...
in the highlands.


Taxonomy and systematics

The
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Todus'' was introduced by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...
in 1760 with the
Jamaican tody The Jamaican tody (''Todus todus'') is a species of bird in the genus '' Todus'' endemic to Jamaica. Local names for the Jamaican tody include ''rasta bird'', ''robin'' and ''robin redbreast''. Taxonomy The Jamaican tody was formally describe ...
(''Todus todus'') as the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
. ''Todus'' is a Latin word for a small bird mentioned by the Roman playwright
Plautus Titus Maccius Plautus ( ; 254 – 184 BC) was a Roman playwright of the Old Latin period. His comedies are the earliest Latin literary works to have survived in their entirety. He wrote Palliata comoedia, the genre devised by Livius Andro ...
and the grammarian
Sextus Pompeius Festus Sextus Pompeius Festus, usually known simply as Festus, was a Ancient Rome, Roman Grammarian (Greco-Roman), grammarian who probably flourished in the later 2nd century AD, perhaps at Narbo (Narbonne) in Gaul. Work He made a 20-volume epitome of V ...
. This name had earlier been used for the Jamaican tody by the Irish physician Patrick Browne in his book ''The Civil and Natural History of Jamaica'' that was published in 1756.


Extant species

Five
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
are recognized:


Former species

* Emperor fairywren (as ''Todus cyanocephalus'') *
Leaden flycatcher The leaden flycatcher (''Myiagra rubecula'') is a species of passerine bird in the family Monarchidae. Around 15 cm (6 in) in length, the male is lustrous azure with white underparts, while the female possesses leaden head, mantle and ...
(as ''Todus rubecula'')


Description

Todies range in weight from 5 to 7 g and in length from 10 to 11.5 cm. They have colourful
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
and resemble kingfishers in their general shape. They have green heads, backs and wings, red throats (absent in immature Puerto Rican, broad-billed, and narrow-billed Todies) with a white and blue-grey stripe on each side, and yellow undertail coverts; the colour of the rest of the undersides is pale and varies according to species. The irises are pale grey. They have long, flattened bills (as do many flycatching birds) with serrated edges; the upper mandible is black and the lower is red with a little black. The legs, and especially the feet, are small. Todies are highly
vocal The human voice consists of sound made by a human being using the vocal tract, including talking, singing, laughing, crying, screaming, shouting, humming or yelling. The human voice frequency is specifically a part of human sound producti ...
, except that the Jamaican tody seldom calls in the non-breeding season (August to November); they give simple, unmusical buzzing notes, beeps, and guttural rattles, puffing their throats out with every call. Their wings produce a "strange, whirring rattle", though mostly when courting or defending territory in the Puerto Rican tody.


Behaviour and ecology

Todies are generally sedentary; the longest single flight known for the broad-billed tody is 40 m. Their activity is greatest in the morning when sunny weather follows rain, and in March and September.


Breeding

Like most of the
Coraciiformes The Coraciiformes are a group of usually colourful birds including the kingfishers, the bee-eaters, the rollers, the motmots, and the todies. They generally have syndactyly, with three forward-pointing toes (and toes 3 & 4 fused at their b ...
, todies nest in tunnels, which they dig with their beaks and feet in steep banks or rotten tree trunks. The tunnel is 30 cm long in the Cuban and narrow-billed Todies, 30 to 60 cm in the broad-billed tody, and ends in a nest chamber, generally not reused. They lay about four round white eggs in the chamber. Both parents incubate but are surprisingly inattentive to the eggs. The young are
altricial Precocial species in birds and mammals are those in which the young are relatively mature and mobile from the moment of birth or hatching. They are normally nidifugous, meaning that they leave the nest shortly after birth or hatching. Altricial ...
and stay in the nest until they can fly. Both parents also care for the nestlings, much more attentively; they may feed each chick up to 140 times per day, the highest rate known among birds.


Food and feeding

Todies eat small prey such as insects and lizards. Insects, particularly grasshoppers, crickets, beetles, bugs, butterflies, bees, wasps, and ants, form the greater part of the diet. Spiders and millipedes may also be taken, as is a small amount of fruit (2% of the diet). Todies typically sit on a low, small branch, singly or in pairs, keeping still or stepping or hopping sideways. When they see prey moving on the lower surface of a leaf, they fly a short distance (averaging 2.2 m in the broad-billed tody and 1.0 m in the
Puerto Rican tody The Puerto Rican tody (''Todus mexicanus''), locally known in Puerto Rican Spanish, Spanish as ''San Pedrito'' ("little Saint Peter"), is a bird Endemism in birds, endemic to the Geography of Puerto Rico, main island of Puerto Rico. In 2022, the ...
), diagonally upward to glean it. They may also take prey from the ground, occasionally chasing it with a few hops.


References


External links


Tody videos
on the Internet Bird Collection {{Taxonbar, from=Q726375 Birds of the Greater Antilles Birds of the Lesser Antilles * Bird genera