''Titanocetus'' (" Titano whale") is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of extinct

cetacea Cetacea (; , ) is an infraorder of aquatic mammals belonging to the order Artiodactyla that includes whales, dolphins and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively c ...

ns closely related to the family

Cetotheriidae Cetotheriidae is a family of baleen whales (parvorder Mysticeti). The family is known to have existed from the Late Oligocene to the Early Pleistocene before going extinct. Although some phylogenetic studies conducted by recovered the living py ...

Discovery

The fossil remains of ''Titanocetus'' were discovered within some marine deposits dating back to the

Serravallian The Serravallian is, in the geologic timescale, an List of time periods, age or a stage (stratigraphy), stage in the middle Miocene epoch (geology), Epoch/series (stratigraphy), Series, which spans the time between 13.82 annum, Ma and 11.63 Ma (m ...

(middle

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

) and belonging to the Fumaiolo Formation (

Republic of San Marino A republic, based on the Latin phrase ''res publica'' ('public affair' or 'people's affair'), is a state in which political power rests with the public (people), typically through their representatives—in contrast to a monarchy. Although a ...

). The whale was then described in 1900 by the Italian

paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...

Giovanni Capellini, who later (1901) named it ''Aulocetus sammarinensis''. Over a century later, in 2006, the paleontologist Michelangelo Bisconti stated that the remains was too different from the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

of the genus '' Aulocetus'' and thus established a new genus, ''Titanocetus''.

Description

This whale was similar in appearance to the living

Balaenopteridae Rorquals () are the largest clade, group of baleen whales, comprising the family (biology), family Balaenopteridae, which contains nine extant taxon, extant species in two genus, genera. They include the largest known animal that has ever lived, ...

, although it was considerably smaller: the

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

barely exceeded one meter in total length, while the entire animal reached around six meters. Considered nowadays to be a primitive member of the

Mysticeti Baleen whales (), also known as whalebone whales, are marine mammals of the parvorder Mysticeti in the infraorder Cetacea (whales, dolphins and porpoises), which use baleen plates (or "whalebone") in their mouths to sieve plankton from the wate ...

, already equipped with

baleen Baleen is a filter feeder, filter-feeding system inside the mouths of baleen whales. To use baleen, the whale first opens its mouth underwater to take in water. The whale then pushes the water out, and animals such as krill are filtered by th ...

, ''Titanocetus'' was a carrier of both modern (i.e. the

rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit ** podium * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ...

, wide and flat) and ancestral characters (i.e. the

squamosal The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestra ...

and

parietal bone The parietal bones ( ) are two bones in the skull which, when joined at a fibrous joint known as a cranial suture, form the sides and roof of the neurocranium. In humans, each bone is roughly quadrilateral in form, and has two surfaces, four bord ...

s, which occupy part of the

temporal fenestra Temporal fenestrae are openings in the temporal region of the skull of some amniotes, behind the orbit (eye socket). These openings have historically been used to track the evolution and affinities of reptiles. Temporal fenestrae are commonly (al ...

References

Bibliography

*Capellini, G. 1900. "Balenottera miocenica della Repubblica di San Marino". ''Atti della Reale Accademia dei Lincei'' 5:233-235. *Capellini, G. 1901. "Balenottera miocenica del Monte Titano Repubblica di S. Marino". ''Memorie della Regia Accademia delle Scienze all'Istituto di Bologna'' 5:237-260. * {{Taxonbar, from=Q15653244 Baleen whales Miocene cetaceans Prehistoric mammals of Europe Monotypic prehistoric placental genera Fossil taxa described in 2006