Timeline Of Zoology
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This is a chronologically organized listing of notable
zoological Zoology ( , ) is the scientific study of animals. Its studies include the anatomy, structure, embryology, Biological classification, classification, Ethology, habits, and distribution of all animals, both living and extinction, extinct, and ...
events and discoveries.


Ancient world

*28000 BC.
Cave painting In archaeology, cave paintings are a type of parietal art (which category also includes petroglyphs, or engravings), found on the wall or ceilings of caves. The term usually implies prehistoric art, prehistoric origin. These paintings were often c ...
s (e.g.
Chauvet Cave The Chauvet-Pont-d'Arc Cave ( ) in the Ardèche department of southeastern France is a cave that contains some of the best-preserved figurative cave paintings in the world, as well as other evidence of Upper Paleolithic life.Clottes (2003b), p. ...
) in
Southern France Southern France, also known as the south of France or colloquially in French as , is a geographical area consisting of the regions of France that border the Atlantic Ocean south of the Marais Poitevin,Louis Papy, ''Le midi atlantique'', Atlas e ...
and northern
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
depict animals such as
mammoths A mammoth is any species of the extinct elephantid genus ''Mammuthus.'' They lived from the late Miocene epoch (from around 6.2 million years ago) into the Holocene until about 4,000 years ago, with mammoth species at various times inhabi ...
in a stylized fashion. *12000–8000 BC. Bubalus Period creation of rock art in the Central Sahara depicting a range of animals including
elephants Elephants are the Largest and heaviest animals, largest living land animals. Three living species are currently recognised: the African bush elephant (''Loxodonta africana''), the African forest elephant (''L. cyclotis''), and the Asian ele ...
,
antelopes The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do no ...
,
rhinoceros A rhinoceros ( ; ; ; : rhinoceros or rhinoceroses), commonly abbreviated to rhino, is a member of any of the five extant taxon, extant species (or numerous extinct species) of odd-toed ungulates (perissodactyls) in the family (biology), famil ...
and
catfish Catfish (or catfishes; order (biology), order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Catfish are common name, named for their prominent barbel (anatomy), barbels, which resemble a cat's whiskers, though not ...
. *10000 BC. Humans (''
Homo sapiens Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
'')
domesticated Domestication is a multi-generational mutualistic relationship in which an animal species, such as humans or leafcutter ants, takes over control and care of another species, such as sheep or fungi, to obtain from them a steady supply of reso ...
dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers. ...
s,
pig The pig (''Sus domesticus''), also called swine (: swine) or hog, is an omnivorous, domesticated, even-toed, hoofed mammal. It is named the domestic pig when distinguishing it from other members of the genus '' Sus''. Some authorities cons ...
s,
sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...
,
goat The goat or domestic goat (''Capra hircus'') is a species of Caprinae, goat-antelope that is mostly kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the ...
s,
fowl Fowl are birds belonging to one of two biological orders, namely the gamefowl or landfowl ( Galliformes) and the waterfowl ( Anseriformes). Anatomical and molecular similarities suggest these two groups are close evolutionary relatives; toget ...
, and other animals in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
,
northern Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...
and the
Near East The Near East () is a transcontinental region around the Eastern Mediterranean encompassing the historical Fertile Crescent, the Levant, Anatolia, Egypt, Mesopotamia, and coastal areas of the Arabian Peninsula. The term was invented in the 20th ...
. *6500 BC. The
aurochs The aurochs (''Bos primigenius''; or ; pl.: aurochs or aurochsen) is an extinct species of Bovini, bovine, considered to be the wild ancestor of modern domestic cattle. With a shoulder height of up to in bulls and in cows, it was one of t ...
, ancestors of domestic
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
, were domesticated in the next two centuries if not earlier (Obre I,
Yugoslavia , common_name = Yugoslavia , life_span = 1918–19921941–1945: World War II in Yugoslavia#Axis invasion and dismemberment of Yugoslavia, Axis occupation , p1 = Kingdom of SerbiaSerbia , flag_p ...
). This was the last major animal to be tamed as a source of milk, meat, power, and leather in the
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
. *3500 BC.
Sumer Sumer () is the earliest known civilization, located in the historical region of southern Mesopotamia (now south-central Iraq), emerging during the Chalcolithic and Early Bronze Age, early Bronze Ages between the sixth and fifth millennium BC. ...
ian animal-drawn wheeled vehicles and plows were developed in
Mesopotamia Mesopotamia is a historical region of West Asia situated within the Tigris–Euphrates river system, in the northern part of the Fertile Crescent. Today, Mesopotamia is known as present-day Iraq and forms the eastern geographic boundary of ...
, the region called the "
Fertile Crescent The Fertile Crescent () is a crescent-shaped region in the Middle East, spanning modern-day Iraq, Israel, Jordan, Lebanon, Palestine, and Syria, together with northern Kuwait, south-eastern Turkey, and western Iran. Some authors also include ...
."
Irrigation Irrigation (also referred to as watering of plants) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has bee ...
was probably done using animal power. Since Sumer had no natural defenses, armies with mounted
cavalry Historically, cavalry (from the French word ''cavalerie'', itself derived from ''cheval'' meaning "horse") are groups of soldiers or warriors who Horses in warfare, fight mounted on horseback. Until the 20th century, cavalry were the most mob ...
and
chariot A chariot is a type of vehicle similar to a cart, driven by a charioteer, usually using horses to provide rapid Propulsion, motive power. The oldest known chariots have been found in burials of the Sintashta culture in modern-day Chelyabinsk O ...
s became important which increased the importance of
equines ''Equus'' () is a genus of mammals in the perissodactyl family (biology), family Equidae, which includes wild horse, horses, Asinus, asses, and zebras. Within the Equidae, ''Equus'' is the only recognized Extant taxon, extant genus, comprising s ...
. *2000 BC. Domestication of the
silkworm ''Bombyx mori'', commonly known as the domestic silk moth, is a moth species belonging to the family Bombycidae. It is the closest relative of '' Bombyx mandarina'', the wild silk moth. Silkworms are the larvae of silk moths. The silkworm is of ...
in
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. *1100 BC. Won Chang (China), first of the Zhou emperors, stocked his imperial
zoological garden A zoo (short for zoological garden; also called an animal park or menagerie) is a facility where animals are kept within enclosures for public exhibition and often bred for Conservation biology, conservation purposes. The term ''zoological garden ...
with
deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...
,
goat The goat or domestic goat (''Capra hircus'') is a species of Caprinae, goat-antelope that is mostly kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the ...
s,
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s, and
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
from many parts of the world. The emperor also enjoyed sporting events with the use of animals. *850 BC.
Homer Homer (; , ; possibly born ) was an Ancient Greece, Ancient Greek poet who is credited as the author of the ''Iliad'' and the ''Odyssey'', two epic poems that are foundational works of ancient Greek literature. Despite doubts about his autho ...
(
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
) wrote the epics
Iliad The ''Iliad'' (; , ; ) is one of two major Ancient Greek epic poems attributed to Homer. It is one of the oldest extant works of literature still widely read by modern audiences. As with the ''Odyssey'', the poem is divided into 24 books and ...
and
Odyssey The ''Odyssey'' (; ) is one of two major epics of ancient Greek literature attributed to Homer. It is one of the oldest surviving works of literature and remains popular with modern audiences. Like the ''Iliad'', the ''Odyssey'' is divi ...
, both of them containing some correct observations on bees and fly maggots, while using animals as
monster A monster is a type of imaginary or fictional creature found in literature, folklore, mythology, fiction and religion. They are very often depicted as dangerous and aggressive, with a strange or grotesque appearance that causes Anxiety, terror ...
s and
metaphor A metaphor is a figure of speech that, for rhetorical effect, directly refers to one thing by mentioning another. It may provide, or obscure, clarity or identify hidden similarities between two different ideas. Metaphors are usually meant to cr ...
s (gross soldiers turned into
pig The pig (''Sus domesticus''), also called swine (: swine) or hog, is an omnivorous, domesticated, even-toed, hoofed mammal. It is named the domestic pig when distinguishing it from other members of the genus '' Sus''. Some authorities cons ...
s by the witch
Circe In Greek mythology, Circe (; ) is an enchantress, sometimes considered a goddess or a nymph. In most accounts, Circe is described as the daughter of the sun god Helios and the Oceanid Perse (mythology), Perse. Circe was renowned for her vast kn ...
). Both epics refer to
mule The mule is a domestic equine hybrid between a donkey, and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two ...
s. *610 BC.
Anaximander Anaximander ( ; ''Anaximandros''; ) was a Pre-Socratic philosophy, pre-Socratic Ancient Greek philosophy, Greek philosopher who lived in Miletus,"Anaximander" in ''Chambers's Encyclopædia''. London: George Newnes Ltd, George Newnes, 1961, Vol. ...
(Greek, 610–545 BC) was a student of
Thales of Miletus Thales of Miletus ( ; ; ) was an Ancient Greek pre-Socratic philosopher from Miletus in Ionia, Asia Minor. Thales was one of the Seven Sages, founding figures of Ancient Greece. Beginning in eighteenth-century historiography, many came to ...
. He was taught that the first life was formed by
spontaneous generation Spontaneous generation is a superseded scientific theory that held that living creatures could arise from non-living matter and that such processes were commonplace and regular. It was hypothesized that certain forms, such as fleas, could ...
in the mud. Later animals came into being by transmutations, left the water, and reached dry land. Man was derived from lower animals, probably aquatic. His writings, especially his poem ''On Nature'', were read and cited by
Aristotle Aristotle (; 384–322 BC) was an Ancient Greek philosophy, Ancient Greek philosopher and polymath. His writings cover a broad range of subjects spanning the natural sciences, philosophy, linguistics, economics, politics, psychology, a ...
and other later philosophers, but are lost now. *563? BC.
Buddha Siddhartha Gautama, most commonly referred to as the Buddha (),* * * was a wandering ascetic and religious teacher who lived in South Asia during the 6th or 5th century BCE and founded Buddhism. According to Buddhist legends, he was ...
(Indian, 563?–483 BC) had gentle ideas on the treatment of animals. He said that animals are held to have intrinsic worth, not just the values they derive from their usefulness to man. *500 BC.
Empedocles Empedocles (; ; , 444–443 BC) was a Ancient Greece, Greek pre-Socratic philosopher and a native citizen of Akragas, a Greek city in Sicily. Empedocles' philosophy is known best for originating the Cosmogony, cosmogonic theory of the four cla ...
of Agrigentum (Greek, 504–433 BC) reportedly rid a town of
malaria Malaria is a Mosquito-borne disease, mosquito-borne infectious disease that affects vertebrates and ''Anopheles'' mosquitoes. Human malaria causes Signs and symptoms, symptoms that typically include fever, Fatigue (medical), fatigue, vomitin ...
by draining nearby swamps. He proposed the theory of the
four humors Humorism, the humoral theory, or humoralism, was a system of medicine detailing a supposed makeup and workings of the human body, adopted by Ancient Greek and Roman physicians and philosophers. Humorism began to fall out of favor in the 17th ce ...
and a natural origin of living things. *500 BC.
Xenophanes Xenophanes of Colophon ( ; ; – c. 478 BC) was a Greek philosopher, theologian, poet, and critic of Homer. He was born in Ionia and travelled throughout the Greek-speaking world in early classical antiquity. As a poet, Xenophanes was known f ...
(Greek, 576–460 BC), a disciple of
Pythagoras Pythagoras of Samos (;  BC) was an ancient Ionian Greek philosopher, polymath, and the eponymous founder of Pythagoreanism. His political and religious teachings were well known in Magna Graecia and influenced the philosophies of P ...
(570–497 BC), first recognized
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s as animal remains and inferred that their presence on mountains indicated the latter had once been beneath the sea. "If horses or oxen had hands and could draw or make statues, horses would represent the forms of gods as horses, oxen as oxen."
Galen Aelius Galenus or Claudius Galenus (; September 129 – AD), often Anglicization, anglicized as Galen () or Galen of Pergamon, was a Ancient Rome, Roman and Greeks, Greek physician, surgeon, and Philosophy, philosopher. Considered to be one o ...
(130–216 AD) revived interest in fossils that had been rejected by Aristotle, and the speculations of Xenophanes were again viewed with favor. *470 BC.
Democritus Democritus (, ; , ''Dēmókritos'', meaning "chosen of the people"; – ) was an Ancient Greece, Ancient Greek Pre-Socratic philosophy, pre-Socratic philosopher from Abdera, Thrace, Abdera, primarily remembered today for his formulation of an ...
of Abdera (Greek, 470–370 BC) made dissections of many animals and humans. He was the first Greek philosopher-scientist to propose a classification of animals, dividing them into blooded animals (Vertebrata) and bloodless animals (Evertebrata). He also held that lower animals had perfected organs and that the brain was the seat of thought. *460 BC.
Hippocrates Hippocrates of Kos (; ; ), also known as Hippocrates II, was a Greek physician and philosopher of the Classical Greece, classical period who is considered one of the most outstanding figures in the history of medicine. He is traditionally referr ...
(Greek, 460–370 BC), the "Father of Medicine," used animal dissections to advance human anatomy. *440 BC.
Herodotus Herodotus (; BC) was a Greek historian and geographer from the Greek city of Halicarnassus (now Bodrum, Turkey), under Persian control in the 5th century BC, and a later citizen of Thurii in modern Calabria, Italy. He wrote the '' Histori ...
of
Halikarnassos Halicarnassus ( ; Latin: ''Halicarnassus'' or ''Halicarnāsus''; ''Halikarnāssós''; ; Carian: 𐊠𐊣𐊫𐊰 𐊴𐊠𐊥𐊵𐊫𐊰 ''alos k̂arnos'') was an ancient Greek city in Caria, in Anatolia.Nile The Nile (also known as the Nile River or River Nile) is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa. It has historically been considered the List of river sy ...
, but much of
ancient Egyptian civilization Ancient Egypt () was a cradle of civilization concentrated along the lower reaches of the Nile River in Northeast Africa. It emerged from prehistoric Egypt around 3150BC (according to conventional Egyptian chronology), when Upper and Lower ...
had already lost to living memory by his time. *427 BC.
Plato Plato ( ; Greek language, Greek: , ; born  BC, died 348/347 BC) was an ancient Greek philosopher of the Classical Greece, Classical period who is considered a foundational thinker in Western philosophy and an innovator of the writte ...
(Greek, 427–347 BC) held that animals existed to serve man, but they should not be mistreated because this would lead people to mistreat others. Others who echoed this opinion are St.
Thomas Aquinas Thomas Aquinas ( ; ; – 7 March 1274) was an Italian Dominican Order, Dominican friar and Catholic priest, priest, the foremost Scholasticism, Scholastic thinker, as well as one of the most influential philosophers and theologians in the W ...
,
Immanuel Kant Immanuel Kant (born Emanuel Kant; 22 April 1724 – 12 February 1804) was a German Philosophy, philosopher and one of the central Age of Enlightenment, Enlightenment thinkers. Born in Königsberg, Kant's comprehensive and systematic works ...
, and
Albert Schweitzer Ludwig Philipp Albert Schweitzer (; 14 January 1875 – 4 September 1965) was a German and French polymath from Alsace. He was a theologian, organist, musicologist, writer, humanitarian, philosopher, and physician. As a Lutheran minister, ...
. *384 BC.
Aristotle Aristotle (; 384–322 BC) was an Ancient Greek philosophy, Ancient Greek philosopher and polymath. His writings cover a broad range of subjects spanning the natural sciences, philosophy, linguistics, economics, politics, psychology, a ...
's (Greek, 384–322 BC) books '' Historia Animalium'' (9 books), ', and ' set the zoological stage for centuries. He emphasized the value of direct observation, recognized law and order in biological phenomena, and derived conclusions inductively from observations. He believed that there was a natural process of animals that ran from simple to complex. He made advances in
marine biology Marine biology is the scientific study of the biology of marine life, organisms that inhabit the sea. Given that in biology many scientific classification, phyla, family (biology), families and genera have some species that live in the sea and ...
, basing his writings on keen observation and rational interpretation as well as conversations with local
Lesbos Lesbos or Lesvos ( ) is a Greek island located in the northeastern Aegean Sea. It has an area of , with approximately of coastline, making it the third largest island in Greece and the List of islands in the Mediterranean#By area, eighth largest ...
fishermen for two years, beginning in 344 BC. His account of male protection of eggs by the barking catfish was scorned for centuries until
Louis Agassiz Jean Louis Rodolphe Agassiz ( ; ) FRS (For) FRSE (May 28, 1807 – December 14, 1873) was a Swiss-born American biologist and geologist who is recognized as a scholar of Earth's natural history. Spending his early life in Switzerland, he recei ...
confirmed Aristotle's description. *323 BC.
Alexander the Great Alexander III of Macedon (; 20/21 July 356 BC – 10/11 June 323 BC), most commonly known as Alexander the Great, was a king of the Ancient Greece, ancient Greek kingdom of Macedonia (ancient kingdom), Macedon. He succeeded his father Philip ...
(Macedonian, 356–323 BC) collected animals when he was not busy conquering the known world. He is credited with the introduction of the
peacock Peafowl is a common name for two bird species of the genus '' Pavo'' and one species of the closely related genus '' Afropavo'' within the tribe Pavonini of the family Phasianidae (the pheasants and their allies). Male peafowl are referred t ...
into Europe. *70 BC. Publius Vergilius Maro (
Virgil Publius Vergilius Maro (; 15 October 70 BC21 September 19 BC), usually called Virgil or Vergil ( ) in English, was an ancient Rome, ancient Roman poet of the Augustan literature (ancient Rome), Augustan period. He composed three of the most fa ...
) (70–19 BC) was a famous Roman poet. His poems ''Bucolics'' (42–37 BC) and ''Georgics'' (37–30 BC) hold much information on animal husbandry and farm life. His ''Aeneid'' (published posthumously) has many references to the zoology of his time. *36 BC. Marcus Terentius
Varro Marcus Terentius Varro (116–27 BCE) was a Roman polymath and a prolific author. He is regarded as ancient Rome's greatest scholar, and was described by Petrarch as "the third great light of Rome" (after Virgil and Cicero). He is sometimes call ...
(116–27 BC) wrote ', a treatise that includes
apiculture Beekeeping (or apiculture, from ) is the maintenance of bee colonies, commonly in artificial beehives. Honey bees in the genus ''Apis (bee), Apis'' are the most commonly kept species but other honey producing bees such as ''Melipona'' stingless be ...
. He also treated the problem of sterility in the
mule The mule is a domestic equine hybrid between a donkey, and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two ...
and recorded a rare instance in which a fertile mule was bred. *50.
Lucius Annaeus Seneca Lucius Annaeus Seneca the Younger ( ; AD 65), usually known mononymously as Seneca, was a Stoicism, Stoic philosopher of Ancient Rome, a statesman, a dramatist, and in one work, a satirist, from the post-Augustan age of Latin literature. Seneca ...
(Roman, 4 BC–65 AD), tutor to Roman emperor
Nero Nero Claudius Caesar Augustus Germanicus ( ; born Lucius Domitius Ahenobarbus; 15 December AD 37 – 9 June AD 68) was a Roman emperor and the final emperor of the Julio-Claudian dynasty, reigning from AD 54 until his ...
, maintained that animals have no reason, just instinct, a "stoic" position. *77.
Pliny the Elder Gaius Plinius Secundus (AD 23/24 79), known in English as Pliny the Elder ( ), was a Roman Empire, Roman author, Natural history, naturalist, and naval and army commander of the early Roman Empire, and a friend of the Roman emperor, emperor Vesp ...
(Roman, 23–79) wrote his ' in 37 volumes. This work is a catch-all of zoological folklore, superstitions, and some good observations. *79.
Pliny the Younger Gaius Plinius Caecilius Secundus (born Gaius Caecilius or Gaius Caecilius Cilo; 61 – ), better known in English as Pliny the Younger ( ), was a lawyer, author, and magistrate of Ancient Rome. Pliny's uncle, Pliny the Elder, helped raise and e ...
(Roman, 62–113), nephew of
Pliny the Elder Gaius Plinius Secundus (AD 23/24 79), known in English as Pliny the Elder ( ), was a Roman Empire, Roman author, Natural history, naturalist, and naval and army commander of the early Roman Empire, and a friend of the Roman emperor, emperor Vesp ...
, inherited his uncle's notes and wrote on beekeeping. *100.
Plutarch Plutarch (; , ''Ploútarchos'', ; – 120s) was a Greek Middle Platonist philosopher, historian, biographer, essayist, and priest at the Temple of Apollo (Delphi), Temple of Apollo in Delphi. He is known primarily for his ''Parallel Lives'', ...
(Roman, 46–120) stated that animals' behavior is motivated by reason and understanding. *131.
Galen Aelius Galenus or Claudius Galenus (; September 129 – AD), often Anglicization, anglicized as Galen () or Galen of Pergamon, was a Ancient Rome, Roman and Greeks, Greek physician, surgeon, and Philosophy, philosopher. Considered to be one o ...
of Pergamum (Greek, 130–216), physician to Roman Emperor
Marcus Aurelius Marcus Aurelius Antoninus ( ; ; 26 April 121 – 17 March 180) was Roman emperor from 161 to 180 and a Stoicism, Stoic philosopher. He was a member of the Nerva–Antonine dynasty, the last of the rulers later known as the Five Good Emperors ...
, wrote on human anatomy from dissections of animals. His texts were used for hundreds of years, gaining the reputation of infallibility. *200 c. Various compilers in post-classical and medieval times added to the ' (or, more popularly, the ''Bestiary''), the major book on animals for hundreds of years. Animals were believed to exist to serve man, if not as food or slaves then as moral examples. *Early third century. Composition of ''De Natura Animalium'' by
Claudius Aelianus Claudius Aelianus (; ), commonly Aelian (), born at Praeneste, was a Roman author and teacher of rhetoric who flourished under Septimius Severus and probably outlived Elagabalus, who died in 222. He spoke Greek so fluently that he was called "h ...
(Roman, 175–235).


Middle Ages

*600. Isidorus Hispalensis (Spanish bishop of
Seville Seville ( ; , ) is the capital and largest city of the Spain, Spanish autonomous communities of Spain, autonomous community of Andalusia and the province of Seville. It is situated on the lower reaches of the Guadalquivir, River Guadalquivir, ...
) (560–636) wrote ', an encyclopedic compendium of ancient knowledge including information on animals that served until the rediscovery of Aristotle and Pliny. Full of errors, it nevertheless was influential for hundreds of years. He also wrote '. *781.
Al-Jahiz Abu Uthman Amr ibn Bahr al-Kinani al-Basri (; ), commonly known as al-Jahiz (), was an Arab polymath and author of works of literature (including theory and criticism), theology, zoology, philosophy, grammar, dialectics, rhetoric, philology, lin ...
(Afro-Arab, 781–868/869), a scholar at
Basra Basra () is a port city in Iraq, southern Iraq. It is the capital of the eponymous Basra Governorate, as well as the List of largest cities of Iraq, third largest city in Iraq overall, behind Baghdad and Mosul. Located near the Iran–Iraq bor ...
, wrote on the influence of environment on animals. *901.
Horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
s came into wider use in those parts of Europe where the
three-field system The three-field system is a regime of crop rotation in which a field is planted with one set of crops one year, a different set in the second year, and left fallow in the third year. A set of crops is ''rotated'' from one field to another. The tech ...
produces grain surpluses for feed, but hay-fed oxen were more economical, if less efficient, in terms of time and labor and remained almost the sole source of animal power in southern Europe, where most farmers continued to use the two-field system. *1225–1244.
Thomas of Cantimpré Thomas of Cantimpré (Latin: Thomas Cantimpratensis or Thomas Cantipratensis) (Sint-Pieters-Leeuw, 1201 – Louvain, 15 May 1272) was a Flemish Region, Flemish Catholic medieval writer, preacher, theologian and a friar belonging to the Dominican ...
‚ (Fleming, 1204?–1275?) wrote ', a major 13th-century encyclopedia. *1244–1248. Frederick II von Hohenstaufen (Holy Roman Emperor) (1194–1250) wrote ' (''The Art of Hunting with Birds'') as a practical guide to
ornithology Ornithology, from Ancient Greek ὄρνις (''órnis''), meaning "bird", and -logy from λόγος (''lógos''), meaning "study", is a branch of zoology dedicated to the study of birds. Several aspects of ornithology differ from related discip ...
. *1244. Vincentius Bellovacensis ( Vincent of Beauvais) (1190–1264) wrote ' (1244–1254), a major encyclopedia. This work comprises three huge volumes, of 80 books and 9,885 chapters. *1254–1323.
Marco Polo Marco Polo (; ; ; 8 January 1324) was a Republic of Venice, Venetian merchant, explorer and writer who travelled through Asia along the Silk Road between 1271 and 1295. His travels are recorded in ''The Travels of Marco Polo'' (also known a ...
(Venetian, 1254–1323) provided information on Asiatic fauna, revealing new animals to Europeans. "
Unicorn The unicorn is a legendary creature that has been described since Classical antiquity, antiquity as a beast with a single large, pointed, spiraling horn (anatomy), horn projecting from its forehead. In European literature and art, the unico ...
s" (which may actually have been rhinos) were reported from southern China, but fantastic animals were otherwise not included. *1255–1270.
Albertus Magnus Albertus Magnus ( 1200 – 15 November 1280), also known as Saint Albert the Great, Albert of Swabia, Albert von Bollstadt, or Albert of Cologne, was a German Dominican friar, philosopher, scientist, and bishop, considered one of the great ...
of
Cologne Cologne ( ; ; ) is the largest city of the States of Germany, German state of North Rhine-Westphalia and the List of cities in Germany by population, fourth-most populous city of Germany with nearly 1.1 million inhabitants in the city pr ...
(Bavarian, 1206?–1280) (Albert von Bollstaedt or St. Albert) wrote '. He promoted Aristotle but also included new material on the perfection and intelligence of animals, especially bees. *1304–1309. Petrus de Crescentii wrote ', a practical manual for agriculture with many accurate observations on
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s and other animals.
Apiculture Beekeeping (or apiculture, from ) is the maintenance of bee colonies, commonly in artificial beehives. Honey bees in the genus ''Apis (bee), Apis'' are the most commonly kept species but other honey producing bees such as ''Melipona'' stingless be ...
was discussed at length. *1492.
Christopher Columbus Christopher Columbus (; between 25 August and 31 October 1451 – 20 May 1506) was an Italians, Italian explorer and navigator from the Republic of Genoa who completed Voyages of Christopher Columbus, four Spanish-based voyages across the At ...
(Italian) arrives in the
New World The term "New World" is used to describe the majority of lands of Earth's Western Hemisphere, particularly the Americas, and sometimes Oceania."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: ...
. New animals soon begin to overload European zoology. Columbus is said to have introduced cattle, horses, and eight pigs from the
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
to
Hispaniola Hispaniola (, also ) is an island between Geography of Cuba, Cuba and Geography of Puerto Rico, Puerto Rico in the Greater Antilles of the Caribbean. Hispaniola is the most populous island in the West Indies, and the second-largest by List of C ...
in 1493, giving rise to virtual devastation of that and other islands. Pigs were often set ashore by sailors to provide food on the ship's later return. Feral populations of hogs were often dangerous to humans. *1519–1520. Bernal Diaz del Castillo (Spanish, 1450?–1500), chronicler of Cortez's conquest of
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, commented on the zoological gardens of
Aztec The Aztecs ( ) were a Mesoamerican civilization that flourished in central Mexico in the Post-Classic stage, post-classic period from 1300 to 1521. The Aztec people included different Indigenous peoples of Mexico, ethnic groups of central ...
ruler Montezuma (1466–1520), a marvel with
parrot Parrots (Psittaciformes), also known as psittacines (), are birds with a strong curved beak, upright stance, and clawed feet. They are classified in four families that contain roughly 410 species in 101 genus (biology), genera, found mostly in ...
s,
rattlesnake Rattlesnakes are venomous snakes that form the genus, genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting sm ...
s, and other animals. *1523.
Gonzalo Fernández de Oviedo y Valdés Gonzalo Fernández de Oviedo y Valdés (August 1478 – 1557), commonly known as Oviedo, was a Spanish soldier, historian, writer, botanist and colonist. Oviedo participated in the Spanish colonization of the West Indies, arriving in the first fe ...
(Spanish, 1478–1557), appointed official historiographer of the
Indies The East Indies (or simply the Indies) is a term used in historical narratives of the Age of Discovery. The ''Indies'' broadly referred to various lands in the East or the Eastern Hemisphere, particularly the islands and mainlands found i ...
in 1523, wrote ''Sumario de la Natural Historia delas Indias'' (Toledo, 1527). He was the first to describe many New World animals, such as the
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a Suidae, pig, with a short, prehensile nose trunk (proboscis). Tapirs inhabit jungle and forest regions of South America, South and Centr ...
,
opossum Opossums () are members of the marsupial order Didelphimorphia () endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 126 species in 18 genera. Opossums originated in South America and entered North A ...
,
manatee Manatees (, family (biology), family Trichechidae, genus ''Trichechus'') are large, fully aquatic, mostly herbivory, herbivorous marine mammals sometimes known as sea cows. There are three accepted living species of Trichechidae, representing t ...
,
iguana ''Iguana'' (, ) is a genus of herbivorous lizards that are native to tropical areas of Mexico, Central America, South America, and the Caribbean. The genus was first described by Austrian naturalist Josephus Nicolaus Laurenti, J.N. Laurenti in ...
,
armadillo Armadillos () are New World placental mammals in the order (biology), order Cingulata. They form part of the superorder Xenarthra, along with the anteaters and sloths. 21 extant species of armadillo have been described, some of which are dis ...
,
anteater Anteaters are the four extant mammal species in the suborder Vermilingua (meaning "worm tongue"), commonly known for eating ants and termites. The individual species have other names in English and other languages. Together with sloths, they ar ...
s,
sloth Sloths are a Neotropical realm, Neotropical group of xenarthran mammals constituting the suborder Folivora, including the extant Arboreal locomotion, arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of move ...
,
pelican Pelicans (genus ''Pelecanus'') are a genus of large water birds that make up the family Pelecanidae. They are characterized by a long beak and a large throat pouch used for catching prey and draining water from the scooped-up contents before ...
, and
hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
s.


Modern world

*1551–1555. Pierre Belon (French, 1517–1564) wrote ' (1551) and ' (1555). This latter work included 110 animal species and offered many new observations and corrections to Herodotus. ' (1555) was his picture book, with improved animal classification and accurate anatomical drawings. In this he published a man's and a bird's skeleton side by side to show the resemblance. He discovered an armadillo shell in a market in
Syria Syria, officially the Syrian Arab Republic, is a country in West Asia located in the Eastern Mediterranean and the Levant. It borders the Mediterranean Sea to the west, Turkey to Syria–Turkey border, the north, Iraq to Iraq–Syria border, t ...
, showing how Muslims were distributing the finds from the New World. *1551.
Conrad Gessner Conrad Gessner (; ; 26 March 1516 – 13 December 1565) was a Swiss physician, naturalist, bibliographer, and philologist. Born into a poor family in Zürich, Switzerland, his father and teachers quickly realised his talents and supported him t ...
(Swiss, 1516–1565) wrote ' (Tiguri, 4 vols., 1551–1558, last volume published in 1587) and gained renown. This work, although uncritically compiled in places, was consulted for over 200 years. He also wrote ' (1553) and ' (1563). *1552 Edward Wotton (English, 1492–1555) published ', a work that influenced Gessner. *1554–1555. Guillaume Rondelet (French, 1507–1566) wrote ' (1554) and ' (1555). He gathered
vernacular Vernacular is the ordinary, informal, spoken language, spoken form of language, particularly when perceptual dialectology, perceived as having lower social status or less Prestige (sociolinguistics), prestige than standard language, which is mor ...
names in hope of being able to identify the animal in question. He did go to print with discoveries that disagreed with Aristotle. *1578. Jean de Lery (French, 1534–1611) was a member of the French colony at
Rio de Janeiro Rio de Janeiro, or simply Rio, is the capital of the Rio de Janeiro (state), state of Rio de Janeiro. It is the List of cities in Brazil by population, second-most-populous city in Brazil (after São Paulo) and the Largest cities in the America ...
. He published ' (1578) with observations on the local fauna. *1585.
Thomas Harriot Thomas Harriot (; – 2 July 1621), also spelled Harriott, Hariot or Heriot, was an English astronomer, mathematician, ethnographer and translator to whom the theory of refraction is attributed. Thomas Harriot was also recognized for his con ...
(English, 1560–1621) was a naturalist with the first attempted English colony in North America, on
Roanoke Island Roanoke Island () is an island in Dare County, bordered by the Outer Banks of North Carolina. It was named after the historical Roanoke, a Carolina Algonquian people who inhabited the area in the 16th century at the time of English colonizat ...
,
North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...
. His ''Brief and True Report of the New Found Land of Virginia'' (1590) describes the black bear, gray squirrel, hare,
otter Otters are carnivorous mammals in the subfamily Lutrinae. The 13 extant otter species are all semiaquatic, aquatic, or marine. Lutrinae is a branch of the Mustelidae family, which includes weasels, badgers, mink, and wolverines, among ...
,
opossum Opossums () are members of the marsupial order Didelphimorphia () endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 126 species in 18 genera. Opossums originated in South America and entered North A ...
,
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the North American, northern or common raccoon (also spelled racoon) to distinguish it from Procyonina, other species of raccoon, is a mammal native to North America. It is the largest ...
,
skunk Skunks are mammals in the family Mephitidae. They are known for their ability to spray a liquid with a strong, unpleasant scent from their anal glands. Different species of skunk vary in appearance from black-and-white to brown, cream or gi ...
,
Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...
and
mule deer The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer. Unlike the related whit ...
, turkeys, and
horseshoe crab Horseshoe crabs are arthropods of the family Limulidae and the only surviving xiphosurans. Despite their name, they are not true crabs or even crustaceans; they are chelicerates, more closely related to arachnids like spiders, ticks, and scor ...
(''Limulus''). *1589. José de Acosta (Spanish, 1539–1600) wrote ' (1589) and ' (1590), describing many animals from the New World previously unknown to Europeans.


17th century

*1600. In Italy a spider scare lead to hysteria and the
tarantella Tarantella () is a group of various Southern Italy, southern Italian Italian folk dance, folk dances originating in the regions of Calabria, Campania, Sicilia, and Apulia. It is characterized by a fast Beat (music), upbeat tempo, usually in Ti ...
dance by which the body cures itself through physical exertions. *1602.
Ulisse Aldrovandi Ulisse Aldrovandi (11 September 1522 – 4 May 1605) was an Italian naturalist, the moving force behind Bologna's botanical garden, one of the first in Europe. Carl Linnaeus and the comte de Buffon reckoned him the father of natural history stud ...
(Italian, 1522–1605) wrote '. This and his other works include many scientific inaccuracies, but he used wing and leg morphology to construct his classification of insects. He is more highly regarded for his ornithological contributions. *1604–1614.
Francisco Hernández de Toledo Francisco Hernández de Toledo (c. 1515 – 28 January 1587) was a naturalist and court physician to Philip II of Spain. He was among the first wave of Spanish Renaissance physicians practicing according to the revived principles formulated by Hipp ...
(Spanish) was sent to study Mexican biota in 1593–1600, by
Philip II of Spain Philip II (21 May 152713 September 1598), sometimes known in Spain as Philip the Prudent (), was King of Spain from 1556, King of Portugal from 1580, and King of Naples and List of Sicilian monarchs, Sicily from 1554 until his death in 1598. He ...
. His notes were published in Mexico in 1604 and 1614, describing many animals to Europeans for the first time:
coyote The coyote (''Canis latrans''), also known as the American jackal, prairie wolf, or brush wolf, is a species of canis, canine native to North America. It is smaller than its close relative, the Wolf, gray wolf, and slightly smaller than the c ...
, buffalo,
axolotl The axolotl (; from ) (''Ambystoma mexicanum'') is a neoteny, paedomorphic salamander, one that Sexual maturity, matures without undergoing metamorphosis into the terrestrial adult form; adults remain Aquatic animal, fully aquatic with obvio ...
,
porcupine Porcupines are large rodents with coats of sharp Spine (zoology), spines, or quills, that protect them against predation. The term covers two Family (biology), families of animals: the Old World porcupines of the family Hystricidae, and the New ...
, pronghorn antelope,
horned lizard ''Phrynosoma'', whose members are known as the horned lizards, horny toads, or horntoads, is a genus of North American lizards and the type genus of the Family (biology), family Phrynosomatidae. Their common names refer directly to their horns or ...
,
bison A bison (: bison) is a large bovine in the genus ''Bison'' (from Greek, meaning 'wild ox') within the tribe Bovini. Two extant taxon, extant and numerous extinction, extinct species are recognised. Of the two surviving species, the American ...
,
peccary Peccaries (also javelinas or skunk pigs) are pig-like ungulates of the family Tayassuidae (New World pigs). They are found throughout Central and South America, Trinidad in the Caribbean, and in the southwestern area of North America. Peccari ...
, and the
toucan Toucans (, ) are Neotropical birds in the family Ramphastidae. They are most closely related to the Semnornis, Toucan barbets. They are brightly marked and have large, often colorful Beak, bills. The family includes five genus, genera and over ...
. He also included figures of many animals for the first time, including the
ocelot The ocelot (''Leopardus pardalis'') is a medium-sized spotted Felidae, wild cat that reaches at the shoulders and weighs between on average. It is native to the southwestern United States, Mexico, Central America, Central and South America, ...
, rattlesnake, manatee, alligator, armadillo, and pelican. *1607 (1612?).
Captain John Smith John Smith ( – 21 June 1631) was an English soldier, explorer, colonial governor, admiral of New England, and author. He was knighted for his services to Sigismund Báthory, Prince of Transylvania, and his friend Mózes Székely. Followin ...
(English), head of the Jamestown colony, wrote ''A Map of Virginia'' in which he describes the physical features of the country, its climate, plants and animals, and inhabitants. He describes the raccoon,
muskrat The muskrat or common muskrat (''Ondatra zibethicus'') is a medium-sized semiaquatic rodent native to North America and an introduced species in parts of Europe, Asia, and South America. The muskrat is found in wetlands over various climates ...
,
flying squirrel Flying squirrels (scientifically known as Pteromyini or Petauristini) are a tribe (biology), tribe of 50 species of squirrels in the family (biology), family Squirrel, Sciuridae. Despite their name, they are not in fact capable of full flight i ...
, and other animals. *1617. Garcilaso de la Vega (Peruvian Spanish, 1539–1617) wrote ''Royal Commentaries of Peru'', containing descriptions of the
condor Condor is the common name for two species of New World vultures, each in a monotypic genus. The name derives from the Quechua language, Quechua ''kuntur''. They are the largest flying land birds in the Western Hemisphere. One species, the And ...
,
ocelot The ocelot (''Leopardus pardalis'') is a medium-sized spotted Felidae, wild cat that reaches at the shoulders and weighs between on average. It is native to the southwestern United States, Mexico, Central America, Central and South America, ...
s, puma,
viscacha Viscacha or vizcacha (, ) are rodents of two genera ('' Lagidium'' and '' Lagostomus'') in the family Chinchillidae. They are native to South America and convergently resemble rabbits. The five extant species of viscacha are: *The Plains vi ...
,
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a Suidae, pig, with a short, prehensile nose trunk (proboscis). Tapirs inhabit jungle and forest regions of South America, South and Centr ...
, rhea,
skunk Skunks are mammals in the family Mephitidae. They are known for their ability to spray a liquid with a strong, unpleasant scent from their anal glands. Different species of skunk vary in appearance from black-and-white to brown, cream or gi ...
,
llama The llama (; or ) (''Lama glama'') is a domesticated South American camelid, widely used as a List of meat animals, meat and pack animal by Inca empire, Andean cultures since the pre-Columbian era. Llamas are social animals and live with ...
, huanaco,
paca A paca ia a rodent in South and Central America. Paca or PACA may also refer to: People * William Paca (1740–1799), a Founding Father of the United States * Paca Blanco (Francisca Blanco Díaz, born 1949), Spanish activist * Paca Navas (Franc ...
, and
vicuña The vicuña (''Lama vicugna'') or vicuna (both , very rarely spelled ''vicugna'', Vicugna, its former genus name) is one of the two wild South American camelids, which live in the high alpine tundra, alpine areas of the Andes; the other cameli ...
. *1620? North American colonists probably introduced the European honeybee, ''
Apis mellifera The western honey bee or European honey bee (''Apis mellifera'') is the most common of the 7–12 species of honey bees worldwide. The genus name ''Apis'' is Latin for 'bee', and ''mellifera'' is the Latin for 'honey-bearing' or 'honey-carrying', ...
'', into Virginia. By the 1640s these insects were also in
Massachusetts Massachusetts ( ; ), officially the Commonwealth of Massachusetts, is a U.S. state, state in the New England region of the Northeastern United States. It borders the Atlantic Ocean and the Gulf of Maine to its east, Connecticut and Rhode ...
. They became feral and advanced through eastern North America before the settlers. *1628.
William Harvey William Harvey (1 April 1578 – 3 June 1657) was an English physician who made influential contributions to anatomy and physiology. He was the first known physician to describe completely, and in detail, pulmonary and systemic circulation ...
(English, 1578–1657) published ' (1628) with the doctrine of the circulation of blood (an inference made by him in about 1616). *1634. William Wood (English) wrote ''New England Prospect'' (1634) in which he describes New England's fauna. *1637. Thomas Morton (English, c. 1579–1647) wrote ''New English Canaan'' (1637) with treatments of 26 species of mammals, 32 birds, 20 fishes and 8 marine invertebrates. *1648. Georg Marcgrave (1610–1644) was a German astronomer working for Johann Moritz, Count Maurice of Nassau, in the Dutch colony set up in northeastern
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
. His ' (1648) contains the best early descriptions of many Brazilian animals. Marcgrave used Tupi names that were later Latinized by Linnaeus in the 13th edition of the ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
''. The biological and linguistic data could have come from Moraes, a Brazilian
Jesuit The Society of Jesus (; abbreviation: S.J. or SJ), also known as the Jesuit Order or the Jesuits ( ; ), is a religious order (Catholic), religious order of clerics regular of pontifical right for men in the Catholic Church headquartered in Rom ...
priest turned apostate. *1651.
William Harvey William Harvey (1 April 1578 – 3 June 1657) was an English physician who made influential contributions to anatomy and physiology. He was the first known physician to describe completely, and in detail, pulmonary and systemic circulation ...
published ' (1651) with the aphorism ' on the title page. *1661.
Marcello Malpighi Marcello Malpighi (10 March 1628 – 30 November 1694) was an Italians, Italian biologist and physician, who is referred to as the "founder of microscopical anatomy, histology and father of physiology and embryology". Malpighi's name is borne by ...
(Italian, 1628–1694) discovered
capillaries A capillary is a small blood vessel, from 5 to 10 micrometres in diameter, and is part of the microcirculation system. Capillaries are microvessels and the smallest blood vessels in the body. They are composed of only the tunica intima (the in ...
(1661), structures predicted to exist by Harvey some thirty years earlier. Malpighi was the founder of
microanatomy Histology, also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures visi ...
. He studied, among other things, the anatomy of the silkworm (1669) and the development of the chick (1672). *1665.
Robert Hooke Robert Hooke (; 18 July 16353 March 1703) was an English polymath who was active as a physicist ("natural philosopher"), astronomer, geologist, meteorologist, and architect. He is credited as one of the first scientists to investigate living ...
(English, 1635–1703) wrote ''Micrographia'' (1665, 88 plates), with his early microscopic studies. He coined the term " cell." *1668. Opening of the Royal Menagerie at
Palace of Versailles The Palace of Versailles ( ; ) is a former royal residence commissioned by King Louis XIV located in Versailles, Yvelines, Versailles, about west of Paris, in the Yvelines, Yvelines Department of Île-de-France, Île-de-France region in Franc ...
. *1668.
Francesco Redi Francesco Redi (18 February 1626 – 1 March 1697) was an Italians, Italian physician, naturalist, biologist, and poet. He is referred to as the "founder of experimental biology", and as the "father of modern parasitology". He was the first perso ...
(Italian, 1621–1697) wrote ' (1668) and ' (1708). His refutation of spontaneous generation in flies is still considered a model in experimentation. *1669.
Jan Swammerdam Jan or Johannes Swammerdam (February 12, 1637 – February 17, 1680) was a Dutch biologist and microscopist. His work on insects demonstrated that the various phases during the life of an insect—Egg (biology), egg, larva, pupa, and adult ...
(Dutch, 1637–1680) wrote ' (1669) describing
metamorphosis Metamorphosis is a biological process by which an animal physically develops including birth transformation or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and different ...
in insects and supporting the performation doctrine. He was a pioneer in microscopic studies. He gave the first description of red blood corpuscles and discovered the valves of
lymph vessels The lymphatic vessels (or lymph vessels or lymphatics) are thin-walled vessels (tubes), structured like blood vessels, that carry lymph. As part of the lymphatic system, lymph vessels are complementary to the cardiovascular system. Lymph vessel ...
. His work was unknown and unacknowledged until after his death. *1672.
Regnier de Graaf Regnier de Graaf (English spelling), original Dutch spelling Reinier de Graaf, or Latinized Reijnerus de Graeff (30 July 164117 August 1673), was a Dutch physician, physiologist and anatomy, anatomist who made key discoveries in reproductive bio ...
(Dutch, 1641–1673) reported that he had traced the human egg from the
ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
down the
fallopian tube The fallopian tubes, also known as uterine tubes, oviducts or salpinges (: salpinx), are paired tubular sex organs in the human female body that stretch from the Ovary, ovaries to the uterus. The fallopian tubes are part of the female reproduct ...
to the
uterus The uterus (from Latin ''uterus'', : uteri or uteruses) or womb () is the hollow organ, organ in the reproductive system of most female mammals, including humans, that accommodates the embryonic development, embryonic and prenatal development, f ...
. What he really saw was the follicle. *1675–1722.
Antonie van Leeuwenhoek Antonie Philips van Leeuwenhoek ( ; ; 24 October 1632 – 26 August 1723) was a Dutch microbiologist and microscopist in the Golden Age of Dutch art, science and technology. A largely self-taught man in science, he is commonly known as " ...
(Dutch, 1632–1723) wrote ', a treatise with early observations made with microscopes. He discovered blood corpuscles, striated muscles, human spermatozoa (1677),
protozoa Protozoa (: protozoan or protozoon; alternative plural: protozoans) are a polyphyletic group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic debris. Historically ...
(1674),
bacteria Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr ...
(1683),
rotifer The rotifers (, from Latin 'wheel' and 'bearing'), sometimes called wheel animals or wheel animalcules, make up a phylum (Rotifera ) of microscopic and near-microscopic Coelom#Pseudocoelomates, pseudocoelomate animals. They were first describ ...
s, etc. *
Martin Lister Martin Lister (12 April 1639 – 2 February 1712) was an English natural history, naturalist and physician. His daughters Anne Lister (illustrator), Anne and Susanna Lister, Susanna were two of his illustrators and engravers. J. D. Woodley, 'L ...
(English, 1639–1712) publishes the first work on spiders based on observation. *1691.
John Ray John Ray Fellow of the Royal Society, FRS (November 29, 1627 – January 17, 1705) was a Christian England, English Natural history, naturalist widely regarded as one of the earliest of the English parson-naturalists. Until 1670, he wrote his ...
(English, 1627–1705) wrote ' (1693), ' (1710), and ''The Wisdom of God Manifested in the Works of the Creation'' (1691). He tried to classify different animal species into groups largely according to their
toe Toes are the digits of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being ''digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being ''plantigrade''; ...
s and
teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
. *1699. Edward Tyson (English, 1650–1708) wrote ' (or ''Anatomy of a Pygmie Compared with that of a Monkey, an Ape and a Man'') (1699), his anatomical study of the
primate Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...
. This was the first detailed and accurate study of the higher apes. Other studies by Tyson include the female
porpoise Porpoises () are small Oceanic dolphin, dolphin-like cetaceans classified under the family Phocoenidae. Although similar in appearance to dolphins, they are more closely related to narwhals and Beluga whale, belugas than to the Oceanic dolphi ...
, male
rattlesnake Rattlesnakes are venomous snakes that form the genus, genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting sm ...
,
tapeworm Eucestoda, commonly referred to as tapeworms, is the larger of the two subclasses of flatworms in the class Cestoda (the other subclass being Cestodaria). Larvae have six posterior hooks on the scolex (head), in contrast to the ten-hooked Ce ...
, roundworm (''Ascaris''),
peccary Peccaries (also javelinas or skunk pigs) are pig-like ungulates of the family Tayassuidae (New World pigs). They are found throughout Central and South America, Trinidad in the Caribbean, and in the southwestern area of North America. Peccari ...
and
opossum Opossums () are members of the marsupial order Didelphimorphia () endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 126 species in 18 genera. Opossums originated in South America and entered North A ...
. *1700. Félix de Azara (Spanish) estimated the feral herds of cattle on the South American
pampas The Pampas (; from Quechua 'plain'), also known as the Pampas Plain, are fertile South American low grasslands that cover more than and include the Argentine provinces of Buenos Aires, La Pampa, Santa Fe, Entre Ríos, and Córdoba; all o ...
at 48 million animals. These animals probably descended from herds introduced by the Jesuits some 100 years earlier. (North America and Australia were to follow in this pattern, where feral herds of cattle and
mustang The mustang is a free-roaming horse of the Western United States, descended from horses brought to the Americas by the Spanish conquistadors. Mustangs are often referred to as wild horses, but because they are descended from once-domesticate ...
s would explode, become pests, and reform the frontier areas.)


18th century

*1705.
Maria Sybilla Merian Maria Sibylla Merian (2 April 164713 January 1717) was a German Entomology, entomologist, naturalist and scientific illustrator. She was one of the earliest European naturalists to document observations about insects directly. Merian was a desce ...
(German, 1647–1717) wrote and illustrated her ' (') (1705). In this book she stated that '' Fulgora lanternaria'' was luminous. *1734–1742. René Antoine Ferchault de Réaumur (French, 1683–1756) was an early entomologist. His ' (6 volumes) shows the best of zoological observation at the time. He invented the glass-fronted bee hive. *1740. Abraham Trembley, Swiss naturalist, discovered the hydra which he considered to combine both animal and plant characteristics. His ' (1744) showed that freshwater polyps of ''Hydra'' could be sectioned or mutilated and still reform. Regeneration soon became a topic of inquiry among Réaumur, Bonnet,
Spallanzani Lazzaro Spallanzani (; 12 January 1729 – 11 February 1799) was an Italian Catholic priest (for which he was nicknamed Abbé Spallanzani), biologist and physiologist who made important contributions to the experimental study of bodily function ...
, and others. *1745. Charles Bonnet (French-Swiss, 1720–1793) wrote ' (1745) and ' (1732). He confirmed
parthenogenesis Parthenogenesis (; from the Greek + ) is a natural form of asexual reproduction in which the embryo develops directly from an egg without need for fertilization. In animals, parthenogenesis means the development of an embryo from an unfertiliz ...
of
aphid Aphids are small sap-sucking insects in the Taxonomic rank, family Aphididae. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white Eriosomatinae, woolly ...
s. *1745. Pierre Louis M. de Maupertuis (French, 1698–1759) went to Lapland to measure the arc of the meridian (1736–1737). Maupertuis was a Newtonian. He generated family trees for inheritable characteristics (e.g.,
haemophilia Haemophilia (British English), or hemophilia (American English) (), is a mostly inherited genetic disorder that impairs the body's ability to make blood clots, a process needed to stop bleeding. This results in people bleeding for a long ...
in European royal families) and showed inheritance through both the male and female lines. He was an early evolutionist and head of the Berlin Academy of Sciences. In 1744 he proposed the theory that molecules from all parts of the body were gathered into the
gonad A gonad, sex gland, or reproductive gland is a Heterocrine gland, mixed gland and sex organ that produces the gametes and sex hormones of an organism. Female reproductive cells are egg cells, and male reproductive cells are sperm. The male gon ...
s (later called "
pangenesis Pangenesis was Charles Darwin's hypothetical mechanism for heredity, in which he proposed that each part of the body continually emitted its own type of small organic particles called gemmules that aggregated in the gonads, contributing heritabl ...
"). ' was published anonymously in 1745. Maupertuis wrote ' in which he suggests a
survival of the fittest "Survival of the fittest" is a phrase that originated from Darwinian evolutionary theory as a way of describing the mechanism of natural selection. The biological concept of fitness is defined as reproductive success. In Darwinian terms, th ...
concept: "Could not one say that since, in the accidental combination of Nature's productions, only those could survive which found themselves provided with certain appropriate relationships, it is no wonder that these relationships are present in all the species that actually exist? These species which we see today are only the smallest part of those which a blind destiny produced." *1748. John Tuberville Needham, an English naturalist, wrote ''Observations upon the Generation, Composition, and Decomposition of Animal and Vegetable Substances'' in which he offers "proof" of spontaneous generation. Needham found flasks of broth teeming with "little animals" after having boiled them and sealed them, but his experimental techniques were faulty. *1749–1804.
Georges-Louis Leclerc, Comte de Buffon Georges-Louis Leclerc, Comte de Buffon (; 7 September 1707 – 16 April 1788) was a French Natural history, naturalist, mathematician, and cosmology, cosmologist. He held the position of ''intendant'' (director) at the ''Jardin du Roi'', now ca ...
(French, 1707–1788) wrote ' (1749–1804 in 44 vols.), which asserted that species were mutable. Buffon also drew attention to vestigial organs. He held that spermatozoa were "living organic molecules" that multiplied in the semen. *1752. Founding of the Schönbrunn Zoo in
Vienna Vienna ( ; ; ) is the capital city, capital, List of largest cities in Austria, most populous city, and one of Federal states of Austria, nine federal states of Austria. It is Austria's primate city, with just over two million inhabitants. ...
, the world's oldest continuously operating zoo. *1753. The
British Museum The British Museum is a Museum, public museum dedicated to human history, art and culture located in the Bloomsbury area of London. Its permanent collection of eight million works is the largest in the world. It documents the story of human cu ...
was founded in the will of Sir
Hans Sloane Sir Hans Sloane, 1st Baronet, (16 April 1660 – 11 January 1753), was an Irish physician, naturalist, and collector. He had a collection of 71,000 items which he bequeathed to the British nation, thus providing the foundation of the British ...
(English (born Ireland), 1660–1753). It would open its doors in 1759. *1758.
Albrecht von Haller Albrecht von Haller (also known as Albertus de Haller; 16 October 170812 December 1777) was a Swiss anatomist, physiologist, naturalist, encyclopedist, bibliographer and poet. A pupil of Herman Boerhaave and Jacob Winslow, he is sometimes r ...
(Swiss, 1708–1777) was one of the founders of modern
physiology Physiology (; ) is the science, scientific study of function (biology), functions and mechanism (biology), mechanisms in a life, living system. As a branches of science, subdiscipline of biology, physiology focuses on how organisms, organ syst ...
. His work on the
nervous system In biology, the nervous system is the complex system, highly complex part of an animal that coordinates its behavior, actions and sense, sensory information by transmitting action potential, signals to and from different parts of its body. Th ...
was revolutionary. He championed animal physiology, along with human physiology. See his textbook ' (1758). *1758.
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
(Swedish, 1707–1778) published the ' whose tenth edition (1758) is the starting point of
binomial nomenclature In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
for zoology. *1759.
Caspar Friedrich Wolff Caspar Friedrich Wolff (18 January 1733 – 22 February 1794) was a German physiologist and embryologist who is widely regarded as one of the pioneers of modern embryology. Life Wolff was born in Berlin, Brandenburg. In 1759 he graduated as an ...
(1733–1794) wrote ' (1759) that disagreed with the idea of
preformation In the history of biology, preformationism (or preformism) is a formerly popular theory that organisms develop from miniature versions of themselves. Instead of assembly from parts, preformationists believed that the form of living things exis ...
. He supported the doctrine of epigenesis as a way to resolve the problem of hybrids (mule, hinny, apemen) in preformation. *1769. Edward Bancroft (English) wrote ''An Essay on the Natural History of Guyana in South America'' (1769) and advanced the theory that flies transmit disease. *1771.
Johann Reinhold Forster Johann Reinhold Forster (; 22 October 1729 – 9 December 1798) was a German Reformed pastor and naturalist. Born in Tczew, Dirschau, Pomeranian Voivodeship (1466–1772), Pomeranian Voivodeship, Polish–Lithuanian Commonwealth (now Tczew, Po ...
(German, 1729–1798) was the naturalist on Cook's second voyage around the world (1772–1775). He published a ''Catalogue of the Animals of North America'' (1771) as an addendum to Kalm's ''Travels''. He also studied the birds of Hudson Bay. *1774. Gilbert White (English) wrote ''The Natural History of Selborne, The natural history and antiquities of Selborne, in the county of Southampton'' (1774) with fine ornithological observations on Animal migration, migration, Territory (animal), territoriality, and Flocking (behavior), flocking. *1774. European lancelet is described. *1775. Johan Christian Fabricius (Danish, 1745–1808) wrote ' (1775), ' (1776), ' (1778), ' (1792–1794, in six vols.), and later publications (to 1805), to make Fabricius one of the world's greatest entomologists. *1780. Lazaro Spallanzani (Italian, 1729–1799) performed In vitro fertilisation, artificial fertilization in the frog, silkmoth and dog. He concluded from filtration experiments that spermatozoa were necessary for fertilization. In 1783 he showed that human digestion was a chemical process since Gastric acid, gastric juices in and outside the body liquefied food (meat). He used himself as the experimental animal. His work to disprove spontaneous generation in microbes was resisted by John Needham (English priest, 1713–1781). *1780. Antoine Lavoisier (French, 1743–1794) and Pierre Laplace (French, 1749–1827) wrote ''Memoir on heat''. Lavoisier, the discoverer of oxygen, concluded that animal Respiration (physiology), respiration was a form of combustion. *1783–1792. Alexandre Rodrigues Ferreira (Brazilian) wrote '. His specimens were taken by Étienne Geoffroy Saint-Hilaire, Saint-Hilaire from Lisbon to the National Museum of Natural History, France, Paris Museum during the Napoleonic invasion of Portugal. *1784. Johann Wolfgang von Goethe (German) wrote ' (1795) that promoted the idea of archetypes to which animals should be compared. *1784. Thomas Jefferson (American) wrote ''Notes on the State of Virginia'' (1784) that refuted some of Buffon's mistakes about New World fauna. As U.S. president, he dispatched the Lewis and Clark Expedition to the American West (1804). *1788. The First Fleet inaugurates British settlement of Australia. Knowledge of Australia's unique zoology, including marsupials and the platypus, would revolutionize Western zoology. *1789? Guillaume Antoine Olivier (French, 1756–1814) wrote ', or ' (1789). *1789. George Shaw (biologist), George Shaw & Frederick Polydore Nodder published ''The Naturalist's Miscellany: or coloured figures of natural objects drawn and described immediately from nature'' (1789–1813) in 24 volumes with hundreds of color plates. *1792. François Huber made original observations on honeybees. In his ' (1792) he noted that the first eggs laid by queen bees develop into Drone (bee), drones if her nuptial flight had been delayed and that her last eggs would also give rise to drones. He also noted that rare worker eggs develop into drones. This anticipated by over 50 years the discovery by Jan Dzierżon that drones come from unfertilized eggs and queen and worker bees come from fertilized eggs. *1793. The National Museum of Natural History, France is founded in Paris. It became a major center of zoological research in the early nineteenth century. *1793. Lazaro Spallanzani (Italian, 1729–1799) conducted experiments on the orientation of bats and owls in the dark. *1793. Christian Konrad Sprengel (1750–1816) wrote ' (1793) that was a major work on insect pollination of flowers, previously discovered in 1721 by Philip Miller (1694–1771), the head gardener at Chelsea and author of the famous ''Gardener's Dictionary'' (1731–1804). *1794. Erasmus Darwin (English, grandfather of Charles Darwin) wrote ''Zoönomia, or the Laws of Organic Life'' (1794) in which he advanced the idea that environmental influences could transform species. *1796–1829. Pierre André Latreille (French, 1762–1833) sought to provide a "natural" system for the classification of animals, in his many monographs on invertebrates. ' (1811) was devoted to insects collected by Alexander Humboldt, Humboldt and Aime Bonpland, Bonpland. *1797-1804. Publication of A History of British Birds by Thomas Bewick and Ralph Beilby in two volumes. *1798. Publication of Thomas Robert Malthus's An Essay on the Principle of Population, a book important to both Darwin and Wallace. *1799. George Shaw (biologist), George Shaw (English) provided the first description of the duck-billed platypus. Everard Home (1802) provided the first complete description. *1799–1803. Alexander von Humboldt (German, 1769–1859) and Aimé Bonpland, Aimé Jacques Alexandre Goujaud Bonpland (French) arrived in Venezuela in 1799. Humboldt's ''Personal Narrative of Travels to the Equinoctial Regions of America during the years 1799–1804'' and ''Kosmos'' were influential in his time. *1799. Georges Cuvier (French, 1769–1832) established comparative anatomy as a field. He also founded the science of paleontology. He wrote ' (1801–1805), ' (1816), ' (1812–1813). He believed in the fixity of species and the Genesis flood narrative, Biblical Flood. His early ' (1798) was influential, but it did not include Cuvier's major contributions to animal classification. *1799. American hunters killed the last
bison A bison (: bison) is a large bovine in the genus ''Bison'' (from Greek, meaning 'wild ox') within the tribe Bovini. Two extant taxon, extant and numerous extinction, extinct species are recognised. Of the two surviving species, the American ...
on the Eastern coast of the United States, in Pennsylvania.


19th Century

*1802. Jean-Baptiste de Lamarck (French, 1744–1829) wrote ' and ' (1809). He was an early evolutionist and organized invertebrate paleontology. While Lamarck's contributions to science include work in meteorology, botany, chemistry, geology, and paleontology, he is best known for his work in invertebrate zoology and his theoretical work on evolution. He published a seven-volume work, ' ("Natural history of animals without backbones;" 1815–1822). *1813–1818. William Charles Wells (Scottish-American, 1757–1817) was the first to recognise the principle of natural selection. He read a paper to the Royal Society in 1813 (but not published until 1818) which used the idea to explain differences between Race (human categorization), human races. The application was limited to the question of how different skin colours arose. *1815. William Kirby (entomologist), William Kirby and William Spence (entomologist), William Spence (English) wrote ''An Introduction to Entomology'' (first edition in 1815). This was the first modern entomology text. *1817. Publication of ''American Entomology'' by Thomas Say, the first work devoted to American insects. A greatly expanded three-volume edition would appear 1824–1828. Say was a systematic zoologist who moved to the utopian community at New Harmony, Indiana, in 1825. Most of his insect collections have been recovered. *1817. Georges Cuvier wrote ''Le Règne Animal'' (Paris). *1817–1820. Johann Baptist von Spix (German, 1781–1826) and Carl Friedrich Philipp von Martius (German) conducted Brazilian zoological and botanical explorations (1817–1820). See their ' (3 vols., 1823–1831). *1817. William Smith (geologist), William Smith, in his ''Strategraphical System of Organized Fossils'' (1817) showed that certain strata have characteristic series of fossils. *1819 William Lawrence (biologist), William Lawrence (English, 1783–1867) published a book of his lectures to the Royal College of Surgeons. The book contains a rejection of Lamarckism (soft inheritance), proto-evolutionary ideas about the origin of mankind, and a denial of the 'Jewish scriptures' (Old Testament). He was forced to suppress the book after the Lord Chancellor refused copyright and other powerful men made threatening remarks. *1819. Malayan tapir, a first species of
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a Suidae, pig, with a short, prehensile nose trunk (proboscis). Tapirs inhabit jungle and forest regions of South America, South and Centr ...
to be discovered, is described. *1824. Publication of the French physician Henri Dutrochet's ''Mémoires pour servir a l'histoire anatomique et physiologique des végétaux et des animaux'' setting forth a physiological theory of the cell. *1824. The Royal Society for the Prevention of Cruelty to Animals (RSPCA) is founded at London. *1824. Founding of the Zoological Journal, the first English-language journal of zoology. The last issue would appear in 1834. *1825. Gideon Mantell (English) wrote "Notice on the ''Iguanodon'', a newly discovered fossil reptile, from the sandstone of Tilgate Forest, in Sussex" (Phil. Trans. Roy, Soc. Lond., 115: 179–186), the first paper on dinosaurs. The name dinosaur was coined by anatomist Richard Owen. *1826. Founding of the Zoological Society of London. *1826–1839. John James Audubon (Haitian-born American, 1785–1851) wrote ''Birds of America'' (1826–1839), with North American bird portraits and studies. See also his posthumously published volume on North American mammals written with his sons and the naturalist John Bachman, ''The Viviparous Quadrupeds of North America'' (1845–1854) with 150 folio plates. *1827. Karl Ernst von Baer (Russian embryologist, 1792–1876) was the founder of comparative embryology. He demonstrated the existence of the mammalian ovum, and he proposed the germ-layer theory. His major works include ' (1827) and ' (1828; 1837). *1827. Description of Sagitta (arrowworm), Sagitta, a genus of chaetognaths. *1828. The Zoological Society of London opens its "zoo" to the public (later known as the London Zoo) for two days a week beginning April 27, 1828, with the first hippopotamus to be seen in Europe since the ancient Romans showed one at the Colosseum. This was the first modern zoo founded for scientific research and education. *1829. James Smithson (English, 1765–1829) donates seed money in his will for the founding of the Smithsonian Institution in Washington, D.C., Washington. *1830–1833. Sir Charles Lyell (English, 1797–1875) writes ''Principles of Geology'' and described the time required for evolution to work. Darwin took this book to sea on HMS Beagle. *1830. Étienne Geoffroy Saint-Hilaire (French, 1772–1844) writes ' (1830). *1830. Founding of the Journal of Zoology, then known as Proceedings of the Zoological Society of London. *1831. Founding of the Magasin de Zoologie, the first French-language zoological journal. *1831–1836. Charles Darwin (English, 1809–1882) and Captain Robert FitzRoy (English) depart for their voyage. Darwin's report is generally known as ''The Voyage of the Beagle''. *1832. Thomas Nuttall (1786–1859) writes ''A Manual of the Ornithology of the United States and Canada'' (1832) that becomes the standard text on the subject for most of the 19th century. *1835. William Swainson (English, 1789–1855) writes ''A Treatise on the Geography and Classification of Animals'' (1835), in which he uses ad hoc land bridges to explain animal distributions. He includes some second-hand observations on Old World army ants. *1835. Founding of the Archiv für Naturgeschichte, the premier German-language journal of natural history with an emphasis on zoology. It would be published until 1926. *1839. Theodor Schwann (German, 1810–1882) writes ' (1839). Schwann established the foundation for cell theory. *1839.
Louis Agassiz Jean Louis Rodolphe Agassiz ( ; ) FRS (For) FRSE (May 28, 1807 – December 14, 1873) was a Swiss-born American biologist and geologist who is recognized as a scholar of Earth's natural history. Spending his early life in Switzerland, he recei ...
(Swiss-American, 1807–1873), an expert on fossil fishes, founds the Museum of Comparative Zoology at Harvard University, and becomes Darwin's North American opposition. He was a popularizer of natural history. His ' (1842–1847) was a pioneering effort. *1840. Jan Evangelista Purkyně, a Czech physiologist in Wrocław proposes that the word "protoplasm" be applied to the formative material of young animal embryos. *1842. Baron Justus von Liebig writes ' in which he suggests that animal heat is produced by combustion, and founds the science of biochemistry. *1843. John James Audubon, age 58, ascends the Missouri River to Fort Union at the mouth of the Yellowstone to sketch wild animals. *1844. Berlin Zoo is founded. *1844. Robert Chambers (journalist), Robert Chambers (Scottish, 1802–1871) writes the ''Vestiges of the Natural History of Creation'' (1844) in which he includes early evolutionary considerations. This book, anonymously published, has a profound effect on Alfred Russel Wallace. *1845. Karl Theodor Ernst von Siebold, von Siebold recognizes Protozoa as single-celled animals. *1848. Josiah C. Nott (American), a physician from New Orleans, publishes his belief that mosquitoes transmit
malaria Malaria is a Mosquito-borne disease, mosquito-borne infectious disease that affects vertebrates and ''Anopheles'' mosquitoes. Human malaria causes Signs and symptoms, symptoms that typically include fever, Fatigue (medical), fatigue, vomitin ...
. *1848. Alfred Russel Wallace (British, 1823–1913) and Henry W. Bates (English, 1825–1892) arrive in the Amazon River, Amazon River valley in 1848. Bates stays until 1859, exploring the upper Amazon. Wallace remains in the Amazon until 1852, exploring the Rio Negro (Amazon), Rio Negro. Wallace writes ''A Narrative of Travels on the Amazon and Rio Negro'' (1853), and Bates writes ''The Naturalist on the River Amazons'' (1863). Later (1854–1862), Wallace travels to the Far East, as he reports in ''The Malay Archipelago'' (1869). *1849. Arnold Adolph Berthold demonstrates by castration and testicular transplant that the Testicle, testis produces a blood-borne substance promoting male Secondary sex characteristic, secondary sexual characteristics. *1850. Thomas Hardwicke (British naturalist) is the first European to discover the lesser panda (''Ailurus fulgens'') in northern India. *1855. Alfred Russel Wallace (English, 1823–1913) publishes ''On the law which has regulated the introduction of new species'' (Ann. Mag. Nat. Hist., September 1855), with evolutionary ideas that drew upon Wallace's experiences in the Amazon. *1857–1881. Henri Milne-Edwards (French, 1800–1885) introduces the idea of physiologic division of labor and writes a treatise on comparative anatomy and physiology (1857–1881). * 1859. Charles Darwin publishes ''On the Origin of Species'', explaining the mechanism of evolution by natural selection and founding the field of evolutionary biology. * 1859. Founding of Copenhagen Zoo. * 1859. Commonwealth of Pennsylvania grants a charter for the founding of the Philadelphia Zoo. However, due to delays caused by the American Civil War the zoo will not open until 1874. * 1864. Louis Pasteur disproves the spontaneous generation of cellular life. * 1864. Founding of the Moscow Zoo. * 1865. Gregor Mendel demonstrates in Pea, pea plants that inheritance follows Mendelian Inheritance, definite rules. The Principle of Segregation states that each organism has two genes per trait, which segregate when the organism makes eggs or sperm. The Principle of Independent Assortment states that each gene in a pair is distributed independently during the formation of eggs or sperm. Mendel's observations went largely unnoticed. * 1868. Founding of the Lincoln Park Zoo in Chicago, Chicago, Illinois. * 1869. Friedrich Miescher discovers nucleic acids in the cell nucleus, nuclei of cells. * 1872. Darwin publishes The Expression of the Emotions in Man and Animals. * 1873. Founding of the Cincinnati Zoo. * 1876. Oskar Hertwig and Hermann Fol independently describe (in sea urchin eggs) the entry of sperm into the egg and the subsequent fusion of the egg and sperm nuclei to form a single new nucleus. * 1876. Founding of the Société zoologique de France. * 1878. Founding of the Zoological Society of Japan. * 1881. Przewalski's horse is described. * 1885. Polypodium hydriforme, an unusual cnidarian, is described. * 1889. Founding of the National Zoological Park (United States) as part of the Smithsonian Institution in Washington, D.C. * 1889. The first species of marsupial mole is described. * 1892. Hans Driesch separates the individual cells of a 2-cell sea urchin embryo, and shows that each cell develops into a complete individual, thus disproving the theory of preformation, and demonstrating that each cell is "totipotent," containing all the hereditary information necessary to form an individual. * 1895. Founding of the International Commission on Zoological Nomenclature, the body which continues to govern zoological nomenclature. * 1899. Founding of the Bronx Zoo. * 1900. Three biologists, Hugo de Vries, Carl Correns, Erich von Tschermak, independently rediscover Mendel's paper on heredity. * 1900. Founding of the Unione Zoologica Italiana. * 1900. First species of Siboglinidae discovered.


20th century


1901–1950

* 1901. Okapi is described for science. * 1905. William Bateson coins the term "genetics" to describe the study of biological inheritance. * 1906. Founding of the Beijing Zoo. * 1907. Ivan Pavlov demonstrates Classical conditioning, conditioned responses with salivating
dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers. ...
s. * 1907. First species of proturans are described. * 1910. Founding of the Saint Louis Zoo. * 1914. Insect suborder Grylloblattodea is discovered. * 1916. Founding of the San Diego Zoo. * 1916. Founding of the Zoological Survey of India. * 1922. Aleksandr Oparin proposes that the Earth's early atmosphere contained methane, ammonia, hydrogen, and Water vapor, water vapour, and that these were the raw materials for the origin of life. * 1931. Founding of Prague Zoo. * 1932. Founding of the Journal of Animal Ecology. * 1932. Establishment of the Bureau of Animal Population at the University of Oxford. * 1934. Brookfield Zoo is founded in Brookfield, Illinois, a suburb of Chicago. * 1935. Konrad Lorenz describes the imprinting (psychology), imprinting behavior of young
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s. * 1936. Founding of the National Zoological Gardens of Sri Lanka. * 1937. In ''Genetics and the Origin of Species'', Theodosius Dobzhansky applies the chromosome theory and population genetics to natural populations in the first mature work of Neo-Darwinism, also called the Modern synthesis (20th century), modern synthesis, a term coined by Julian Huxley. * 1938. A living coelacanth is found off the coast of southern Africa. * 1940. Donald Griffin and Robert Galambos announce their discovery of Animal echolocation, echolocation by bats.


1951–2000

* 1952. American Developmental biology, developmental biologists Robert Briggs (scientist), Robert Briggs and Thomas Joseph King, Thomas King Cloning, clone the first vertebrate by transplanting nuclei from leopard frog embryos into enucleated eggs. * 1952. Monoplacophorans, so far considered extinct, are found to be still present today. * 1955. First species from crustacean class Cephalocarida is discovered. * 1956. First gnathostomulids, a group of animals currently ranked as a phylum, are described. * 1959. Founding of the National Zoological Park Delhi in New Delhi, India. * 1960. Jane Goodall begins her chimpanzee research. * 1961. Joan Oró discovers that concentrated solutions of ammonium cyanide in water can produce the nucleotide adenine, a discovery that opened the way for new theories on the origin of life. * 1963. Founding of the National Zoo of Malaysia in Selangor. * 1963. Premier of the popular American zoological documentary series Wild Kingdom on the NBC television network. 140 episodes would appear before the series ended in 1988. * 1964. The International Union for Conservation of Nature (IUCN) issues its first IUCN Red List of species threatened by extinction. * 1967. John Gurdon used nuclear transplantation to cloning, clone an African clawed frog; first cloning of a vertebrate using a nucleus from a fully Cellular differentiation, differentiated adult cell. * 1972. Stephen Jay Gould and Niles Eldredge propose "punctuated equilibrium," a theory which states that the fossil record is an accurate depiction of the pace of evolution, with long periods of "stasis" (little change) punctuated by brief periods of rapid change and species Divergent evolution, divergence. * 1973. Passage of the U. S. Endangered Species Act of 1973. * 1977. Riftia pachyptila, the giant tube worm, discovered. * 1980. Founding of PETA, People for the Ethical Treatment of Animals. * 1980. Lobatocerebridae, a group of simple, unsegmented annelids with relatively complex brain, are described. * 1981. The first extant members of remipedia, remipedes, a new class of crustaceans, are discovered. * 1983. A new class of crustaceans, Tantulocarida, is proposed. * 1983. First species of loriciferans are described. * 1985. The primatologist Dian Fossey is murdered by poachers. * 1986. Sea daisies, a group of unusual starfishes, are discovered. * 1990. American entomologist E. O. Wilson and German entomologist Bert Hölldobler publish The Ants. The next year it will win the Pulitzer Prize for non-fiction, the only zoology textbook ever to do so. * 1992. Saola, a new species of ungulate, is discovered in Vietnam. * 1995. Symbion pandora, the first known species from the phylum Cycliophora, is described by Reinhardt Kristensen and Peter Funch. * 1996. Dolly the sheep is the first adult mammal to be successfully cloned. * 1999. Indonesian coelacanth, the second species of coelacanth, is described. * 2000. Reinhardt Kristensen and Peter Funch describe Limnognathia maerski, the first known species from the animal phylum Micrognathozoa. * 2000. Kikiki huna, the smallest known flying
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
, is described.


21st century

* 2002. First species from
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
suborder Mantophasmatodea are described. * 2002. Founding of the Zoological Society of Nigeria. * 2003. Tiburonia granrojo, known also as Big Red Jellyfish, a deep-sea species of jellyfish, is described. * 2004. Osedax, known also as bone-eating worm, a deep-sea species of Siboglinidae that feeds on bones of whale carcasses, is described. * 2005. Mimic octopus is described. * 2005. Malo maxima, a very venomous species of cubozoan, is described. * 2006. Chondrocladia, Ping-pong Tree Sponge, an unusual species of carnivorous sponge, is described. * 2006. Wunderpus photogenicus, Wunderpus is described. * 2007. Malo kingi, a tiny, but very venomous species of cubozoan, is described. * 2011. Halicephalobus mephisto, a species of nematode living 3.6 km under the surface of Earth in gold mines in South Africa, is discovered. * 2011. Bonaire banded box jellyfish, a species of cubozoan, is described. * 2012. Chondrocladia lyra, an unusual species of carnivorous sponge, is described. * 2013. Edwardsiella andrillae, an unusual species of sea anemone that lives attached to sea ice, is described. * 2016. Scolopendra cataracta, the first known species of amphibious centipede, is described. * 2016. Four new species from subphylum Xenoturbellida are described. Only a single species from Xenoturbellida (Xenoturbella bocki) was known before. * 2017. Tapanuli orangutan is found to be a distinct species. * 2017. Mariana snailfish, a deep-sea species of fish, is described. * 2018. Hoilungia, Hoilungia hongkongensis is described, the first species of placozoan to be discovered since 1883. * 2019. Polyplacotoma mediterranea is described, the first species from the placozoan class Polyplacotomia. * 2021. Eumillipes persephone, a species of millipede with up to 1,306 legs and the first myriapod known to have more than 1,000 legs is discovered.


See also

* Timeline of biology and organic chemistry * List of megafauna discovered in modern times * International Institute for Species Exploration * Monterey Bay Aquarium Research Institute * Lazarus taxon


References


External links


Mc-Graw HillWonders of Nature in the Menagerie of Blauw Jan in Amsterdam, as observed by Jan Velten around 1700Exotic Animals in Eighteenth-Century BritainZoologica
Göttingen State and University Library {{DEFAULTSORT:Timeline Of Zoology Zoology timelines, History of zoology