Timeline Of Plesiosaur Research
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This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation,
taxonomic 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme of classes (a taxonomy) and the allocation ...
revisions, and cultural portrayals of
plesiosaurs The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...
, an order of
marine reptiles Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. Only about 100 of the 12,000 extant reptile species and subspecies are classed as marine reptiles, including marine ...
that flourished during the
Mesozoic Era The Mesozoic Era is the era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Periods. It is characterized by the dominance of archosaurian reptiles such as the dinosaurs, and of gymnosperms s ...
. The first scientifically documented plesiosaur fossils were discovered during the early
19th century The 19th century began on 1 January 1801 (represented by the Roman numerals MDCCCI), and ended on 31 December 1900 (MCM). It was the 9th century of the 2nd millennium. It was characterized by vast social upheaval. Slavery was Abolitionism, ...
by
Mary Anning Mary Anning (21 May 1799 – 9 March 1847) was an English fossil collector, fossil trade, dealer, and palaeontologist. She became known internationally for her discoveries in Jurassic marine fossil beds in the cliffs along the English Cha ...
. Plesiosaurs were actually discovered and described before
dinosaurs Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
. They were also among the first animals to be featured in artistic reconstructions of the ancient world, and therefore among the earliest prehistoric creatures to attract the attention of the lay public. Plesiosaurs were originally thought to be a kind of primitive transitional form between marine life and terrestrial
reptiles Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
. However, now plesiosaurs are recognized as highly derived marine reptiles descended from terrestrial ancestors. Early researchers thought that plesiosaurs laid
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
s like most reptiles. They commonly imagined plesiosaurs crawling up beaches and burying eggs like
turtles Turtles are reptiles of the order Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtle ...
. However, later opinion shifted towards the idea that plesiosaurs gave live birth and never went on dry land: the biggest genera may have been too heavy to go on land at all, but smaller genera could've been capable. Plesiosaur locomotion has been a source of continuous controversy among paleontologists. The earliest speculations on the subject during the 19th century saw plesiosaur swimming as analogous to the paddling of modern
sea turtles Sea turtles (superfamily Chelonioidea), sometimes called marine turtles, are reptiles of the order Testudines and of the suborder Cryptodira. The seven existing species of sea turtles are the flatback, green, hawksbill, leatherback, loggerh ...
. During the 1920s opinion shifted to the idea that plesiosaurs swam with a
rowing Rowing is the act of propelling a human-powered watercraft using the sweeping motions of oars to displace water and generate reactional propulsion. Rowing is functionally similar to paddling, but rowing requires oars to be mechanically a ...
motion. However, a paper published in 1975 that once more found support for sea turtle-like swimming in plesiosaurs. This conclusion reignited the controversy regarding plesiosaur locomotion through the late 20th century. In 2011,
F. Robin O'Keefe F is the sixth letter of the Latin alphabet. F may also refer to: Science and technology Mathematics * F or f, the number 15 in hexadecimal and higher positional systems * ''p'F'q'', the hypergeometric function * F-distribution, a conti ...
and Luis M. Chiappe concluded the debate on plesiosaur reproduction, reporting the discovery of a gravid female plesiosaur with a single large embryo preserved inside her.


Prescientific

Associated remains of plesiosaurs and animals like the diving
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
''
Hesperornis ''Hesperornis'' (meaning "western bird") is a genus of cormorant-like Ornithuran that spanned throughout the Campanian age, and possibly even up to the early Maastrichtian age, of the Late Cretaceous period. One of the lesser-known discoverie ...
'' or the
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
''
Pteranodon ''Pteranodon'' (; from and ) is a genus of pterosaur that included some of the largest known flying reptiles, with ''P. longiceps'' having a wingspan of over . They lived during the late Cretaceous geological period of North America in presen ...
'' may have inspired legends about conflict between Thunder Birds and Water Monsters told by the Native Americans of
Kansas Kansas ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the west. Kansas is named a ...
and
Nebraska Nebraska ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Ka ...
.


18th century

1719 *
William Stukeley William Stukeley (7 November 1687 – 3 March 1765) was an English antiquarian, physician and Anglican clergyman. A significant influence on the later development of archaeology, he pioneered the scholarly investigation of the prehistoric ...
described the first partial skeleton of a plesiosaur, brought to his attention by the great-grandfather of
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
, Robert Darwin of Elston.


19th century


1810s

1811 *
Mary Anning Mary Anning (21 May 1799 – 9 March 1847) was an English fossil collector, fossil trade, dealer, and palaeontologist. She became known internationally for her discoveries in Jurassic marine fossil beds in the cliffs along the English Cha ...
discovered some plesiosaur fossils in
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...
.


1820s

1821 *
Henry De la Beche Sir Henry Thomas De la Beche KCB, FRS (10 February 179613 April 1855) was an English geologist and palaeontologist, the first director of the Geological Survey of Great Britain, who helped pioneer early geological survey methods. He was the ...
and William Conybeare are the first to name a plesiosaurian species: ''
Plesiosaurus dolichodeirus ''Plesiosaurus'' (Greek: ' ('), near to + ' ('), lizard) is a genus of extinct, large marine sauropterygian reptile that lived during the Early Jurassic. It is known by nearly complete skeletons from the Lias Group, Lias of England. It is disting ...
''. 1822 * Parkinson coined the name ''
Plesiosaurus priscus ''Plesiosaurus'' (Greek: ' ('), near to + ' ('), lizard) is a genus of extinct, large marine sauropterygian reptile that lived during the Early Jurassic. It is known by nearly complete skeletons from the Lias of England. It is distinguishable by ...
'' for some of the remains used by de la Beche and Conybeare as the basis for ''Plesiosaurus''. This species is currently regarded as of dubious taxonomic value. 1823 ''December'' *
Mary Anning Mary Anning (21 May 1799 – 9 March 1847) was an English fossil collector, fossil trade, dealer, and palaeontologist. She became known internationally for her discoveries in Jurassic marine fossil beds in the cliffs along the English Cha ...
discovered a nearly complete ''Plesiosaurus'' skeleton near Lyme Regis. This specimen would later be catalogued as BMNH 22656. ''c. December'' * Around the same time as the discovery of BMNH 22656, another Plesiosaurus specimen was discovered at the same site. The specimen was donated to the Oxford University museum and is probably the specimen known today as OXFUM J.10304. 1824 * Conybeare described the new species name ''Plesiosaurus dolichodeirus'' for the Plesiosaurus discovered by Anning. As the first species name given to a distinctive and well preserved Plesiosaurus skeleton it has come to be regarded as both the type specimen of ''Plesiosaurus dolichodeirus'' specifically and of the genus Plesiosaurus as a whole. 1829 * Mary Anning collected the ''Plesiosaurus dolichodeirus'' specimen now known as BMNH R.1313.


1830s

1837 * Buckland in Conybeare described the new species Plesiosaurus macrocephalus. 1839 * De la Beche illustrated a work titled "''
Duria Antiquior ''Duria Antiquior, a more ancient Dorset'' is the first pictorial representation of a scene of prehistoric life based on evidence from fossil reconstructions, a genre now known as paleoart. The first version was a watercolour painted in 1830 b ...
''", meaning "A More Ancient
Dorset Dorset ( ; Archaism, archaically: Dorsetshire , ) is a Ceremonial counties of England, ceremonial county in South West England. It is bordered by Somerset to the north-west, Wiltshire to the north and the north-east, Hampshire to the east, t ...
" for fossil hunter Mary Anning. This work, which prominently features plesiosaurs, has been regarded as the first attempt to accurately
reconstruct Reconstruction may refer to: Politics, history, and sociology *Reconstruction (law), the transfer of a company's (or several companies') business to a new company *''Perestroika'' (Russian for "reconstruction"), a late 20th century Soviet Union ...
the
Mesozoic The Mesozoic Era is the Era (geology), era of Earth's Geologic time scale, geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Period (geology), Periods. It is characterized by the dominance of archosaurian r ...
world through an artistic medium. *
von Meyer Meyer is a German surname, often but not always used with the Nobiliary particle "von" (of). Notable people sharing this surname include: * Christian Erich Hermann von Meyer (1801–1869), German palaeontologist, see m:de:Hermann von Meyer * ...
described ''
Pistosaurus ''Pistosaurus'' (exact etymology uncertain) is an extinct genus of aquatic sauropterygian reptile closely related to plesiosaurs. Fossils have been found in France and Germany, and date to the Middle Triassic. It contains a single species, ''Pi ...
''. ''
Pistosaurus ''Pistosaurus'' (exact etymology uncertain) is an extinct genus of aquatic sauropterygian reptile closely related to plesiosaurs. Fossils have been found in France and Germany, and date to the Middle Triassic. It contains a single species, ''Pi ...
'' is believed to be a transitional form linking plesiosaurs to their basal
sauropterygia Sauropterygia ("lizard flippers") is an extinct taxon of diverse, aquatic diapsid reptiles that developed from terrestrial ancestors soon after the end-Permian extinction and flourished during the Triassic before all except for the Plesiosau ...
n forebears.


1840s

1840 * Owen described the species now known as '' Colymbosaurus trochanterius'', '' Eurycleidus arcuatus'', and ''
Thalassiodracon hawkinsii ''Thalassiodracon'' (tha-LAS-ee-o-DRAY-kon) is an extinct genus of plesiosauroid from the Pliosauridae that was alive during the Late Triassic-Early Jurassic (Rhaetian-Hettangian) and is known exclusively from the Lower Lias of England. The type ...
''. 1841 * Owen described the new species ''
Pliosaurus brachydeirus ''Pliosaurus'' (meaning 'more lizard') is an extinct genus of thalassophonean pliosaurid known from the Late Jurassic (Kimmeridgian and Tithonian stages) of Europe and South America. This genus has contained many species in the past but recent re ...
'', '' Pliosaurus brachyspondylus''. and '' Polyptychodon interruptus''. 1842 *
Sir Richard Owen Sir Richard Owen (20 July 1804 – 18 December 1892) was an English biologist, comparative anatomist and palaeontologist. Owen is generally considered to have been an outstanding naturalist with a remarkable gift for interpreting fossils. ...
formally named the
pliosaurs Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive tooth ...
. 1846 * Stutchbury described the species now known as '' Atychodracon megacephalus''. 1848 * The trustees of the British Museum of Natural History bought the type specimen of Plesiosaurus from the estate of the first duke of Buckingham, Richard Glenville. The museum catalogued the specimen as BMNH 22656.


1860s

1863 * Carte and Bailey described the species now known as Rhomaleosaurus cramptoni. 1864 *
Jules Verne Jules Gabriel Verne (;''Longman Pronunciation Dictionary''. ; 8 February 1828 – 24 March 1905) was a French novelist, poet and playwright. His collaboration with the publisher Pierre-Jules Hetzel led to the creation of the ''Voyages extraor ...
depicted a relict plesiosaur's defeat in combat against an
ichthyosaur Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs", although the term is also used for wider clades in which the order resides. Ichthyosaurians thrived during much of the Mesozoic era; based on fo ...
in ''Journey to the Center of the Earth''. 1865 in paleontology, 1865 * Owen described the species now known as ''Eretmosaurus rugosus'' and ''Microcleidus homalospondylus''. * Seeley described the species now known as ''Microcleidus macropterus''. * Owen described the species now known as ''Archaeonectrus rostratus''. 1867 in paleontology, 1867 * An army surgeon named Dr. Theophilus Turner discovered the fossils of a large animal in the Pierre Shale of Kansas, USA. The remains represented the first nearly complete plesiosaur specimen from North America. Turner gave some of its vertebrae to a member of the Union Pacific Railroad's survey named John LeConte. LeConte sent the vertebrae to Edward Drinker Cope for study. Cope recognized the find as a significant plesiosaur discovery and wrote to Turner asking him to excavate and ship the fossils to him. ''March, mid'' * Cope erected the new genus and species ''Elasmosaurus platyurus'' for the fossils sent by Turner in a rushed descriptive manuscript written within two weeks of obtaining them. ''March 24'' * Cope presented his findings regarding ''Elasmosaurus'' to a meeting of the Academy of Natural Sciences in Philadelphia, Pennsylvania. 1868 in paleontology, 1868 * Cope described the new species ''Elasmosaurus platyurus''. * Cope's description of ''Elasmosaurus'' was formally published. ''September'' * William E. Webb and others collected and shipped a plesiosaur specimen to Cope. 1869 in paleontology, 1869 * Seeley described the species now known as ''Liopleurodon pachydeirus'' and ''Peloneustes philarchus''. * Cope named the plesiosaur specimen collected by Webb ''Polycotylus latipinnis.'' ''August'' * Cope prepared a preprint for the Transactions of the American Philosophical Society of his ''Elasmosaurus'' description, including reconstruction of the animal with a short neck and very long tail. The manuscript was then distributed to other scholars.


1870s

1870 in paleontology, 1870 ''March 8'' * Cope's mentor Joseph Leidy gave a presentation reporting his recent discovery that Cope's reconstruction of ''Elasmosaurus'' positioned the skull at the end of the tail rather than the end of the neck. * Leidy's discovery embarrassed Cope, who began spreading notice of an unspecified error in his ''Elasmosaurus'' description with an offer to replace it with a corrected version and its second volume. ''November'' * O. C. Marsh collected a better an additional specimen of ''Polycotylus'' in Kansas that was better preserved than the type described by Cope. The specimen is now catalogued as YPM 1125. 1871 in paleontology, 1871 * Phillips described the species now known as ''Pliosaurus macromerus''. * Phillips described the species now known as ''Cryptoclidus eurymerus''. * Cope described the species now known as ''Hydralmosaurus serpentinus''. * Cope inaccurately referred to ''Polycotylus'' as "the first true plesiosauroid found in America." 1872 in paleontology, 1872 * Cope imagined elasmosaurs feeding by craning their necks above the water and striking downward at fish long distances from their bodies. * B. F. Mudge discovered ten articulated vertebrae in the Fairport Chalk of Kansas that he mistook for ichthyosaur remains. These fossils are now catalogued as KUVP 1325. 1873 in paleontology, 1873 * Sauvage described the new species ''Liopleurodon ferox''. * Joseph Savage discovered a second, better preserved ''Trinacromerum "anonymum"'' in Kansas. 1874 in paleontology, 1874 * Hector described the new species ''Mauisaurus haasti''. * Seeley described the new species ''Muraenosaurus leedsi''. * B. F. Mudge discovered fragments of a large elasmosaur skeleton in the Fort Hays Limestone of Kansas. * Mudge and Williston excavated the remains another large Kansan plesiosaur, this one from the Smoky Hill Chalk. The specimen may be a ''Styxosaurus snowii'' and is currently catalogued as YPM 1644. It was the first plesiosaur Mudge had ever found with gastroliths and the first plesiosaur encountered by Williston in general. * Hector reported the presence of elasmosaur remains in New Zealand. * Seeley published a paper intended to help improve the state of science's understanding of plesiosaur shoulder girdle anatomy, which had been muddled by the poor preservation of the fossils many early paleontologists had to rely on for their observations. 1875 in paleontology, 1875 * Cope once more portrayed elasmosaurs as feeding by fishing from a distance with heads held above the waterline. 1876 in paleontology, 1876 * Blake in Tate and Blake described the new species ''Eretmosaurus, Eretmosaurus dubius'' and ''Plesiosaurus longirostris''. 1877 in paleontology, 1877 * Richard Lydekker, Lydekker described the species now known as ''Simolestes indicus''. * Mudge discussed the gastroliths of YPM 1644 in a scientific publication. He concluded that plesiosaur exploited gastroliths to assist in breaking down food the way many modern birds and reptiles do. 1879 in paleontology, 1879 * Lydekker described the species now known as ''Cryptoclidus richardsoni''.


1880s

1880 in paleontology, 1880 * Oxford acquired the Misses Philpot collection, which included the type specimen of ''Plesiosaurus macromus''. The museum catalogued this specimen as OXFUM J.28587. 1881 in paleontology, 1881 * Sollas described the species now known as ''Attenborosaurus conybeari''. 1882 in paleontology, 1882 * The British Museum of Natural History purchased the Edgerton collection, which included the complete ''Plesiosaurus dolichodeirus'' jaw now known as BMNH R.255. 1884 in paleontology, 1884 ''Spring'' * The Smithsonian bought a partial plesiosaur skeleton from Charles Sternberg. The specimen is now catalogued as USNM 4989 and would later serve as the type specimen of the new genus and species ''Brachauchenius lucasi''. 1888 in paleontology, 1888 * Harry Seeley mistakenly claimed to have discovered several fossil plesiosaur embryos. * F. W. Cragin named the genus and species ''Trinacromerum bentonianum'' from Kansas.


1890s

1890 in paleontology, 1890 * In an article published in the New York Herald, Marsh brought up Cope's anatomic reversal of ''Elasmosaurus''. * E. P. West excavated a skull and partial neck belonging to the elasmosaur that would come to be named ''Styxosaurus snowii''. The specimen is now catalogued as KUVP 1301. * Williston described the species now known as ''Styxosaurus snowii''. 1892 in paleontology, 1892 * Seeley described the species now known as Muraenosaurus beloclis. 1893 in paleontology, 1893 * Marsh described the new species ''Pantosaurus striatus''. * Charles H. Sternberg obtained two large elasmosaur vertebrae that would later serve as the type specimen of ''Elasmosaurus sternbergi''. The specimen is now catalogued as KUVP 1312. * F. W. Cragin discovered a partial plesiosaur skeleton and associated gastroliths in what is now recognized as the Kiowa Shale. This specimen is now catalogued as KUVP 1305 and would later be named ''Plesiosaurus mudgei''. * Williston argued that plesiosaurs ingested gastroliths only accidentally or to relieve "food craving[s]". However, he also observed that the rocks used as gastroliths were more similar to rocks 400–500 miles away in Iowa or the Black Hills of South Dakota than those of the local geology. 1894 in paleontology, 1894 * Cragin described the new species Plesiosaurus mudgei for KUVP 1305. 1895 in paleontology, 1895 * Dames described the species now known as ''Seeleysaurus guilelmiimperatoris''. 1897 in paleontology, 1897 * With guidance from Edward Drinker Cope, paleo-artist Charles R. Knight illustrated an ''Elasmosaurus platyurus'' eating a fish. The elasmosaur's neck was erroneously looped into an anatomically impossible figure 8 configuration that evoked the image of "a Pythonidae, python grasping at its prey". 1898 in paleontology, 1898 * Knight described the new species ''Megalneusaurus rex''. 1899 in paleontology, 1899 * A man named Andrew Crombie discovered a fossil jaw fragment with six teeth in Queensland, Australia. The specimen would become the type for the genus ''Kronosaurus''.


20th century


1900s

1900 in paleontology, 1900 * Knight described the species now known as ''Tatenectes laramiensis''. * George F. Sternberg discovered the plesiosaur specimen now known as KUVP 1300 that would later serve as the type specimen of ''Dolichorhynchops osborni''. 1902 in paleontology, 1902 * Williston described the new species ''Dolichorhynchops osborni''. 1903 in paleontology, 1903 * Williston made several changes to plesiosaur taxonomy. One of these was the description of the new genus and species ''Brachauchenius lucasi'', whose type specimen was a partial skeleton discovered in Kansas. This specimen is now catalogued as USNM 4989. He also described the new species ''Trinacromerum anonymum'' based on the vertebral series discovered by Mudge in 1872. This specimen is now known as KUVP 1325. Lastly, Williston regarded ''Plesiosaurus mudgei'' as a junior synonym of the species Plesiosaurus gouldii. He also commented on the ongoing debate regarding plesiosaur gastroliths, acknowledging the possibility that they were used for ballast while also maintaining openness to his 1893 suggestion that the stones were ingested accidentally. 1904 in paleontology, 1904 * Barnum Brown hypothesized that plesiosaurs used their gastroliths in a gizzard-like organ (biology), organ to grind up their invertebrate prey since they had no grinding or crushing teeth to do that job for them. * Harvard paleontologist Charles R. Eastman, "offended" by Brown's claim that plesiosaurs had a gizzard, criticized the idea in print. * Williston responded to Eastman, reasserting the evidence for plesiosaur gastroliths by noting that by this time at least 30 specimens containing them had been found. 1906 in paleontology, 1906 * Williston described several new taxa and specimens. One of these was the new species ''Elasmosaurus nobilis''. Williston also described the new ''Elasmosaurus'' species ''Elasmosaurus sternbergi, E. sternbergi'' based on the vertebrae discovered by Charles H. Sternberg in 1893. He remarked that these fossils were the largest elasmosaur vertebrae that he had ever seen. Lastly, Williston described Marsh's ''Polycotylus'', YPM 1125. 1907 in paleontology, 1907 * Williston reported the discovery of another ''Brachauchenius'' specimen, although this one was discovered in Paleontology in Texas, Texas. * Williston argued that ''Brachauchenius lucasi'' was closely related to ''Liopleurodon ferox''. 1909 in paleontology, 1909 * Andrews described the new species ''Simolestes vorax''. and ''Tricleidus seeleyi''. * Watson described the new species ''Sthenarasaurus dawkins''.


1910s

1910 in paleontology, 1910 * Fraas described the species now known as ''Rhomaleosaurus victor''. 1911 in paleontology, 1911 * Andrews described the species now known as ''Leptocleidus capensis''. 1913 in paleontology, 1913 * Brown described the new species ''Leurospondylus ultimus''. 1914 in paleontology, 1914 * Williston criticized portrayals of long-necked plesiosaurs as having unnaturally flexible necks. * Wegner described the new species ''Brancasaurus brancai''. * Williston observed that the semicircular canals inside a plesiosaur's ear were well developed, giving them a good sense of balance and coordination. 1919 in paleontology, 1919 * The Smithsonian obtained the ''Tylosaurus'' specimen with preserved polycotylid stomach contents from Charles Sternberg. The ''Tylosaurus'' is catalogued as USNM 8898 and its last supper as USNM 9468.


1920s

1922 in paleontology, 1922 * Andrews described the new species ''Leptocleidus superstes''. * Sternberg observed that being contained in the stomach of a mosasaur might have helped ensure the preservation of the polycotylid now known as USNM 9468 by protecting it from scavenging sharks. 1923 in paleontology, 1923 * Huene described the species now known as ''Hydrorion brachypterygius''. 1924 in paleontology, 1924 * Heber Longman described ''Kronosaurus queenslandicus'' based on the jaw fragment found there by Andrew Crombie in 1899. 1926 in paleontology, 1926 * George F. Sternberg discovered a third specimen of ''Dolichorhynchops osborni'' in Kansas. 1929 in paleontology, 1929 * More ''Kronosaurus'' fossils were discovered in central Queensland near the site of the type specimen's discovery.


1930s

1930 in paleontology, 1930 * Swinton described the new species ''Macroplata tenuiceps''. 1931 in paleontology, 1931 * George F. Sternberg and M. V. Walker discovered a well-preserved large elasmosaur specimen. 1931 in paleontology, 1931-1932 in paleontology, 1932 * Harvard University dispatched a fossil hunting expedition to Queensland, Australia. In Army Downs they discovered a nearly complete specimen of ''Kronosaurus''. 1934 in paleontology, 1934 * The "surgeon's photograph" of the Loch Ness monster was hoaxed, cementing the association between plesiosaurs and the Cryptid, mythical beast. c. 1935 in paleontology, 1935 * The University of Nebraska State Museum bought the elasmosaur specimen discovered by Sternberg and Walker in 1931. The specimen is now catalogued as UNSM 1195. 1935 in paleontology, 1935 * Russell described the new species ''Trinacromerum kirki''. 1936 in paleontology, 1936 * A specimen of ''Trinacromerum'' was discovered in a roadside exposure of the Greenhorn Formation in Kansas. The specimen is now catalogued as KUVP 5070. 1938 in paleontology, 1938 * A large pliosaur skeleton was found on the banks of Russia's Volga River. However, the specimen was damaged during the excavation and only the skull and chest region were successfully extracted.


1940s

1940 in paleontology, 1940 * A complete specimen of ''Plesiosaurus conybeari'', including preserved soft tissues, was destroyed in a bombing raid against Bristol. Fortunately, a cast of the specimen survived in the British Museum. * White described the new species Seeleyosaurus holzmadensis. 1941 in paleontology, 1941 * Cabrera described the new species ''Aristonectes parvidens''. 1942 in paleontology, 1942 * Young described the new species ''Sinopliosaurus weiyuanensis''. 1943 in paleontology, 1943 * Welles described the new species ''Aphrosaurus furlongi'', ''Morenosaurus stocki'', ''Thalassomedon haningtoni'', ''Fresnosaurus drescheri'', and ''Hydrotherosaurus alexandrae''. * Welles argued that plesiosaurs did have flexible necks after all. 1944 in paleontology, 1944 * Elmer S. Riggs named a new species of ''Trinacromerum'', ''T. willistoni''. The type specimen had been found by a construction crew working on US Highway 81, who donated it to the University of Kansas University of Kansas Museum of Paleontology, Museum of Paleontology. * Riggs described the new species ''Trinacromerum willistoni'' based on the 1936 discovery KUVP 5070. 1948 in paleontology, 1948 * Soviet paleontologist Nestor Novozhilov described the Volga pliosaur as a new species, ''Pliosaurus rossicus''. * Novozhilov described the species now known as ''Pliosaurus irgisensis''. 1949 in paleontology, 1949 * Welles described the species now known as ''Libonectes morgani''. * de la Torre and Rojas described the species now known as ''Vinialesaurus caroli''.


1950s

:1950s in paleontology, 1950s * Alfred Sherwood Romer helped mount the ''Kronosaurus'' discovered in Queensland by the 1930s Harvard expedition for the University's Museum of Comparative Zoology. The poorly preserved bones required a significant amount of plaster for the restoration, earning the specimen the mocking nickname "Plasterosaurus". The final mount was 42 feet long, probably due to Romer overestimating the number of vertebrae in its spine; a more likely length is about 35 feet. 1950 in paleontology, 1950 * Fossil hunters Robert and Frank Jennrich serendipitously discovered a partial ''Brachauchenius'' skeleton when looking for sharks teeth. * Shuler, like Williston in 1914, found elasmosaurs to have relatively inflexible necks. He also found elasmosaurs to have stereoscopic vision, which would have been useful for hunting small prey. ''October'' * George Sternberg excavated the ''Brachauchenius'' discovered by the Jennriches. This specimen, now known as FHSM VP-321, was both larger and better preserved than the ''Brachauchenius'' type specimen. Although it was put on display soon after discovery, it would not be described for the scientific literature for nearly 50 years. 1952 in paleontology, 1952 * Welles argued that the "Elasmosaurus sternbergi" type specimen was actually pliosaur vertebrae. 1954 in paleontology, 1954 * A private landowner in Kansas donated some ''Elasmosaurus'' vertebrae to the Sternberg Museum. These fossils are now catalogued as FHSM VP-398.


1960s

1960 in paleontology, 1960 * Tarlo described the new species ''Pliosaurus andrewsi''. 1962 in paleontology, 1962 * Welles described the species now known as ''Callawayasaurus colombiensis''. * Welles reported the presence of elasmosaur remains in South America. * Chatterjee and Zinsmeister reported the presence of elasmosaur remains in Antarctica. 1967 in paleontology, 1967 * Barney H. Newman and Lambert Beverly Halstead Tarlo argued that long-necked plesiosaur flippers could only move horizontally, and while maneuverable, they were confined to surface waters by an inability to dive. * South Australian opal miners John and Molly Addyman discovered a plesiosaur whose remains had been converted to opal.


1970s

1971 in paleontology, 1971 * Beverly Halstead reclassified the Volga pliosaur, ''Pliosaurus rossicus'', to the genus ''Liopleurodon''. * Paul Johnston discovered plesiosaur fossils in a roadside exposure of the Greenhorn Formation in Kansas. During the excavation the dig site was scouted by two suspicious men. After a break from digging the Johnston team returned to find all of the fossils crudely extracted from the rock except for a flipper that the team had reburied. Based on the flipper, the stolen plesiosaur could be identified as ''Trinacromerum bentonianum''. 1975 in paleontology, 1975 * Jane Ann Robinson published a paper on plesiosaur locomotion concluding that they really did swim by "underwater flight" like
sea turtles Sea turtles (superfamily Chelonioidea), sometimes called marine turtles, are reptiles of the order Testudines and of the suborder Cryptodira. The seven existing species of sea turtles are the flatback, green, hawksbill, leatherback, loggerh ...
or penguins. 1976 in paleontology, 1976 * Ochev described the species now known as ''Georgiasaurus penzensis''. 1977 in paleontology, 1977 * Robinson publishes follow up research to her previous publication on plesiosaur locomotion. This second paper notably concluded that plesiosaurs were incapable of leaving the water.


1980s

1980 in paleontology, 1980 * Dong described the new species ''Bishanopliosaurus youngi''. 1981 in paleontology, 1981 * Michael Alan Taylor published a paper concluding that plesiosaurs would have been capable of moving on land after all because their spinal column was too arched for their lungs to collapse. * Brown described the new species ''Kimmerosaurus langhami''. * Brown emended the species ''Plesiosaurus guilelmiiperatoris'' originally described by Dames in 1895. * Taylor argued that plesiosaurs used their gastroliths to adjust buoyancy or to help stay level and balanced while swimming. 1982 in paleontology, 1982 * Samuel F. Tarsitano and Jürgen Riess published a paper harshly critical of Robinson's previous work on plesiosaur locomotion. However, while criticizing Robinson's work they were reluctant to make any positive claims of their own, concluding that the details of plesiosaur locomotion were "unknown". * Richard A. Thulborn published the results of his recent re-examination of the purported plesiosaur embryos discovered by Harry Govier Seeley. Thulborn concluded that Seeley's supposed embryos were actually nodules of mudstone and shale derived from sediments that once filled in a crustacean burrow system and were not even animal body fossils. * Delair described the new species ''Bathyspondylus swindoniensis''. 1984 in paleontology, 1984 * The partial remains of a large pliosaur, initially mistaken for a dinosaur, were discovered near Aramberri, Mexico. 1985 in paleontology, 1985 * Zhang described the new species ''Yuzhoupliosaurus chengjiangensis''. 1986 in paleontology, 1986 * A South Australian opal miner named Joe Vida discovered the skeleton of a juvenile plesiosaur whose remains had converted to opal. Its preparator, Paul Willis (paleontologist), Paul Willis nicknamed it Eric. An entrepreneur named Sid Londish bought the specimen and funded its preparation, but went bankrupt. When the specimen was put up for auction fear spread that a potential buyer might break the specimen down for its gemstone value. A television drive was arranged on behalf of the Australian Museum. The Museum succeeded in raising 340,000 dollars to buy the specimen, whose gemstone value was about $300,000. Eric was later identified as a specimen of ''Leptocleidus''. * Wiffen and Moisley described the new species ''Tuarangisaurus keysei''. 1987 in paleontology, 1987 * Judith Massare published an analysis of plesiosaur feeding habits. She concluded that the long-necked plesiosauroids ate soft prey. ''Liopleurodon'' and its relatives, on the other hand, had teeth resembling those of killer whales and probably ate larger, bonier prey. * Orville Bonner discovered a specimen of ''Dolichorhynchops osborni'' that was later seen to preserve developing young inside it. 1988 in paleontology, 1988 * Judy Massare analyzed Mesozoic marine reptile swimming abilities and found that long-necked plesiosaurs would have been significantly slower than pliosaurs due to excess drag incurred from the length of the neck. * The Los Angeles County Museum of Natural History acquired the ''Dolichorhynchops osborni'' specimen discovered by Bonner and catalogued it as LACMNH 129639. 1989 in paleontology, 1989 * Beverly Halstead published a paper suggesting that plesiosaurs swam using all four flippers paired with an undulatory motion of the body comparable to a sea lion's. * Nakaya reported the presence of elasmosaur remains in Japan.


1990s

1990 in paleontology, 1990 * The world's smallest plesiosaur, between four and five feet in length, was discovered near Charmouth on the Dorset, Dorset Coast. * Sciau et al. described the species now known as ''Occitanosaurus tournemirensis''. * Gasparini and Spalleti described the new species ''Sulcusuchus erraini''. * Stewart noted a relative paucity of plesiosaur fossils from the lower portions of the Smoky Hill Chalk in a manuscript for the Society of Vertebrate Paleontology. *Tim Tokaryk of the Royal Saskatchewan Museum discovers a new kind of plesiosaur, Dolichorhynchops herschelensis near Herschel, Saskatchewan. ''May'' * J. D. Stewart, accompanied by Everhart, discovered a nearly complete ''Dolichorhynchops'' rear flipper in the lower Smoky Hill Chalk. Unfortunately it was too late to correct the erroneous statements in his aforementioned paper regarding the supposed rarity of plesiosaurs in the lower Smoky Hill Chalk. The flipper is now catalogued as LACMNH 148920. ''October'' * Stewart's paper, complete with his now-erroneous statements, was published in the Niobrara Chalk Excursion Guidebook in honor of the society's 50th anniversary meeting in Lawrence that year. 1991 in paleontology, 1991 * Ralph E. Molnar published suspicion that the "''Kronosaurus queenslandicus''" specimen discovered by the Harvard expedition might be a distinct species. * Several Elasmosaurus vertebrae and gastroliths were found near the site where the type specimen of the genus itself was excavated. * Cruikshank and others hypothesized that plesiosaurs could smell and taste water that "passively" flowed through their nasal passages while they swam. 1992 in paleontology, 1992 * Hampe described the new species ''Kronosaurus boyacensis''. * Everhart discovered some fragmentary plesiosaur fossils in the lower Smoky Hill Chalk of Kansas. Some of the fossils seemed to have been partially digested. The remains were later catalogued as FHSM VP-13966. * Everhart showed the partially digested fossils to J. D. Stewart, who recognized them as pieces of a plesiosaur skull. The fossils are now catalogued as * Everhart and his wife helped excavated a ''Styxosaurus snowii'' specimen in Kansas. During the dig Mrs. Everhart discovered an additional partial plesiosaur skeleton. 1993 in paleontology, 1993 * Robert T. Bakker nicknamed the long-necked plesiosaurs "swan lizards". * Robert T. Bakker argued that plesiosaurs suffered several major extinction events throughout the Mesozoic, rather than continuously diversifying until the Cretaceous–Paleogene extinction event, end-Cretaceous mass extinction. * Robert T. Bakker argued that ''Dolichorhynchops'' and its relatives became the most common kind of short-necked plesiosaur in the Cretaceous Western Interior Seaway after the ichthyosaurs became extinct. Further, they convergently evolved many traits similar to those of ichthyosaurs, such as long snouts and large eyes. * Tony Thulborn and Susan Turner examined the crushed skull of the long-necked plesiosaur ''Woolungasaurus'' and found the presence of tooth marks left by some giant predator. They hypothesized that a ''Kronosaurus'' was the culprit. * Storrs, like Williston and Shuler before him, argued that long-necked plesiosaurs had relatively inflexible necks. * Rothschild and Martin reported the presence of the remains of a fossilized fetus preserved in the abdomen of a ''Dolichorhynchops osborni''. 1994 in paleontology, 1994 * Glenn W. Storrs formally described the world's smallest plesiosaur for the scientific literature. * South Dakota School of Mines paleontologist James Martin excavated and described a very young pliosaur specimen. The specimen was in what would have been the middle of the Cretaceous Western Interior Seaway. It is more plausible to think that this young individual was born in the seaway itself rather than surviving the extremely long swim from shore. So, the specimen provided circumstantial evidence for live birth and possibly even parental care by pliosaurs in the Cretaceous Western Interior Seaway. However, since there were no adult fossils present this interpretation is speculative. Further, the fossil site had been heavily vandalized before discovery by scientists. A large hole was found near the baby pliosaur that could have once held the bones of its mother or other pod members. * An amateur fossil hunter named Simon Carpenter discovered a 7-foot-long '' Pliosaurus brachyspondylus'' skull in claypits of the Blue Circle cement works near Westbury, Wiltshire, Westbury, England. More of the skeleton was found in the vicinity and this specimen came to be regarded as the best preserved pliosaur ever found; it is held by Bristol Museum & Art Gallery, Bristol Museum. * A man named Alan Dawn discovered a previously unknown kind of pliosaur in the Middle Jurassic Oxford Clay of Cambridgeshire, England. Godefroit described the new species ''Simolestes keileni''. * Ken Carpenter recognized the plesiosaur specimen discovered by Pamela Everhart in 1992 as one of the largest known specimens of ''Dolichorhynchops osborni'', now catalogued as CMC VP-7055. 1996 in paleontology, 1996 * Carpenter published a review of the Cretaceous short-necked plesiosaurs known from western North America. In this paper he both revised these plesiosaurs' taxonomy as well as offering observations on their biostratigraphy and evolution. Carpenter described the new genus and species Plesiopleurodon wellesi. He also argued against the prevailing trend to treat ''Dolichorhynchops'' ''Trinacromerum'' as taxonomic synonyms by observing that they could be distinguished based on their skull anatomy. However, he did conclude that the Trinacromerum species T. anonymum and T. willistoni were junior synonyms of T. bentonianum. In his remarks on short-necked plesiosaur evolution, Carpenter argued that polycotylids were more closely related to long-necked plesiosaurs than pliosaurs.Everhart (2005); "Pliosaurs and Polycotylids", page 144. He observed that ''Trinacromerum bentonianum'' seems to have existed from the late Cenomanian to the Turonian. This represents a span of time approximating 3.3 million years. He found ''Dolichorhynchops osborni'' to have had an even longer lifespan, from the middle Turonian to the early Campanian., or roughly 4 million years. His research also suggested that there was a span of time during the life of the Western Interior Seaway in which it was not inhabited by polycotylids. He also reported that the ''Dolichorhynchops'' specimen KUVP 40001 from the Pierre Shale of South Dakota may have achieved the extraordinary length of 23 feet. The large size of the Pierre Shale ''Dolichorhynchops'' compared to those of the earlier Smoky Hill Chalk suggested to Carpenter that these plesiosaurs were evolving larger body sizes over time. In fact the Pierre Shale specimens of ''Dolichorhynchops'' were nearly as large as ''Brachauchenius lucasi''. Carpenter described a particularly large specimen of that latter taxon in this paper as well, specifically FHSM VP-321. His study of ''Brachauchenius'' led him to concur with Williston that it was closely related to ''Liopleurodon ferox''. * Arthur R. I. Cruickshank, David M. Martill, and Leslie Noe formally named the new Oxford Clay pliosaur ''Pachycostasaurus dawni''. The researchers noticed that its bones were very dense. So dense, they speculated it would naturally sink in the water and spent most of its time feeding on soft bodied animals living near the seafloor. 1997 in paleontology, 1997 * Cruickshank and Long described the new species ''Leptocleidus clemai''. * Gasparini described the new species ''Maresaurus coccai''. * Liggett and others reported the discovery of a giant plesiosaur flipper from the Greenhorn Limestone of Kansas. Although a significant portion of the specimen was missing, it implied a life length of more than 2 m. The researchers tentatively attributed the flipper to ''Brachauchenius lucasi''. The specimen is now catalogued as FHSM VP-13997. *Two fossilized skeletons of Dolichorhynchops herschelensis are discovered near Herschel, Saskatchewan at the Ancient Echoes Interpretive Centre- only the second and third specimens to have ever been found. 1998 in paleontology, 1998 * Tamaki Sato and Kazushige Tanabe reported that a partial long necked plesiosaur skeleton had been discovered on the banks of Hokkaido's Obirashibe River. Although they could not identify its exact species, the specimen was significant because its stomach contents provided clues to long-necked plesiosaur diets. This individual's gut preserved the beaks of thirty small ammonites. * John A. Long bemoaned the fact that the putative "''Kronosaurus queenslandicus''" uncovered by a Harvard team during the early 1930s had still not been formally described for the scientific literature. * Michael Everhart and Glenn Storrs excavated additional ''Elasmosaurus'' ribs, vertebrae and gastroliths at the site of the 1991 discovery. * Long reported the presence of elasmosaur remains in Australia. 1999 in paleontology, 1999 * Carpenter published a summary of the elasmosaur fossils discovered in the Smoky Hill Chalk. * Storrs published a revision of ''Elasmosaurus'' taxonomy. He reinterpreted the ''Elasmosaurus nobilis'' type specimen as indeterminate elasmosaurid remains. He also reinterpreted the "Elasmosaurus" sternbergi type specimen as two cervical and one dorsal vertebrae rather than two dorsal vertebrae as Williston had reported in his original description. However, Storrs did agree that it was an elasmosaur specimen rather than a pliosaur as argued by Welles in 1952.


21st century


2000s

2000 in paleontology, 2000 * Theagarten Lingham-Soliar published further criticism of Robinson's interpretation of the biomechanics of plesiosaur locomotion. * O'Keefe described the new species Hauffiosaurus zanoni. * Michael Everhart re-examined UNSM 1195. * Lingham-Soliar argued that plesiosaur hind-flippers weren't mobile or muscular enough to help propel them through the water. * Everhart published a study of the gastroliths associated with the elasmosaur specimen KUVP 129744 from Kansas. The specimen was associated with roughly 13.1 kg of gastroliths. The largest of these was 17 cm long and 1.4 kg in weight. Everhart would later compare its size to that of a softball and observe that not only was it one of the largest known plesiosaur gastroliths, but also one of the largest gastroliths from any animal. ''November'' * ''The Advertiser (Adelaide), The Advertiser'', a newspaper based in Adelaide, Australia bought the Addyman opalized plesiosaur specimen for $25,000 and donated it to the South Australian Museum. A paleontologist at the museum named Ben Kear identified it as a member of the genus ''Leptocleidus''. The two foot long specimen was the smallest specimen of the genus ever found and probably a baby. 2001 in paleontology, 2001 * David J. Cicimiurri and Michael J. Everhart published a study of the ''Styxosaurus snowii'' specimen NJSM 15435, which preserved both stomach contents and gastroliths. Among the stomach contents were remains of the bony fish Enchodus. By this point in time at least fifteen different plesiosaur specimens were known with preserved stomach contents. The researchers observed that the Enchodus remains preserved in NJSM 15435 were an example of shifting dietary preferences in plesiosaurs, who fed primarily on cephalopods for most of their evolutionary history, before coming to rely more heavily on fishes during the Late Cretaceous. They also noted that some of NJSM 15435's gastroliths were scarred by rounded chips and arc-shaped marks. These were likely inflicted by contact with other gastroliths during the churning of the animal's stomach, and constituted physical evidence that plesiosaurs used their gastroliths to help break down their food during digestion. Cicimurri and Everhart disputed the hypothesis that plesiosaurs used their gastroliths for ballast on the grounds that swallowing and vomiting such stones would be relatively difficult for the long-necked forms and their feeding grounds may have been hundreds of miles from sources of stones. * Everhart resumed the study of the partially digested plesiosaur skull bones, FHSM VP-13966. He sought the expertise of Ken Carpenter due to his relevant 1996 paper on short-necked plesiosaurs. Carpenter identified the bones as probable ''Dolichorhynchops'' remains. * Noe published another study of ''Pachycostasaurus''. He changed his mind regarding its diet. Where previously he believed it to feed on soft-bodied animals, the robust and "heavily ornamented" build of its teeth suggested it fed on harder, bonier prey. ''September'' * Eberhard Frey, Celine Bachy, and Wolfgang Stinnesbeck gave a presentation on the Aramberri pliosaur remains to the European Workshop on Vertebrate Paleontology in Florence, Italy. The paleontologists could not identify its species. ''September 11'' * Everhart was forced to cancel plans to examined the ''Tylosaurus'' specimen USNM 8898 and its polycotylid dinner USNM 9468 due to the September 11th terrorist attacks. ''November'' * Everhart was finally able to examine the tylosaur specimen with the polycotylid stomach contents. 2001–2002 in paleontology, 2002 * Robin O'Keefe publishes a series of papers examining the aspect ratios of plesiosaur flippers and comparing them to the wings of birds, bats, and aircraft. O'Keefe concluded that long-necked plesiosaurs were long-distance swimmers, while pliosaurs were more maneuverable. 2002 in paleontology, 2002 * Cruickshank and Fordyce described the new species ''Kaiwhekea katiki''. * Druckenmiller described the new species ''Edgarosaurus muddi''. * Michael Everhart examined FHSM VP-398 and found Sternberg's original note revealing that these fossils had been collected at the same site as the 1991 Elasmosaurus discovery. Everhart realized that the remains discovered there collectively represented most of the bones that had been missing from the Elasmosaurus type specimen. He inferred that they may represent fragments that fell off of the decomposing type carcass while it was adrift, before its final burial and fossilization. * An elasmosaur specimen with over 600 associated gastroliths was discovered in the Pierre Shale of Nebraska. The specimen is now catalogued as UNSM 1111–002. ''December 30'' * The BBC began making sensational claims about the Aramberri pliosaur, claiming that the unidentified pliosaur was a ''Liopleurodon'', that it was 65 feet long, that its teeth were the size of machetes, and its jaws powerful enough to crush granite. 2003 in paleontology, 2003 * The University of Nebraska led the excavation of a long-necked plesiosaur skeleton on local Santee Sioux land. The Santee people requested that the skeleton be mounted and displayed with a plaque acknowledging them as the source of the fossils and as having given permission for the University of Nebraska State Museum, museum to display the remains. However, the museum claims it could not honor the request as it did not have the funding to mount the skeleton for display, and it further claimed that the land the fossils were recovered from was of "disputed" ownership. * Mulder and others reported the presence of elasmosaur remains in Europe. * Sato described the new genus and species ''Terminonatator ponteixensis''. In his study of the animal's skeleton, he found that the vertebral discs in the neck were flat on both sides and packed tightly together. He estimated that there would have been only about 0.5 cm of cartilaginous padding between these discs. These observations provided additional evidence for a lack of flexibility in plesiosaur necks. * Everhart argued contrary to Carpenter's 1996 paper that polycotylids were present throughout the life of the Western Interior Seaway. * Everhart finally described the partially digested partial plesiosaur skull he discovered in 1992. These were among the earliest known plesiosaur fossils in the Smoky Hill Chalk. He has since concluded that the animal that partially digested the remains was probably a shark, which would go on to vomit them up before they were buried and preserved. * Bardet and others described the new species ''Thililua longicollis''. 2004 in paleontology, 2004 * Michael Everhart found Charles H. Sternberg's account of the discovery of the ''Elasmosaurus sternbergi'' type specimen in his 1932 book. This allowed Everhart to verify the specimen's geographic and stratigraphic provenance. * Everhart argued that the greater abundance of arc shaped marks and rounded divots in plesiosaur gastroliths compared to rocks deposited by ancient rivers and sea shores was evidence for their use in the breakdown of plesiosaurs' food. * Everhart redescribed the ''Tylosaurus'' specimen USNM 8898 and its polycotylid dinner USNM 9468. Contrary to Sternberg's original assessment of the stomach contents as representing a "huge plesiosaur" Everhart found it to be a young polycotylid only about 2-2.5 m long. * Noe et al. described the new species ''Pliosaurus portentificus''. 2005 in paleontology, 2005 * Sato described the new species ''Dolichorhynchops herschelensis''. * Sachs described the species now known as ''Eromangasaurus australis''. * Buchy et al. described the new species ''Manemergus anguirostris''. *The plesiosaur remains found at Ancient Echoes Interpretive Centre#History & Timeline, Ancient Echoes Interpretive Centre near Herschel, Saskatchewan, a completely new species, Dolichorhynchops herschelensis, by Dr. Tamaki Sato, a Japanese vertebrate paleontologist. 2006 in paleontology, 2006 * Buchy described the new species ''Libonectes atlasense''. * Kear described the new species ''Umoonasaurus demoscyllus''. * Kear described the new species ''Opallionectes andamookaensis''. * Sato et al. described the new species ''Futabasaurus suzukii''. 2007 in paleontology, 2007 * Albright et al. described the new species ''Eopolycotylus rankini''. * Albright et al. described the new species ''Palmula quadratus''. 2008 in paleontology, 2008 2009 in paleontology, 2009


2010s

2010 in paleontology, 2010 * Sennikov and Arkhangelsky described the new genus and species ''Alexeyisaurus karnoushenkoi''. * Smith and Vincent described the new genus ''Meyerasaurus''. 2011 in paleontology, 2011 *
F. Robin O'Keefe F is the sixth letter of the Latin alphabet. F may also refer to: Science and technology Mathematics * F or f, the number 15 in hexadecimal and higher positional systems * ''p'F'q'', the hypergeometric function * F-distribution, a conti ...
and Luis M. Chiappe concluded the debate on plesiosaur reproduction, reporting the discovery of a gravid female plesiosaur with a single large embryo preserved inside her. They concluded that like marine mammals, but unlike many reptiles, plesiosaurs had a K-selected reproductive strategy. * Berezin described the new genus and species ''Abyssosaurus nataliae'' * Benson and others described the new species ''Hauffiosaurus tomistomimus'' * Ketchum and Benson described the new genus and species ''Marmornectes candrewi'' * Schwermann and Sander described the new genus and species ''Westphaliasaurus simonsensii '' * Vincent and others described the new genus and species ''Zarafasaura oceanis'' 2012 in paleontology, 2012 * Kubo, Mitchell and Henderson described the new genus and species ''Albertonectes vanderveldei''. * Vincent and Benson described the new genus and species ''Anningasaura lymense''. * Benson, Evans and Druckenmiller described the new genera and species ''Avalonnectes arturi'', ''Eoplesiosaurus antiquior'', and ''Stratesaurus taylori''. * Knutsen, Druckenmiller and Hurum described the new genus and species ''Djupedalia engeri''. * McKean described the new species ''Dolichorhynchops tropicensis''. * Smith, Araújo and Mateus described the new genus and species ''Lusonectes sauvagei''. * Knutsen, Druckenmiller and Hurum described the new species ''Pliosaurus, Pliosaurus funkei'' * Knutsen, Druckenmiller and Hurum described the new genus ''Spitrasaurus'' and two species, ''S. wensaasi'', and ''S. larseni''. 2013 in paleontology, 2013 * Benson and others described the new genus ''Hastanectes'' and the new genus and species ''Vectocleidus pastorum''. * Vincent, Bardet and Mattioli described the new genus and species ''Cryonectes neustriacus'' * Hampe described the new genus and species ''Gronausaurus wegneri'' * Schumacher, Carpenter and Everhart described the new genus and species ''Megacephalosaurus eulerti'' * Benson and others described the new ''Pliosaurus'' species ''P. carpenteri'', ''P. kevani'', and ''P. westburyensis''. 2014 in paleontology, 2014 * Otero and others described the new species ''Aristonectes quiriquinensis''. * Gasparini and O’Gorman described the new species ''Pliosaurus, Pliosaurus patagonicus''. 2015 in paleontology, 2015 * Cau and Fanti described the new genus and species ''Anguanax zignoi''. * Smith described the new genus ''Atychodracon''. * Araújo and others described the new genus and species ''Cardiocorax mukulu''. * O’Gorman and others described the new genus and species ''Vegasaurus molyi''. * A study on the teeth replacement patterns during the ontogeny in Pliosauridae, pliosaurids is published by Sassoon, Foffa & Marek (2015). 2016 in paleontology, 2016 * Otero and others described the new genus and species ''Alexandronectes, Alexandronectes zealandiensis''. * O’Gorman described the new genus ''Kawanectes''. * Páramo and others described the new genus and species ''Stenorhynchosaurus, Stenorhynchosaurus munozi''. * Cheng and others described the new genus ''Dawazisaurus''. * Klein and others described the new species ''Lariosaurus vosseveldensis''. * Efimov, Meleshin and Nikiforov described the new species ''Polycotylus sopozkoi''. * A reassessment of fossils attributed to the genus ''Polyptychodon'' is published by Madzia (2016), who considers the type species of this genus, ''P. interruptus'', to be ''nomen dubium'', and the genus ''Polyptychodon'' to be a wastebasket taxon. * O'Gorman (2016) provides a new diagnosis for ''Fresnosaurus drescheri'' and describes additional Plesiosauria, plesiosaur material from the Late Cretaceous (Maastrichtian) Moreno Formation (California, USA), which he interprets as representing the first Aristonectinae, aristonectine plesiosaur reported from the Northern Hemisphere. * A redescription of the holotype specimen of ''Brancasaurus brancai'' and a study on the phylogenetic relationships of the species is published by Sachs, Hornung & Kear (2016), who consider the species ''Gronausaurus wegneri'' to be a Synonym (taxonomy), junior synonym of ''B. brancai''. 2017 in paleontology, 2017 * Gómez-Pérez and Noè described the new genus and species ''Acostasaurus pavachoquensis''. * Sachs, Hornung, and Kear described the new genus and species ''Lagenanectes richterae''. * Fischer and others described the new genus and species ''Luskhan itilensis''. * Frey and others described the new genus and species ''Mauriciosaurus fernandezi''. * Serratos, Druckenmiller, and Benson described the new genus and species ''Nakonanectes bradti''. * Wintrich and others described the new genus and species ''Rhaeticosaurus mertensi''. * Smith and Araújo described the new genus and species ''Thaumatodracon wiedenrothi''. * A study on the mechanisms generating vertebral counts and their regionalisation during embryo development that were responsible for high plasticity of the body plan of
sauropterygia Sauropterygia ("lizard flippers") is an extinct taxon of diverse, aquatic diapsid reptiles that developed from terrestrial ancestors soon after the end-Permian extinction and flourished during the Triassic before all except for the Plesiosau ...
ns is published by Soul & Benson (2017). * A study on the function of the long neck in Plesiosauria, plesiosaurs as indicated by the anatomy of the neck is published by Noè, Taylor & Gómez-Pérez (2017). * A study on the large, paired openings in the neck vertebrae of plesiosaurs and their implications for inferring the anatomy of the vascular system in the neck of plesiosaurs is published by Wintrich, Scaal & Sander (2017). * A study on the swimming method of plesiosaurs is published by Muscutt ''et al.'' (2017). * An assessment of the completeness of the plesiosaur fossil record is published by Tutin & Butler (2017). * A description of a new specimen of ''Colymbosaurus svalbardensis'' from the Tithonian–Berriasian Agardhfjellet Formation (Svalbard, Norway), a reevaluation of the diagnostic features of the species and a study on its phylogenetic relationships is published by Roberts ''et al.'' (2017). * A study on the tooth formation cycle in Elasmosauridae, elasmosaurid plesiosaurs is published by Kear ''et al.'' (2017). * A redescription of the holotype specimen of ''Tuarangisaurus keyesi'' and a study on the phylogenetic relationships of the species is published by O'Gorman ''et al.'' (2017). * A study on the anatomy of the vertebra of ''Vegasaurus molyi'' and its implications for the anatomy of the nervous system of the species is published by O'Gorman & Fernandez (2017). * A study on the skeletal Morphology (biology), morphology and histology of a Prenatal development#Definitions of periods, perinatal Aristonectinae, aristonectine Plesiosauria, plesiosaur specimen recovered from the Lopez de Bertodano Formation (Seymour Island, Antarctica) is published by O'Gorman, Talevi & Fernández (2017). * A redescription of the anatomy of the holotype skull of ''Morturneria seymourensis'' is published by O'Keefe ''et al.'' (2017). * A reappraisal and a study on the phylogenetic relationships of ''Mauisaurus'' is published by Hiller ''et al.'' (2017). * ''Libonectes atlasense'' is redescribed by Sachs & Kear (2017), who consider this species to be likely Synonym (taxonomy), synonymous with ''Libonectes morgani''. * An elasmosaurid specimen closely related to ''Vegasaurus molyi'', ''Kawanectes lafquenianum'', ''Morenosaurus stocki'' and aristonectines is described from the Late Cretaceous (late Maastrichtian) Lopez de Bertodano Formation (Antarctica) by O’Gorman & Rodolfo Coria, Coria (2017), who name a new elasmosaurid clade Weddellonectia. 2018 in paleontology, 2018 * Sachs and Kear described the new genus and species ''Arminisaurus schuberti''. * O’Gorman, Gasparini and Spalletti described the new species ''Pliosaurus almanzaensis''. * Páramo-Fonseca, Benavides-Cabra and Gutiérrez described the new genus and species ''Sachicasaurus vitae''. * De Miguel Chaves, Ortega and Pérez‐García described the new genus and species ''Paludidraco multidentatus''. * A study aiming to estimate metabolic rates and bone growth rates in eosauropterygians, especially in Plesiosauria, plesiosaurs, is published by Fleischle, Wintrich & Sander (2018). * A study on the variability of the skull Morphology (biology), morphology in ''Simosaurus gaillardoti'' is published by de Miguel Chaves, Ortega & Pérez-García (2018). * An incomplete mandible of a large-bodied predatory Plesiosauria, plesiosaur is described from the Lower Cretaceous (Barremian) Deister Formation (Germany) by Sachs ''et al.'' (2018). * The first Jurassic plesiosaur from Antarctica is described from the Upper Jurassic Ameghino (= Nordensköld) Formation (Antarctic Peninsula) by O’Gorman ''et al.'' (2018). * Morphology (biology), Morphologically diverse Pliosauridae, pliosaurid teeth are described from the Upper Jurassic (Tithonian) of the Kheta river basin (Eastern Siberia, Russia) and from the Lower Cretaceous (Berriasian and Valanginian) of the Volga region (European Russia) by Zverkov ''et al.'' (2018), who argue that their findings challenge the hypothesis that only one lineage of pliosaurids crossed the Jurassic–Cretaceous boundary. * Complete mandible of ''Kronosaurus queenslandicus'' is described from the Albian Allaru Formation, Allaru Mudstone (Australia) by Holland (2018). * Description of the skull bones of ''Abyssosaurus nataliae'' from the Cretaceous (Hauterivian) of Chuvashia (Russia) is published by Berezin (2018), who also revises the species diagnosis. * A study on a specimen of ''Cryptoclidus eurymerus'' from the Middle Jurassic (Callovian) of Peterborough (United Kingdom), with the left forelimb injured by a predator causing the loss of use of this limb but which nevertheless survived for some time after that injury, is published by Rothschild, Clark & Clark (2018), who also evaluate the implications of this specimen for the various hypotheses on plesiosaur propulsion. * A study on the range of motion of the neck of an exceptionally preserved specimen of ''Nichollssaura borealis'' is published by Nagesan, Henderson & Anderson (2018). * A study on the morphology of ''Thililua longicollis'' and on the phylogenetic relationships of members of the family Polycotylidae is published by Fischer ''et al.'' (2018), who name a new clade Occultonectia. * Two new plesiosaur specimens, including a specimen of the species ''Libonectes morgani'' (otherwise known from North American fossils), are described from the Upper Cretaceous (Turonian) deposits of Goulmima (Morocco) by Allemand ''et al.'' (2018). * Description of a skull and partial postcranial skeleton of a juvenile Elasmosauridae, elasmosaurid from the Upper Cretaceous Tahora Formation (New Zealand), referred to the species ''Tuarangisaurus keyesi'', is published by Otero ''et al.'' (2018). * An exceptionally well-preserved elasmosaurid basicranium, providing new information on the anatomy of the skull of elasmosaurids, is described from the Upper Cretaceous (lower Campanian) Rybushka Formation (Russia) by Zverkov, Averianov & Popov (2018). * Redescription of ''Aristonectes quiriquinensis'', providing new information on the anatomy of this species, is published by Otero, Soto-Acuña & O'keefe (2018). * Cranial material of a non-aristonectine elasmosaurid plesiosaur is described from the Upper Cretaceous (Maastrichtian) Cape Lamb Member of the Snow Hill Island Formation (Vega Island, Antarctica) by O'Gorman ''et al.'' (2018). * New elasmosaurid specimen is described from the upper Maatrichtian horizons of the Lopez de Bertodano Formation (Antarctica) by O’Gorman ''et al.'' (2018), representing one of the youngest non-aristonectine weddellonectian elasmosaurid specimens from Antarctica reported so far, documenting the presence of at least two different non-aristonectine elasmosaurids in Antarctica during the late Maastrichtian, and confirming the coexistence of aristonectine and non-aristonectine elasmosaurids in Antarctica until the end of the Cretaceous. * Redescription of the holotype of ''Styxosaurus snowii'' and a study on the phylogenetic relationships of this species is published by Sachs, Lindgren & Kear (2018). 2019 in paleontology, 2019 * Pathological fusions of neck vertebrae are reported in four plesiosaur specimens from different geological horizons by Sassoon (2019). * A study on the morphology of the teeth and skull of ''Megacephalosaurus eulerti'', and on their implications for assessing the phylogenetic relationships of this species, will be published by Madzia, Sachs & Lindgren (2019). * New plesiosaur fossils are described from the Barremian levels of the Arcillas de Morella Formation (Spain) by Quesada ''et al.'' (2019), including the first Leptocleididae, leptocleidid fossil reported from the Iberian Peninsula. * A study on the skull Morphology (biology), morphology of two specimens of ''Dolichorhynchops bonneri'' from the Pierre Shale of South Dakota, as well as on the phylogenetic relationships of this species, is published by Morgan & O'Keefe (2019). * A study on bone histology and ontogeny of the gravid specimen of ''Polycotylus latipinnus'' displayed at the Los Angeles County Museum of Natural History, and on its implications for interpreting a histological growth series in ''Dolichorhynchops bonneri'', is published by O’Keefe ''et al.'' (2019). * Skull and neck bones of an Elasmosauridae, elasmosaurid plesiosaur are described from the Cenomanian Hegushi Formation (Japan) by Utsunomiya (2019), representing the oldest confirmed elasmosaurid in Japan and in East Asia. * Páramo Fonseca and others described the new genus and species ''Leivanectes bernardoi''. * Vincent and Storrs described the new genus and species ''Lindwurmia thiuda''. * Vincent and others described the new species ''Microcleidus melusinae''.


2020s

2020 in paleontology, 2020 * Roberts and others described the new genus and species ''Ophthalmothule cryostea''. 2023 in paleontology, 2023 * Sachs and others referred ''Simolestes keileni'' to a new genus ''Lorrainosaurus''. * Clark and others describe the new genus and species ''Unktaheela specta'' and move the species "''Dolichorhynchops''" ''bonneri'' to the new genus ''Martinectes'' and "''Dolichorhynchops''" ''tropicensis'' to the new genus ''Scalamagnus''. 2024 in paleontology, 2024 * Sachs, Eggmaier and Madzia described a new species and genus of plesiosauroid plesiosaur, ''Franconiasaurus, Franconiasaurus brevispinus'', from the Toarcian deposits of Germany.


See also

* Timeline of paleontology * Timeline of ichthyosaur research * Timeline of mosasaur research * List of plesiosaurs


Footnotes


References


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External links

* {{Portal bar, Paleontology, History of Science Paleontology timelines, plesiosaur research Plesiosaurs Plesiosaur-related lists Zoology timelines, plesiosaur research