Timothy M. Caro (born 1951) is a British
evolutionary ecologist known for his work on
conservation biology
Conservation biology is the study of the conservation of nature and of Earth's biodiversity with the aim of protecting species, their habitats, and ecosystems from excessive rates of extinction and the erosion of biotic interactions. It is an ...
,
animal behaviour
Ethology is the scientific study of animal behaviour, usually with a focus on behaviour under natural conditions, and viewing behaviour as an evolutionarily adaptive trait. Behaviourism as a term also describes the scientific and objective ...
,
anti-predator defences in animals, and the function of
zebra
Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: the Grévy's zebra (''Equus grevyi''), plains zebra (''E. quagga''), and the mountain zebra (''E. ...
stripes. He is the author of several textbooks on these subjects.
Life
Caro was born in 1951 to artists
Anthony Caro
Sir Anthony Alfred Caro (8 March 192423 October 2013) was an English abstract art, abstract sculptor whose work is characterised by assemblages of metal using 'Found object, found' industrial objects. His style was of the modernist school, hav ...
and
Sheila Girling
Sheila May Girling (1 July 1924 – 14 February 2015) was a British artist, and wife of the sculptor Anthony Caro.
Girling was born in Birmingham and studied at the Royal Academy. Her first solo exhibitions were in Canada but she also exhibited w ...
. Caro gained his bachelor's degree in
zoology
Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and ...
at
Cambridge University
, mottoeng = Literal: From here, light and sacred draughts.
Non literal: From this place, we gain enlightenment and precious knowledge.
, established =
, other_name = The Chancellor, Masters and Schola ...
in 1973, and his doctorate in
psychology
Psychology is the scientific study of mind and behavior. Psychology includes the study of conscious and unconscious phenomena, including feelings and thoughts. It is an academic discipline of immense scope, crossing the boundaries betwe ...
at the
University of St Andrews
(Aien aristeuein)
, motto_lang = grc
, mottoeng = Ever to ExcelorEver to be the Best
, established =
, type = Public research university
Ancient university
, endowment ...
in 1979. He was a professor of wildlife biology at
University of California, Davis
The University of California, Davis (UC Davis, UCD, or Davis) is a public land-grant research university near Davis, California. Named a Public Ivy, it is the northernmost of the ten campuses of the University of California system. The inst ...
, in the departments of population biology and wildlife and fish conservation biology. He is currently a professor of biology at the
University of Bristol
The University of Bristol is a Red brick university, red brick Russell Group research university in Bristol, England. It received its royal charter in 1909, although it can trace its roots to a Society of Merchant Venturers, Merchant Venturers' sc ...
. He has studied the colour
polymorphism
Polymorphism, polymorphic, polymorph, polymorphous, or polymorphy may refer to:
Computing
* Polymorphism (computer science), the ability in programming to present the same programming interface for differing underlying forms
* Ad hoc polymorphis ...
of
coconut crab
The coconut crab (''Birgus latro'') is a species of terrestrial hermit crab, also known as the robber crab or palm thief. It is the largest terrestrial arthropod in the world, with a weight of up to . It can grow to up to in width from the tip ...
s, the
conservation
Conservation is the preservation or efficient use of resources, or the conservation of various quantities under physical laws.
Conservation may also refer to:
Environment and natural resources
* Nature conservation, the protection and managem ...
of fragments of
forest
A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
, and the function of
coloration in mammals, especially
zebra
Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: the Grévy's zebra (''Equus grevyi''), plains zebra (''E. quagga''), and the mountain zebra (''E. ...
stripes.
How the zebra got its stripes

Caro's team found evidence that zebra stripes help to reduce biting by
tabanid flies, but no reliable support for traditionally held hypotheses about the function of zebra stripes including
camouflage
Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
,
predator avoidance, heat management, or social interaction.
He evaluated 18 different proposed explanations for the stripes, devising and carrying out quantitative tests to compare them. The evolutionary ecologist Tim Birkhead, writing in the ''Times Higher Education'', praised Caro's 2006 book ''Zebra Stripes'' as "an exemplary study", calling it "one long argument", a phrase used by
Darwin
Darwin may refer to:
Common meanings
* Charles Darwin (1809–1882), English naturalist and writer, best known as the originator of the theory of biological evolution by natural selection
* Darwin, Northern Territory, a territorial capital city i ...
of his ''
On the Origin of Species
''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life''),The book's full original title was ''On the Origin of Species by Me ...
'', summarizing it as "in essence a 300-page scientific paper".
Karin Brulliard, writing in ''
The Washington Post
''The Washington Post'' (also known as the ''Post'' and, informally, ''WaPo'') is an American daily newspaper published in Washington, D.C. It is the most widely circulated newspaper within the Washington metropolitan area and has a large n ...
'' under the headline "To figure out why the zebra got its stripes, this researcher dressed up like one", portrays Caro in a zebra costume "not used in his fieldwork", but also in a tailor-made striped suit in the Tanzanian bush, as well as in pelts of zebra and the unstriped wildebeest. The newspaper reports Caro as "absolutely convinced" that he has found the right explanation.
Matthew Cobb, writing in ''
New Scientist
''New Scientist'' is a magazine covering all aspects of science and technology. Based in London, it publishes weekly English-language editions in the United Kingdom, the United States and Australia. An editorially separate organisation publish ...
'', recalls
Rudyard Kipling
Joseph Rudyard Kipling ( ; 30 December 1865 – 18 January 1936)''The Times'', (London) 18 January 1936, p. 12. was an English novelist, short-story writer, poet, and journalist. He was born in British Raj, British India, which inspired much o ...
's children's book, ''
Just So Stories
''Just So Stories for Little Children'' is a 1902 collection of origin stories by the British author Rudyard Kipling. Considered a classic of children's literature, the book is among Kipling's best known works.
Kipling began working on the ...
'', in which the zebra got his stripes by standing half-in, half-out of the shadows "with the slippery-slidy shadows of the trees" on his body. Cobb calls ''Zebra Stripes'' a marvellous book and predicts it will encourage a generation to "tackle evolutionary biology's remaining enigmas, with or without the help of Kipling."
Michael Lemonick, writing in ''
The New Yorker
''The New Yorker'' is an American weekly magazine featuring journalism, commentary, criticism, essays, fiction, satire, cartoons, and poetry. Founded as a weekly in 1925, the magazine is published 47 times annually, with five of these issue ...
'' echoed the just-so-story theme.
Works
*
*
*
*
*
References
External links
UC Davis faculty profileUniversity of Bristol faculty profile
{{DEFAULTSORT:Caro, Tim
1951 births
Living people
British ecologists
Evolutionary biologists
Camouflage researchers
University of California, Davis faculty
Alumni of the University of Cambridge
Alumni of the University of St Andrews
Academics of the University of Bristol