Thrushes
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The thrushes are a
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
, Turdidae, with a worldwide distribution. The family was once much larger before biologists reclassified the former subfamily Saxicolinae, which includes the chats and European robins, as
Old World flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, bluethroat (''Luscinia svecica'') and north ...
s. Thrushes are small to medium-sized ground living birds that feed on insects, other invertebrates, and fruit. Some unrelated species around the world have been named after thrushes due to their similarity to birds in this family.


Characteristics

Thrushes are plump, soft-plumaged, small to medium-sized birds that inhabit wooded areas and often feed on the ground. The smallest thrush may be the shortwings, which have ambiguous alliances with both thrushes and
Old World flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, bluethroat (''Luscinia svecica'') and north ...
s. The lesser shortwing averages . The largest thrush is the
great thrush The great thrush (''Turdus fuscater'') is a species of bird in the family Turdidae. It is found in Bolivia, Colombia, Ecuador, Peru, and Venezuela. It is considered as the largest thrush in South America. The great thrush's size distinguishes it ...
at and ; the larger, commonly recognized blue whistling thrush is an
Old world flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, bluethroat (''Luscinia svecica'') and north ...
. The Amami thrush might, however, grow larger than the
great thrush The great thrush (''Turdus fuscater'') is a species of bird in the family Turdidae. It is found in Bolivia, Colombia, Ecuador, Peru, and Venezuela. It is considered as the largest thrush in South America. The great thrush's size distinguishes it ...
. Most species are grey or brown in colour, often with speckled underparts. They are
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores we ...
, but most species also eat worms,
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial molluscs, terrestrial gastropod mollusks that have gastropod shell, shel ...
s, and fruit (usually
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone fruit, stone or pit (fruit), pit although many wikt:pip#Etymology 2, pips or seeds may be p ...
). Many species are permanently resident in warm climates, while others migrate to higher latitudes during the summer, often over considerable distances. Thrushes build cup-shaped nests, sometimes lining them with mud. They lay two to five speckled eggs, sometimes laying two or more
clutch A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...
es per year. Both parents help raise the young. In almost all cases, the nest is placed on a branch; the only exceptions are the three species of
bluebird The bluebirds are a North American group of medium-sized, mostly insectivorous or omnivorous passerine birds in the genus ''Sialia'' of the thrush family (Turdidae). Bluebirds are one of the few thrush genera in the Americas. Bluebirds lay an ...
, which nest in holes.


Ecology

Turdidae species spread the seeds of plants, contributing to the dispersal of many species and the recovery of ecosystems. Plants have limited
seed dispersal In spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...
mobility away from the parent plant and consequently rely upon a variety of
dispersal vector A dispersal vector is an agent of biological dispersal that moves a dispersal unit, or organism, away from its birth population to another location or population in which the individual will reproduce. These dispersal units can range from pollen t ...
s to transport their propagules, including both
abiotic In biology and ecology, abiotic components or abiotic factors are non-living chemical and physical parts of the environment that affect living organisms and the functioning of ecosystems. Abiotic factors and the phenomena associated with them und ...
and biotic vectors. Seeds can be dispersed away from the parent plant individually or collectively, as well as dispersed in both space and time. Many bats and birds rely heavily on fruits for their diet, including birds in the families Cotingidae,
Columbidae Columbidae is a bird Family (biology), family consisting of doves and pigeons. It is the only family in the Order (biology), order Columbiformes. These are stout-bodied birds with small heads, relatively short necks and slender bills that in ...
,
Trogonidae The trogons and quetzals are birds in the order Trogoniformes which contains only one family, the Trogonidae. The family Trogonidae contains 49 species in seven genera. The fossil record of the trogons dates back 49 million years to the Early ...
, Turdidae, and
Ramphastidae Toucans (, ) are Neotropical birds in the family Ramphastidae. They are most closely related to the Toucan barbets. They are brightly marked and have large, often colorful bills. The family includes five genera and over 40 different species. ...
. While eating fruit, these animals swallow seeds and then later regurgitate them or pass them in their faeces. Such ornithochory has been a major mechanism of seed dispersal across ocean barriers. Other seeds may stick to the feet or feathers of birds and in this way may travel long distances. Seeds of grasses, spores of algae, and the eggs of molluscs and other invertebrates commonly establish in remote areas after long journeys of this sort. The Turdidae have a great ecological importance because some populations migrate long distances and disperse the seeds of endangered plant species at new sites, helping to eliminate inbreeding and increasing the genetic diversity of local flora.


Taxonomy

The family Turdidae was introduced (as Turdinia) by the French
polymath A polymath or polyhistor is an individual whose knowledge spans many different subjects, known to draw on complex bodies of knowledge to solve specific problems. Polymaths often prefer a specific context in which to explain their knowledge, ...
Constantine Samuel Rafinesque Constantine Samuel Rafinesque-Schmaltz (; 22 October 178318 September 1840) was a French early 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ult ...
in 1815. The taxonomic treatment of this large family has varied significantly in recent years. Traditionally, the Turdidae included the small Old World species, like the
nightingale The common nightingale, rufous nightingale or simply nightingale (''Luscinia megarhynchos''), is a small passerine bird which is best known for its powerful and beautiful song. It was formerly classed as a member of the thrush family Turdidae, ...
and
European robin The European robin (''Erithacus rubecula''), known simply as the robin or robin redbreast in the British Isles, is a small insectivorous passerine bird that belongs to the Old World flycatcher family Muscicapidae. It is found across Europe, ea ...
in the subfamily Saxicolinae, but most authorities now place this group in the
Old World flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, bluethroat (''Luscinia svecica'') and north ...
family
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, bluethroat (''Luscinia svecica'') and north ...
.
Molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
analysis has shown that the family Turdidae is a member of the superfamily Muscicapoidea and is
sister A sister is a woman or a girl who shares parents or a parent with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to ref ...
to the family
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, bluethroat (''Luscinia svecica'') and north ...
. The two families diverged in the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
around 17 million years ago. The family formerly included more species. At the time of the publication of the third edition of
Howard and Moore Complete Checklist of the Birds of the World The ''Howard and Moore Complete Checklist of the Birds of the World'' is a book by Richard Howard and Alick Moore which presents a list of the bird species of the world. It was the first single-volume world bird list to include subspecies names, ...
in 2003, the genera '' Myophonus'', '' Alethe'', ''
Brachypteryx ''Brachypteryx'' is a genus of passerine birds in the family Muscicapidae containing ten species known as shortwings, that occurs in southeast Asia. Shortwings are small birds with long legs, finely pointed bills, short tails and short rounded w ...
'' and '' Heinrichia'' were included in Turdidae. Subsequent
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
studies have shown that the species in these four genera are more closely related to species in the family
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, bluethroat (''Luscinia svecica'') and north ...
. As a consequence, these four genera are now placed in Muscicapidae. In contrast, the genus '' Cochoa'' which had previously been placed in Muscicapidae, was shown to belong in Turdidae.


Genera

The family contains 193 species, which are divided into 17 genera: * '' Grandala'' – grandala * '' Sialia'' – bluebirds (3 species) * '' Stizorhina'' – rufous thrushes (2 species) * '' Neocossyphus'' – ant thrushes (2 species) * '' Pinarornis'' – boulder chat * ''
Myadestes ''Myadestes'' is a genus of Solitaire (bird), solitaires, medium-sized mostly Insectivore, insectivorous birds in the thrush (bird), thrush family, Turdidae. They are found in the Americas and Hawaii, Hawaii, where several island species have be ...
'' – solitaires (13 species, including two recently extinct) * '' Chlamydochaera'' – fruithunter * '' Cochoa'' – cochoas (4 species) * '' Ixoreus'' – varied thrush * '' Ridgwayia'' – Aztec thrush * '' Cichlopsis'' – rufous-brown solitaire * '' Entomodestes'' – solitaires (2 species) * '' Hylocichla'' – wood thrush * '' Catharus'' – typical American thrushes and nightingale-thrushes (13 species) * '' Zoothera'' – Asian thrushes (22 species, including one recently extinct) * '' Geokichla'' – (21 species) * '' Turdus'' – true thrushes (104 species, including two recently extinct)


Cooking

The thrush is one of the many kinds of small bird that have in the past been trapped and eaten in much of Europe; the practice is now rare. Among traditional ways of cooking thrush were with
polenta Polenta (, ) is an Italian cuisine, Italian dish of boiled cornmeal that was historically made from other grains. It may be allowed to cool and solidify into a loaf that can be baked, fried or Grilling, grilled. The variety of cereal used is ...
or grilled on a skewer, in Italy; with
juniper Junipers are coniferous trees and shrubs in the genus ''Juniperus'' ( ) of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere as far south ...
berries in Belgium; and made into a
pâté Pâté ( , , ) is a forcemeat. Originally, the dish was cooked in a pastry case; in more recent times it is more usually cooked without pastry in a terrine. Various ingredients are used, which may include meat from pork, poultry, fish or bee ...
or terrine. The French cook and cookery writer Marie-Antoine Carême recommended cooking thrushes in crépinettes and serving with sauce Périgueux.


References


External links


Thrush videos
on the Internet Bird Collection
High-resolution photo gallery of around 100 species.
{{Authority control * Taxa named by Constantine Samuel Rafinesque