Thomas Holtz
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Thomas Richard Holtz Jr. (born September 13, 1965) is an American vertebrate
palaeontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
, author, and principal lecturer at the
University of Maryland The University of Maryland, College Park (University of Maryland, UMD, or simply Maryland) is a public land-grant research university in College Park, Maryland, United States. Founded in 1856, UMD is the flagship institution of the Univ ...
's Department of Geology. He has published extensively on the
phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ...
,
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
,
ecomorphology Ecomorphology or ecological morphology is the study of the relationship between the ecological, ecological role of an individual and its morphology (biology), morphological adaptations. The term "morphological" here is in the anatomical context. Bo ...
, and locomotion of terrestrial predators, especially on
tyrannosaurid Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to fifteen genera, including the eponymous ''Tyrannosaurus''. The exact number of genera ...
s and other
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s.Holtz, ''Thomas R. Holtz Jr.''. He wrote the book ''
Dinosaurs Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
'' and is the author or co-author of the chapters "Saurischia", "Basal Tetanurae", and "Tyrannosauroidea" in the second edition of ''
The Dinosauria ''The Dinosauria'' is an encyclopedia on dinosaurs, edited by paleontologists David B. Weishampel, Peter Dodson, and Halszka Osmólska. It has been published in two editions by the University of California Press, with the first edition in 1990 ...
''. He has also been consulted as a scientific advisor for the ''
Walking with Dinosaurs ''Walking with Dinosaurs'' is a 1999 six-part nature documentary television miniseries created by Tim Haines and produced by the BBC Science Unit, the Discovery Channel and BBC Worldwide, in association with TV Asahi, ProSieben and France 3. ...
'' BBC series as well as the
Discovery Discovery may refer to: * Discovery (observation), observing or finding something unknown * Discovery (fiction), a character's learning something unknown * Discovery (law), a process in courts of law relating to evidence Discovery, The Discovery ...
special ''
When Dinosaurs Roamed America ''When Dinosaurs Roamed America'' (sometimes shortened to ''When Dinosaurs Roamed'' outside of the United States) is a two-hour American television program (produced in the style of a traditional nature documentary) that first aired on the Disco ...
'', and has appeared in numerous documentaries focused on prehistoric life, such as ''
Jurassic Fight Club ''Jurassic Fight Club'' (titled ''Dinosaur Secrets'' in Australia and the United Kingdom) is a paleontology-based television miniseries on the History Channel which premiered in the United States in July 2008. ''Jurassic Fight Club'' was hosted ...
'' on
History History is the systematic study of the past, focusing primarily on the Human history, human past. As an academic discipline, it analyses and interprets evidence to construct narratives about what happened and explain why it happened. Some t ...
and ''
Monsters Resurrected ''Monsters Resurrected'' is an American edutainment television series that premiered on September 13, 2009, on the Discovery Channel. The program reconstructs extinct animals of both Mesozoic and Cenozoic. It is also called ''Mega Beasts''. Epi ...
'', ''
Dinosaur Revolution ''Dinosaur Revolution'' is a four-part American nature documentary produced by Creative Differences. It utilizes computer-generated imagery to portray dinosaurs and other animals from the Mesozoic era. The program was originally aired on the Disc ...
'' and ''
Clash of the Dinosaurs ''Clash of the Dinosaurs'' is a four-part television mini-series produced by Dangerous LTD for Discovery Channel. The show premiered on December 6, 2009, with the first two episodes scheduled back-to-back. ''Clash of the Dinosaurs'' was negative ...
'' on Discovery. Holtz is also the director of the Science and Global Change Program within the
College Park Scholars The University of Maryland, College Park (University of Maryland, UMD, or simply Maryland) is a public land-grant research university in College Park, Maryland, United States. Founded in 1856, UMD is the flagship institution of the Universi ...
living-learning community at the University of Maryland, College Park.


Early life

Holtz was born in California. He has stated that he wanted to be a dinosaur as a child, but upon learning that it is impossible to turn into a dinosaur he shifted his goals to study them. He attended
Johns Hopkins University The Johns Hopkins University (often abbreviated as Johns Hopkins, Hopkins, or JHU) is a private university, private research university in Baltimore, Maryland, United States. Founded in 1876 based on the European research institution model, J ...
, where he met his future wife. He then attended Yale, where he met
John Ostrom John Harold Ostrom (February 18, 1928 – July 16, 2005) was an American paleontologist who revolutionized the modern understanding of dinosaurs. Ostrom's work inspired what his pupil Robert T. Bakker has termed a " dinosaur renaissance". Begin ...
, who served as his academic advisor.


Theories

Holtz has come up with several new theories and hypotheses about the dinosaurs' classification. For example, he coined the terms
Maniraptoriformes Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyra ...
and
Arctometatarsus An arctometatarsalian organism is one in which the proximal part of the middle metatarsal is pinched between the surrounding metatarsals. The trait appears to be highly homoplastic, common in certain sorts of dinosaurs accustomed to running (among ...
. He also proposed two classification systems for theropods. The first is the clade
Arctometatarsalia Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia, Europe and North ...
, made up of
tyrannosauroids Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent b ...
,
ornithomimosaurs Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period (geology), Period of Laurasia (now Asia ...
, and
troodontids Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinos ...
, because all of these
coelurosaurs Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyran ...
had pinched middle
metatarsal The metatarsal bones or metatarsus (: metatarsi) are a group of five long bones in the midfoot, located between the tarsal bones (which form the heel and the ankle) and the phalanges ( toes). Lacking individual names, the metatarsal bones are ...
bones in their feet. In this proposed classification system, the tyrannosauroids were supposedly basal to a clade known as Bullatosauria, which was made up of the
Troodontidae Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinos ...
and the
Ornithomimosauria Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia, Europe and No ...
. These two groups were purported to form a
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
, because they both shared a common characteristic; which was a skull capsule. However, later on, both of these classification systems were found to be
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
, or "artificial", clades. For example, troodontids are now known to be deinonychosaurs, or "raptors", closely related to dromaeosaurids and birds. The discovery of basal tyrannosauroids, such as ''
Guanlong ''Guanlong'' (冠龍) is a genus of extinct proceratosaurid tyrannosauroid dinosaur from the Late Jurassic of China. The taxon was first described in 2006 by Xu Xing ''et al.'', who found it to represent a new taxon related to ''Tyrannosaurus' ...
'', which lacked an arctometatarsus, also helped to disprove this theory. And, eventually, the "skull capsule" in troodontids and ornithomimosaurs was found to be an example of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
, causing the clade Bullatosauria to be abandoned. Holtz was also a key figure in the discovery that tyrannosauroids were not
carnosaur Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods. While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Carn ...
s, as had been previously believed by most palaeontologists, but rather large
coelurosaur Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, ty ...
s. One of the first scientists to theorize this, Holtz contributed greatly to the debunking of a
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
Carnosauria that contained Megalosauroidea, Allosauroidea and Tyrannosauroidea. However, fellow vertebrate paleontologist Oliver Rauhut is known to recover a monophyletic Carnosauria that contains both Megalosauroidea and Allosauroidea in many of his publications, although like most researcher he agrees that Tyrannosauroids are likely Coelurosaurians as based on the conclusions made by Holtz.


Selected publications

* * * * * * *


Footnotes


References

* * {{DEFAULTSORT:Holtz, Thomas R. Jr. Living people American paleontologists University of Maryland, College Park faculty 1965 births